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Stuttgarter Beiträge zur Naturkunde 

Serie A (Biologie) 

Herausgeber: 

Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart 

Stuttgarter Beitr. Naturk. Ser. A Nr. 639 120 S. Stuttgart, 1. 8. 2002 

A Bibliography of Terrestrial Isopods 

(Crustacea: Isopoda: Oniscidea) 

By Helmut Schmalfuss and Karin Wolf-Schwenninger, Stuttgart 

Summary 

A reference list is presented containing most scientific publications on the biology of terrestrial 

isopods (suborder Oniscidea) published in a European language until the year 2000. Not 

considered are popular accounts, textbooks, faunal lists and preliminary reports if these are 

summarised in later publications, and publications before 1850 if they do not contain descriptions 

of new species. 

Zusammenfassung 

Die vorliegende Bibliografie enthält die meisten wissenschaftlichen Publikation über die 

Biologie der Landisopoden (Unterordnung Oniscidea), die in einer europäischen Sprache bis 

zum Jahre 2000 veröffentlicht wurden. Nicht berücksichtigt sind populärwissenschaftliche 

Artikel, Lehrbücher, Faunenlisten und vorläufige Berichte, wenn diese in einer späteren Publikation 

zusammengefasst werden, und Publikationen vor 1850, wenn sie keine Beschreibungen 

von neuen Arten enthalten. 

1. Introduction 

In the present list we tried to compile a complete bibliography of scientific publications 

on the isopodan suborder Oniscidea, covering all biological research fields. 

Such a list on a group of animals comprising some 3800 described species will, of 

course, never be complete. We shall certainly have missed a number of publications, 

especially in those more general topics as physiology, genetics, biochemistry, endocrinology 

etc. Nevertheless we think it will be useful for every student of this taxon 

to dispose of such a list which can be scrutinised for publications of a certain 

field of interest. This should justify to publish a bibliography, the reason to do it 

was, however, another one. We intend to publish in the near future a world list of all 

described species of terrestrial isopods, with detailed bibliographies for every species. 

This will be done in several parts, and the present bibliography will have to be 

used as the reference list.

2 stuttgarter beiträge zur naturkunde Ser. A, Nr. 639 

In the present list we have, on purpose, not included: 

1. Popular accounts without new scientific information; 

2. textbooks and treatises which do not contain new data or conceptions; 

3. publications consisting only of faunistic records of common or widespread species; 

4. reports on excursions with lists of the collected species if these are summarised 

in later publications; 

5. preliminary reports on results treated in detail in later publications; 

6. unpublished theses (Ph. D. or other); 

7. electronic publications not published in a printed form; 

8. publications that do not contain at least a title and summary written in Latin or 

Cyrillic letters (as e. g. papers written in Chinese). In the case of Russian publications 

the titles are either transcribed in Latin letters, or they are translated into English 

and put in square brackets; 

9. publications before 1850 if they do not contain descriptions of new species; 

most of this older literature is compiled in the bibliography of BUDDE-LUND 1885. 

The list contains publications that have appeared until the year 2000. 

For journals abbreviations are used according or comparable to those given in the 

World List of Scientific Periodicals. Other publications are cited at full length. 

The arrangement of publications from the same author(s) and year (a, b, c. . ..) is, 

for practical reasons, not necessarily chronological. 

2. Publications on terrestrial Isopoda (suborder Oniscidea) 

ABBOT, C. (1918): Reactions of land isopods to light. – J. exp. Zool. 27: 193–246; Philadelphia. 

– (1940): Shore isopods: Niches occupied, and degrees of transition toward land life with 

special reference to the family Ligydidae. – Proceedings 6 th Pacific Science Congress 3: 

505–511; Java. 

ABE, T. & IKEHARA, S. (1981): Ecological distribution of terrestrial isopods in the subtropical 

uninhabited islands (Uotsuri-jima and Minami-kojima, the Senkaku Islands, Japan). – 

Bull. Coll. Sci. Univ. Ryukyus 31: 61–70; Naha. 

ABRAHAM, A. & WOLSKY, A. (1929): Über die Nerven und Nervenendigungen der Antennen 

einiger Landasseln. – Zool. Anz. 84: 316–323; Leipzig. 

– & – (1930 a): Über ein neues Sinnesorgan der Landisopoden. – Zool. Anz. 87: 87–93; 

Leipzig. 

– & – (1930 b): Die Geruchsorgane der Landisopoden. – Z. Morph. Ökol. Tiere 17: 

441–463; Berlin. 

ABSOLON, K. (1933): O temnostnich balkanskych Titanethech. – Prˇíroda, Brno 26: 49–54; 

Brno. 

ABSOLON, K. & STROUHAL, H. (1932): Protonethes ocellatus nov. gen., nov. spec., eine neue 

Höhlen-Trichoniscide. – Zool. Anz. 101: 17–28; Leipzig. 

ACCOLA, S., TOMESCU, N., BERCIU, C. & RUSU, C. (1993): The reproduction of Trachelipus 

difficilis rotundatus. – Revue roum. Biol. (Biol. anim.) 38: 115–121; Bucharest. 

ADAMKEWICZ, S. (1969): Color polymorphism in the land isopod Armadillidium nasatum. – 

Heredity 24: 249–264; London. 

AFONSO, O. (1978): Une nouvelle espèce d’isopode troglobie du Portugal. – Publções Inst. 

Zool. Dr. Auguste Nobre 140: 9–23; Coimbra. 

AHMED, M. (1974): Studies on certain species of terrestrial Isopoda from Basrah region, Iraq. 

– Bull. Basrah nat. Hist. Mus. 1: 17–42; Basrah. 

AKAHIRA, Y. (1956): The function of thoracic processes found in females of the common 

woodlouse, Porcellio scaber. – J. Fac. Sci. Hokkaido Univ. (6) 12: 493–498; Sapporo.

schmalfuss et al., bibliography of terrestrial isopods 3 

– (1957): Effects of certain physiological solutions on the embryo of a terrestrial isopod, 

Porcellio scaber. – J. Fac. Sci. Hokkaido Univ. (6) 13: 359–363; Sapporo. 

AL-DABBAGH, K. & BLOCK, W. (1981): Population ecology of a terrestrial isopod in two 

Breckland grass heaths. – J. Anim. Ecol. 50: 61–77; Cambridge. 

AL-DABBAGH, K. & MARINA, B. (1986): The effect of starvation and breeding conditions on 

the respiration rate of Metoponorthus pruinosus. – Ekol. pol. 33: 705–714; Warsaw. 

ALEXANDER, C. (1969): Structure and properties of mechanoreceptors in the pereiopods of 

Ligia oceanica. – Comp. Biochem. Physiol. 29: 1197–1205; London. 

– (1970): Studies on the nervous system of an isopod crustacean, Ligia oceanica. – Comp. 

Biochem. Physiol. 33: 323–332; London. 

– (1971): Observations on receptor mechanisms in Ligia oceanica. – Comp. Biochem. 

Physiol. 40 A: 339–347; London. 

– (1972): Locomotion in the isopod crustacean, Ligia oceanica. – Comp. Biochem. Physiol. 

42 A: 1039–1047; London. 

– (1977): Antennal sense organs in the isopod Ligia oceanica. – Mar. Behav. Physiol. 5: 

61–77; London. 

ALIKHAN, M. (1968 a): The behaviour of Metoponorthus pruinosus in relation to humidity.- 

Curr. Sci. 37: 78; Bangalore. 

– (1968 b): The physiology of the wood-louse, Porcellio laevis. II. The occurrence of the 

goblet cells in the gut epithelium. – Curr. Sci. 37: 499–500; Bangalore. 

– (1968 c): The internal anatomy of the woodlouse, Metoponorthus pruinosus. – Can. J. 

Zool. 46: 321–327; Ottawa. 

– (1969): The physiology of the wood-louse, Porcellio laevis. I. Studies on the gut epithelium 

cytology and its relation to the maltase secretion. – Can. J. Zool. 47: 65–75; Ottawa. 

– (1972 a): Haemolymph and hepatopancreas copper in Porcellio laevis. – Comp. Biochem. 

Physiol. 42 A: 823–832; London. 

– (1972 b): The fine structure of the midgut epithelium in the wood-louse, Porcellio laevis. 

– Crustaceana, Suppl. 3: 101–109; Leyden. 

– (1972 c): Changes in the hepatopancreas metabolic reserves of Porcellio laevis during 

starvation and the moult-cycle. – Am. Midl. Nat. 87: 503–514; Notre Dame. 

– (1973): Ionic content of the haemolymph in Porcellio laevis. – Comp. Biochem. Physiol. 

44 A: 47–54; London. 

– (1981): Haemolymph pressure regulation in Porcellio laevis. – Archs int. Physiol. Biochim. 

89: 295–301; Liège. 

– (1983): Oxygen consumption and haemolymph pressure measurements in Armadillidium 

vulgare. – J. crust. Biol. 3: 25–33; Lawrence. 

– (1984): Haemodynamics in terrestrial Isopoda. – Symp. zool. Soc. London 53: 217–232; 

London. 

– (1991): Effects of temperature and copper and nickel accumulations on the metabolic 

rate and membrane electrical potential of the hepatopancreas cells of Oniscus asellus. – 

In: JUCHAULT, P. & MOCQUARD, J.(eds.): The Biology of Terrestrial Isopods III. Proceedings 

of the Third International Symposium on the Biology of Terrestrial Isopods, 

pp. 149–155; Poitiers. 

ALIKHAN, M. & PANI, A. (1989): Magnesium and manganese uptake and accumulation in the 

body tissue and their effect on the oxygen and nitrogen metabolism in Porcellio spinicornis. 

– Monitore zool. ital. (N. S.) Monogr. 4: 211–225; Florence. 

ALIKHAN, M. & SALEEM, M (1977): The role of phosphomonoesterases in the development 

and the moult cycle in Porcellio laevis. – Zool. Pol. 26: 176–185; Lwow. 

ALIKHAN, M. & STORCH, V. (1990): Copper and nickel uptake and accumulation, and their effects 

on redox and electrical potentials of hepatopancreatic cells of Oniscus asellus. – 

Can. J. Zool. 68: 651–655; Ottawa. 

AL-JETLAWI, A. & NAIR, G. (1994): Breeding and population biology of Armadillo officinalis 

in Benghazi, Libya. – J. arid Environm. 27: 241–255; London. 

ALLEE, W. (1926): Studies in animal aggregations: Causes and effects of bunching in land isopods. 

– J. exp. Zool. 45: 255–277; Philadelphia. 

ALLRED, D. & MULAIK, S. (1965): Two isopods of the Nevada Test Site. – Gt Basin Nat. 25: 

43–47; Provo.


ALLSPACH, A. (1987): Zum Vorkommen von Armadillidium nasatum in der Bundesrepublik 

Deutschland. – Hess. faun. Briefe 7: 20–27; Darmstadt. 

– (1989): Neunachweise von vier Landasselarten für Deutschland. – Stuttg. Beitr. Naturk. 

(Ser. A) 436: 8 pp.; Stuttgart. 

– (1992): Die Landasseln Hessens. – Natursch. heute 12: 1–146; Wetzlar. 

– (1996): The terrestrial isopods of the Bükk National Park. – In: The Fauna of the Bükk 

National Park, pp. 71–74; Budapest. 

ALLSPACH, A. & BAUSCHMANN, G. (1987): Stand der Erfassung bei Landasseln in Hessen. – 

Natursch. heute 14: 71–77; Wetzlar. 

ALLSPACH, A. & SZLAVECZ, K. (1990): The terrestrial isopod fauna of the Bátorliget Nature 

Reserves. – In: The Bátorliget Nature Reserves – after fourty years, pp. 251–257; Budapest. 

AMANIEU, M. (1960): Recherches écologiques sur les faunes des plages abritées de la région 

d’Arcachon. – Helgoländer wiss. Meeresunters. 19: 455–557; Kiel. 

– (1965): Cycle reproducteur annuel à Arcachon d’une population maritime de Porcellio 

scaber scaber. – C. r. hebd. Séanc. Acad. Sci. Paris 268: 5907–5910; Paris. 

AMMON, L. VON (1882): Ein Beitrag zur Kenntnis der vorweltlichen Asseln. – Sber. bayer. 

Akad. Wiss. (math.-phys. Kl.) 12: 507–550; Munich. 

AMOURIQ, L. (1967): Rôle des effets de groupe et de masse sur la croissance, la maturité sexuelle, 

la fécondité des jeunes de Porcellio scaber. – Bull. Soc. zool. Fr. 92: 177–185; Paris. 

ANDERSON, R. (1978): Armadillidium album in Co. Down. – Ir. Nat. J. 19: 202–203; Belfast. 

ANDERSSON, Å. (1960 a): A case of intersexuality in Benthana longicornis. – Ark. Zool. 12: 

415–419; Uppsala. 

– (1960 b): South American terrestrial isopods in the collection of the Swedish State Museum 

of Natural History. – Ark. Zool. 12: 537–570; Uppsala. 

ANDO, Y. (1996): Life history patterns of two coexisting isopods (Ligidium japonicum and Ligidium 

koreanum) in a Cruyptomeria japonica plantation of southwestern Japan. – Edaphologia 

56: 31–46 [no further data found]. 

ANDREEV, S. (1970): Beitrag zur Untersuchung der Landasseln Bulgariens. Neue Art der Gattung 

Trichoniscus. – C. r. Acad. bulg. Sci 23: 1135–1138; Sofia. 

– (1972): Beitrag zur Kenntnis der Landasseln in Bulgarien. II. – Izv. zool. Inst. Sofiya 34: 

177–188; Sofia. 

– (1984): Contribution à l’étude des isopodes terrestres de la Grèce. 1. Sur une nouvelle 

espèce cavernicole du genre Alpioniscus. – Biol. gallo-hellen. 11: 61–64; Athens. 

– (1985 a): Contribution à l’étude des isopodes terrestres de Grèce. 2. Cordioniscus antiparosi 

n. sp. de l’île Antiparos. – Crustaceana 49: 161–163; Leyden. 

– (1985 b): Deux nouvelles espèces cavernicoles du genre Trichoniscus de Bulgarie. – Acta 

zool. bulg. 27: 46–51; Sofia. 

– (1986 a): Cordioniscus bulgaricus sp. n. – premier représentant de la famille Styloniscidae 

en Bulgarie. – Acta zool. bulg. 31: 67–70; Sofia. 

– (1986 b): Contribution à l’étude des isopodes terrestres de la Grèce. 3. Sur trois nouvelles 

espèces des genres Cordioniscus, Alpioniscus et Trichoniscus et nouvelles données sur 

les isopodes terr. – Biol. gallo-hellen. 11: 153–164; Athens. 

– (1997): Contribution à l’étude des isopodes terrestres de Grèce. 6. Cordioniscus kalimnosi 

n. sp. – Hist. nat. bulg. 7: 13–16; Sofia. 

– (1998): Bulgarian endemic species of Isopoda. Balkan endemic species of Isopoda. Rare 

species of Isopoda. – In: MEINE, C. (ed.): Bulgaria’s biological Diversity: Conservation 

Status and Needs Assessment. Appendix 12, 13 and 14: pp. 132–133; Sofia. 

– (2000): Aperçu sur le genre Trichoniscus en Bulgarie et description d’une nouvelle espèce 

– Trichoniscus garevi n. sp. – Hist. nat. bulg. 11: 39–46; Sofia. 

ANDREEV, S. & BOZAROVA, F. (2000): Type material of Isopoda (Crustacea: Oniscidea, 

Anthuridae) from the collections of the National Museum of Natural History in Sofia. 

– Hist. nat. bulg. 11: 25–32; Sofia. 

ANDREEV, S. & TABACARU, I. (1972): Sur une nouvelle espèce du genre Trichoniscus de Bulgarie, 

Trichoniscus raitchevi n. sp. – C. r. Acad. bulg. Sci. 25: 385–388; Sofia. 

ANGELINI, G. (1935): Osservazioni sulla struttura del mesointestino di Armadillidium. – Memorie 

Soc. tosc. Sci. nat. 45: 1–15; Pisa. 

ANSENNE, A., COMPERE, P. & GOFFINET, G. (1988): Organisation, sclerotisation et composi-


tion minérale des sclerites d’une espèce d’isopode marin et de quatre espèces d’isopodes 

terrestres oniscoides. – IFREMER, Actes de Colloques 8: 39–45; Issy-les-Moulineaux. 

ANTHOLZ, I. & WEIDEMANN, G. (1985): Rekultivierung als ökologisches Problem: 7. Präferenzverhalten 

von Landasseln und Besiedlungserfolg. – Verh. Ges. Ökol. 13: 791–794; 

Göttingen. 

ARAUJO, P. DE (1999): Two new species of Alboscia from Rio Grande do Sul, Brasil. – Crustaceana 

72: 487–496; Leyden. 

ARAUJO, P. DE & BUCKUP, L.(1994 a): Nova especie de Trichorhina do sul do Brasil. – Iheringia 

(Ser. Zool.) 77: 129–134; Porto Alegre. 

– & – (1994 b): Two new species of terrestrial Isopoda from southern Brasil. – Spixiana 17: 

269–274; Munich. 

– & – (1996 a): Occorrência de Nagurus no sul do Brasil. – Nauplius 4: 161–163; Rio Grande. 

– & – (1996 b): Novos registros e uma espécie nova de Trichorhina do sul do Brasil. – Revista 

brasil. Zool. 13: 799–810; São Paulo. 

ARAUJO, P. DE & BUCKUP, L. & BOND-BUCKUP, G. (1996): Isópodos terrestres de Santa Catarina 

e Rio Grande so Sul, Brasil. – Iheringia (Ser. Zool.) 81: 111–134; Porto Alegre. 

ARAUJO, P. DE & BUENO, A. (1998): Novos registros de isópodos terrestres do sul do Brasil: 

Porcellionidae e Trichoniscidae. – Nauplius 6: 185–186; Rio Grande. 

ARAUJO, P. DE & LEISTIKOW, A. (1999): Philosciids with pleopodal lungs from Brazil, with descriptions 

of a new species. – Contr. Zool. 68: 109–141; The Hague. 

ARAUJO, P. DE & ZARDO, C. (1996): Uma nova espécie de Balloniscus do sul do Brasil. – Revista 

brasil. Zool. 12: 785–790; São Paulo. 

ARCANGELI, A. (1910 a): Armadillidium peraccai. Contributo alla migliore conoscenza di 

questo isopode italiano. – Processi verb. soc. tosc. Sci. nat. 1910: 1–8; Pisa. 

– (1910 b): Armadillidium Gestroi. Contributo alla migliore conoscenza di questo isopode 

italiano. – Monitore zool. ital. 21: 13–27; Florence. 

– (1911): Sopra due specie di Armadillidium della fauna italiana. – Memorie Soc. tosc. Sci. 

nat. 27: 187–204; Pisa. 

– (1913): Isopodi terrestri nuovi o poco noti di Italia. – Monitore zool. ital. 24: 183–202; 

Florence. 

– (1914 a): Escursioni zoologiche del Dr. ENRICO FESTA[VE] [VEE]nell’Isola di Rodi. 

Isopodi. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 28: 1–22; Torino. 

– (1914 b): La collezione di isopodi terrestri del R. Museo di Zoologia degli Invertebrati 

di Firenze. – Atti Soc. ital. Sci. nat. 52: 445–486; Milano. 

– (1921): Note isopodologiche. Il genere Platyarthrus. – Atti Soc. ital. Sci. nat. 60: 

189–210; Milano. 

– (1922 a): Isopodi terrestri del Museo Zoologico della R. Università di Napoli. – Annuar. 

Mus. zool. R. Univ. Napoli (N. S.) 5: 1–8; Naples. 

– (1922 b): Primo contributo alla fauna degli isopodi degli Astroni. – Annuar. Mus. zool. 

R. Univ. Napoli (N. S.) Suppl. 10: 1–2; Naples. 

– (1923 a): Revisione del gruppo degli haplophthalmi, isopodi terrestri. – Archo zool. ital. 

10: 259–322 and plates 7–8; Naples. 

– (1923 b): Cylisticus Pierantonii, nuova specie di isopodo dell’humus. – Boll. Musei 

Zool. Anat. comp. r. Univ. Torino (N. S. N. 2) 38: 1–4; Torino. 

– (1923 c): Caratteri sessuali secondari e conformazione dell’apparato copulatore di Philoscia 

elongata nel periodo riproduttivo. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 

(N. S. N. 3) 28: 1–7 and plate 1; Torino. 

– (1923 d): Armadillidium zangherii, nuova specie di isopodo terrestre italiano. – Boll. 

Lab. Zool. gen. agr. R. Scuola Agric. Portici 17: 131–136 and 1 plate; Portici. 

– (1924 a): Per una migliore conoscenza della fauna iopodologica della Libia. – Monitore 

zool. ital. 35: 223–233; Florence. 

– (1924 b): Contributo alla conoscenza degli isopodi della Catalogna. – Trab. Mus. Cienc. 

nat. Barcelona 4: 3–29 and plates 1–5; Barcelona. 

– (1924 c): Osservazioni sopra l’exopodite dei pleopodi del genere Tracheoniscus. – Boll. 

Lab. Zool. gen. agr. R. Scuola Agric. Portici 17: 176–186; Portici. 

– (1924 d): Gli isopodi terrestri della Sardegna. – Boll. Musei Zool. Anat. comp. r. Univ. 

Torino 39: 13–75 and plates 4–5; Torino.


– (1925 a): Ermafroditismo e partenogenesi negli isopodi terrestri (prima nota). – Monitore 

zool. ital. 36: 105–122; Florence. 

– (1925 b): Isopodi terrestri della Spagna settentrionale ed orientale, raccolti dal Dr. F. 

HAAS negli anni 1914–1919. – Abh. senckenb. naturforsch. Ges. 39: 131–137; Frankfurt/Main. 

– (1926 a): Contributo alla conoscenza della fauna isopodologica delle terre circonstanti 

all’alto Adriatico. – Atti Mus. civ. Stor. nat. Trieste 11: 1–62; Trieste. 

– (1926 b): Zoologische Ergebnisse einer Reise nach den Pelagischen Inseln und Sizilien. 

8. Isopoda. – Senckenbergiana 8: 265–270; Frankfurt/Main. 

– (1927 a): Paraperiscyphis Caligarii nuova specie di isopodo terrestre di Sumatra. – Boll. 


– (1927 b): Isopodi terrestri raccolti nell’Estremo Oriente dal Prof. FILIPPO SILVESTRI. – 

Boll. Lab. Zool. gen. agr. R. Scuola Agric. Portici 20: 169–211; Portici. 

– (1927 c): Revisione dei generi degli isopodi terrestri. I a nota. Sopra alcuni generi di Africa 

e di America. – Atti Soc. ital. Sci. nat. 66: 126–141; Milano. 

– (1927 d): Osservazioni sopra l’apparato copulatore degli isopodi terrestri. – Ric. Morf. 

Biol. anim. 1: 3–41. 

– (1928): Ligidium Ghigii n.sp. – Boll. Musei Lab. Zool. Anat. comp. r. Univ. Genova 8: 

1–2; Genoa. 

– (1929 a): Ricerche faunistiche nelle isole dell’Egeo. Isopodi. – Archo zool. ital. 13: 

259–268; Naples. 

– (1929 b): Specie nuove o poco note del genere Periscyphis ed osservazioni sulle cavità incubatorie 

degli isopodi terrestri. – Annuar. Mus. zool. R. Univ. Napoli 5: 1–20 and plate 

7; Naples. 

– (1929 c): Isopodi terrestri raccolti nel 1925 in Italia dal Dr. E. DUDICH ed esistenti nel 

Museo Nazionale Ungherese (Budapest). – Annls hist.-nat. Mus. natn. hung. 26: 60–71 

and plate 1; Budapest. 

– (1929 d): Pseudalbinismo in isopodi e fattori dello stesso. – Natura, Milano 20: 1–8; Milano. 

– (1929 e): Isopodi terrestri raccolti in Cuba dal Prof. F. SILVESTRI. – Boll. Lab. Zool. gen. 

agr. R. Scuola Agric. Portici 23: 129–148; Portici. 

– (1930 a): Isopodi terrestri raccolti nelle Isole Canarie dal Prof. FILIPPO SILVESTRI. – Boll. 


– (1930 b): Contributo alla conoscenza del „micorgenton“ di Costa Rica. – Boll. Lab. 

Zool. gen. agr. R. Scuola Agric. Portici 25: 1–29; Portici. 

– (1931 a): Gli isopodi italiani che sono stati finora erroneamente diagnosticati come Porcellio 

monticola e Porcellio lugubris. – Boll. Lab. Zool. agr. Bachic. R. Ist. sup. agr. Milano 

3: 1–23; Parma. 

– (1931 b): Circoniscus Bezzii, nuova specie di isopodo terrestre del Brasile. – Boll. Zool. 

2: 115–122; Naples. 

– (1931 c): Sul rapporto numerico dei sessi negli isopodi terrestri. – Boll. Musei Zool. 

Anat. comp. r. Univ. Torino 41: 1–34; Torino. 

– (1931 d): L’ermafroditismo negli isopodi terrestri. – Archo zool. ital. 17: 165–256; Naples. 

– (1931 e): Musoniscus romanorum, nuovo genere e nuova specie di isopodo terrestre di 

Cirenaica. – Boll. Lab. Zool. gen. agr. R. Scuola Agric. Portici 25: 39–43; Portici. 

– (1931 f): Due nuove specie del genere Rhyscotus. – Boll. Lab. Zool. gen. agr. R. Scuola 

Agric. Portici 25: 30–38; Portici. 

– (1932 a): Tracheoniscus Pierantonii n. sp. Nuovo isopodo umicolo del Trentino. – Annuar. 

Mus. zool. R. Univ. Napoli (N. S.) 6: 1–3; Naples. 

– (1932 b): Primo contributo alla conoscenza degli isopodi del Parco Nazionale del Gran 

Paradiso. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 42, No. 22: 1–18 and 6 plates; 

Torino. 

– (1932 c): Synarmadilloides roccatii è sinonimo di Gerutha pila. – Boll. Musei Zool. 

Anat. comp. r. Univ. Torino 42, No. 17: 1–2; Torino. 

– (1932 d): Isopodi terrestri di Dominica (Piccole Antille). – Boll. Musei Zool. Anat. 

comp. r. Univ. Torino 42, No. 18: 1–5 and 4 plates; Torino. 

– (1932 e): Sopra alcune specie di Hemilepistus. – Boll. Musei Zool. Anat. comp. r. Univ. 

Torino 42, No. 25: 1–10 and 1 plate; Torino.


– (1932 f): Escursione zoologica all’oasi di Marrakesch nell’aprile 1930. Isopodi terrestri. 

– Boll. Zool. 3: 225–232 and plate 2; Naples. 

– (1932 g): Considerazioni sopra la validità dei nomi generici Armadillo, Armadillidium, 

Oniscus, Porcellio. – Boll. Zool. 3: 123–127; Naples. 

– (1932 h): Sopra i caratteri che si debbono prendere in considerazione nel descrivere le 

specie di isopodi terrestri. Le amenità di un isopodologo. – Boll. Zool. 3: 273–284; Naples. 

– (1932 i): Isopodi terrestri raccolti dal Prof. SILVESTRI nel Nord-America. – Boll. Lab. 


– (1932 k): Isopodi terrestri raccolti in Somalia dal Dott. GUISEPPE RUSSO. – Boll. Lab. 


– (1932 l): Nuova specie di isopodo terrestre. Platyarthrus Haplophthalmoides. – Boll. 


– (1932 m): Isopodi terrestri raccolti in Albania del Dott. PIETRO PARENZAN. – Atti Accad. 

scient. veneto-trent.-istriana 22: 75–85 and plate 2; Padova. 

– (1932 n): Isopodi terrestri raccolti dal Dott. PARENZAN in una caverna presso Promontore 

(dintorni di Pola, Istria). – Atti Ist. veneto Sci. 91: 1039–1048 and plates 1–9; Venice. 

– (1932 o): Porcellionidi nuovi o poco noti d’Italia. Correzioni ed aggiunte. – Boll. Lab. 

Zool. agr. Bachic. R. Ist. sup. agr. Milano 4: 5–26; Parma. 

– (1932 p): Isopodi umicoli raccolti nel Trentino. – Studi trent. Sci. nat. 13: 229–233; Trento. 

– (1933 a): Per una migliore conoscenza di alcune specie di isopodi terrestri in rapporto 

alla loro distribuzione geografica. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 43: 

47–62; Torino. 

– (1933 b): Isopodi terrestri delle isole Samoa e considerazioni sopra la distribuzione geografica 

del genere Armadillo. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 43: 25–33 

and plates 1–4; Torino. 

– (1933 c): Due nuove specie cieche della famiglia Armadillidae. – Archo zool. ital. 19: 

389–403 and plates 5–6; Naples. 

– (1933 d): Isopodi terrestri di Somalia. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 

43: 63–69 and plates 1–6; Torino. 

– (1934 a): Nuovi contributi alla conoscenza della fauna delle isole dell’Egeo. III. Isopodi 

terrestri. – Boll. Lab. Zool. gen. agr. R. Scuola Agric. Portici 28: 37–69; Portici. 

– (1934 b): Trois Porcellio de la Grande Salvage et des Canaries. – Bull. Mus. natn. Hist. 

nat. Paris 6: 250–251; Paris. 

– (1934 c): Note di revisione sulla famiglia Armadillidae. – Boll. Musei Zool. Anat. comp. 

r. Univ. Torino 44: 83–119 and plates 1–4; Torino. 

– (1934 d): Due specie ed un genere di isopodi terrestri nuovi per la Libia. – Boll. Musei 

Zool. Anat. comp. r. Univ. Torino 44: 213–220 and plates 1–5; Torino. 

– (1934 e): Le specie di Armadilllidium della Sicilia. – Boll. Musei Zool. Anat. comp. r. 

Univ. Torino 44: 173–191 and plates 1–4; Torino. 

– (1934 f): Isopodi terrestri raccolti nel Caracorum dalla Spedizione di S.A.R. il Duca di 

Spoleto. – Annali r. Ist. sup. agr. Milano 1: 1–23 and plates 1–3; Parma. 

– (1934 g): Anaphiloscia sicula, nuova specie di isopodo umicolo della Sicilia. – Boll. Lab. 


– (1935 a): Isopodi del Museo civico di Storia naturale di Milano. – Atti Soc. ital. Sci. nat. 

74: 204–222 and plates 1–7; Milano. 

– (1935 b): Sopra alcuni sottogeneri di Armadillidium. – Boll. Musei Zool. Anat. comp. r. 


– (1935 c): Eoniscus simplicissimus, un pigmeo degli isopodi terrestri, appartenente a nuova 

specie, nuovo genere, nuova famiglia. – Boll. Zool. 6: 203–212; Naples. 

– (1935 d): Paraschizidium hispanum, nuova specie di isopodo umicolo della famiglia Armadillidiidae. 

– Eos 11: 173–176; Madrid. 

– (1935 e): Isopodi terrestri di caverne della Spagna. – Eos 10: 171–195; Madrid. 

– (1935 f): Isopodi terrestri di Sumatra raccolte dal Sig. J. C. VAN DER MEER MOHR. – 

Miscnea zool. sumatr. 98: 1–4; Medan. 

– (1936 a): Isopodi terrestri dell’Isola di Cipro. – Boll. Musei Zool. Anat. comp. r. Univ. 

Torino 45: 5–25; Torino.


– (1936 b): Isopodi terrestri. Correzioni aggiunte, critiche. – Boll. Musei Zool. Anat. 

comp. r. Univ. Torino 45: 257–279; Torino. 

– (1936 c): Porcellio flavocinctus e Porcellio pelseneeri n. sp. – Mém. Mus. r. Hist. nat. 

Belg. (2. Ser.) 3: 185–196; Brussels. 

– (1936 d): Atlantidium barretoi, isopodo terrestre dell’Isola di Madera. Nuova specie e 

nuovo genere della famiglia Armadillidiidae. – Rossegna faun. 3: 44–46; Rome. 

– (1936 e): Atlantidium, nuovo genere della famiglia Armadillidiidae. – Boll. Musei Zool. 


– (1936 f): Un genere e due specie nuovi di isopodi terrestri del Brasile. – Archo zool. ital. 

23: 201–208 and plate 4; Naples. 

– (1936 g): Gli isopodi terrestri del Portogallo. – Boll. Lab. Zool. gen. agr. R. Scuola 

Agric. Portici 29: 1–39; Portici. 

– (1936 h): Specie nuove di isopodi terrestri della Spagna raccolti dal Prof. F. SILVESTRI. – 

Boll. Lab. Zool. gen. agr. R. Scuola Agric. Portici 29: 40–46; Portici. 

– (1937): Nuovi contributi alla conoscenza della fauna delle isole italiane dell’Egeo. VIII. 

Isopodi terrestri (2a nota). – Boll. Lab. Zool. gen. agr. R. Scuola Agric. Portici 30: 

75–86; Portici. 

– (1938 a): Philoscia elongata e Chaetophiloscia hastata appartengono alla stessa specie. 

Dimostrazione sperimentale. – Atti Accad. Sci. Torino 73: 1–5; Torino. 

– (1938 b): Tylos Latreillii, suoi biotopi, sua area di diffusione. – Boll. Musei Zool. Anat. 

comp. r. Univ. Torino 46: 139–151 and plates 1–6; Torino. 

– (1938 c): Il genere Spelaeonethes. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 46: 

157–168 and plates 1–7; Torino. 

– (1938 d): Isopodi raccolti in Asia Minore dal Sig. HENRI GADEAU DE KERVILLE. – In: 

Voyage zoologique d’HENRI GADAEU DE KERVILLE en Asie Mineure (Avril-Mai 1912), 

vol. 1: 29–65; Rouen. 

– (1938 e): Note di revisione del genere Androniscus. 1. Il gruppo roseus. – Boll. Musei 

Zool. Anat. comp. r. Univ. Torino 46: 103–120 and plates 1–4; Torino. 

– (1938 f): Androniscus dentiger è sempre provvisto di occhi. – Boll. Musei Zool. Anat. 


– (1938 g): Titanethes albus e Titanethes dahli. – Grotte Ital. (Ser. 2) 2: 44–52; Trieste. 

– (1938 h): Titanethes albus e Titanethes dahli appartengono alla stessa specie che può 

avere o no organi simbiotici o micetomi nel sesso maschile. – Atti Accad. Sci. Torino 73: 


– (1939 a): Crustacea, Isopoda. – In: Missione biologica nel Paese dei Borana. Raccolte 

zoologiche, vol. 3: 399–415; Rome. 

– (1939 b): Androniscus dentiger in Cirenaica. – Boll. Musei Zool. Anat. comp. r. Univ. 

Torino 47: 121–123 and plates 1–2; Torino. 

– (1939 c): A proposito di sedicenti ghiandole di Titanethes. – Boll. Musei Zool. Anat. 


– (1939 d): Per una migliore conoscenza del genere Mesoniscus e della sua posizione sistematica. 

– Boll. Musei Zool. Anat. comp. r. Univ. Torino 47: 7–36 and plates 1–8; Torino. 

– (1939 e): Esperimenti male condotti e male interpretati per giudicare specie del genere 

Hyloniscus. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 47: 173–176; Torino. 

– (1939 f): Sopra alcune forme della famiglia Armadillidiidae erroneamente interpretate. – 

Boll. Musei Zool. Anat. comp. r. Univ. Torino 47: 169–171; Torino. 

– (1939 g): Murgeoniscus anellii, nuovo genere e nuova specie di isopodo terrestre cavernicolo 

italiano. – Grotte Ital. (Ser. 2 a) 3: 1–8 and plates 9–15; Trieste. 

– (1939 h): Sopra mutazioni ottenute sperimentalmente in isopodi terrestri. Nota I. – Atti 

Accad. Sci. Torino 74: 1–35; Torino. 

– (1939 i): Sopra i pigmenti degli isopodi terrestri. – Atti Accad. Sci. Torino 74: 3–11; Torino. 

– (1939 j): Sardoniscus pusillus, nuovo genere e nuova specie di isopodo terrestre di Sardegna. 

– Boll. Musei Zool. Anat. comp. r. Univ. Torino 47: 125–129 and plates 1–2; Torino. 

– (1939 k): I generi Oroniscus e Strouhaloniscus. – Boll. Musei Zool. Anat. comp. r. Univ. 

Torino 47: 157–168; Torino.


– (1939 l): Sopra presunte razze e sottospecie di isopodi terrestri ottenute in domesticità. 

– Boll. Musei Zool. Anat. comp. r. Univ. Torino 47: 105–108; Torino. 

– (1940 a): Isopodi terrestri dell’Africa orientale italiana. – Riv. Biol. colon. 3: 381–385; 

Rome. 

– (1940 b): Il genere Alpioniscus. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 48: 


– (1940 c): Sopra alcune specie di Armadillidium appartenenti al sottogenere „Marginiferum“. 

– Boll. Musei Zool. Anat. comp. r. Univ. Torino 48: 163–171 and plates 1–6; 

Torino. 

– (1940 d): Note di revisione del genere Androniscus. – Boll. Musei Zool. Anat. comp. r. 


– (1941): Crustacea, Isopoda. – In: Missione biologica Sagan-Omo, vol. 12: 5–18; Rome. 

– (1942 a): Appunti sopra Ligia italica ed i suoi spermatofori.- Archo ital. Anat. Embriol. 

47: 1–6; Florence. 

– (1942 b): Porcellio monticola e Porcellio verhoeffi. – Boll. Musei Zool. Anat. comp. r. 

Univ. Torino 49: 109–111; Torino. 

– (1942 c): Ermafroditismo ed intersessualitá in crostacei isopodi terrestri. – Boll. Musei 

Zool. Anat. comp. r. Univ. Torino 49: 139–149 and plate 1; Torino. 

– (1942 d): Porcellio (Lucasius) leptotrichoides, nuova specie di crostaceo isopodo terrestre 

dell’isola Grande Salvage. – Boll. Musei Zool. Anat. comp. r. Univ. Torino 49: 

223–225 and plate 1; Torino. 

– (1945): Considerazioni sopra l’origine delle forme cavernicole degli isopodi terrestri. – 

Archo zool. ital., Suppl. 34: 115–147; Naples. 

– (1946 a): Due specie nuove di triconiscidi della sottofamiglia Haplophthalminae: Calconiscellus 

(Cypholambrana) Malanchinii e Leucocyphoniscus Torrii. – Boll. Ist. Mus. 

Zool. Univ. Torino 1: 19–21 and plates 1–4; Torino. 

– (1946 b): Come viene applicata la geologia da uno zoologo per risolvere un problema 

zoogeografico. – Boll. Ist. Mus. Zool. Univ. Torino 1: 15–17; Torino. 

– (1946 c): Appunti sulla riproduzione degli isopodi terrestri. – Boll. Ist. Mus. Zool. Univ. 

Torino 1: 5–13; Torino. 

– (1947): Raccolte faunistiche compiute nel Gargano da A. GHIGI e F. P. POMINI. II. Isopodi 

terrestri. – Acta pontif. Acad. Sci. 12: 93–114; Rome. 

– (1947 b): La posizione sistematica del genere Helleria – Archo zool. ital. 31: 373–389; 

Naples. 

– (1947 c): Un rappresentante della famiglia Rhyscotidae in Asia. – Atti Accad. Sci. Torino 

81–82: 1–2; Torino. 

– (1948 a): Il genere Halophiloscia nella regione mediterranea. – Archo zool. ital. 33: 

437–428; Naples. 

– (1948 b): Sopra una particolare mutazione rufinica nel genere Armadillidium. – Archo 

zool. ital. 33: 203–211; Naples. 

– (1948 c): Sopra la existenza e la variabilità del distretto respiratorio negli exopoditi dei 

pleopodi del genera Philoscia. – Archo zool. ital. 33: 195–202; Naples. 

– (1948 d): Schizidiinae sottofamiglia di Armadillidiidae. – Boll. Ist. Mus. Zool. Univ. Torino 

1: 211–272; Torino. 

– (1948 e): Crostacei isopodi terrestri della Grotte di Bossea (Mondovi). – Boll. Ist. Mus. 

Zool. Univ. Torino 1: 23–28; Torino. 

– (1948 f): Il processo della amitosi nei crostacei isopodi. – Boll. Ist. Mus. Zool. Univ. Torino 

1: 125–136; Torino. 

– (1948 g): Una specie nuova del genere Toradjia. – Boll. Ist. Mus. Zool. Univ. Torino 1: 


– (1948 h): Osservazioni sopra la muta dei crostacei isopodi terrestri. – Boll. Ist. Mus. 


– (1949): La famiglia Rhyscotidae. – Boll. Ist. Mus. Zool. Univ. Torino 2: 5–36 and plate 

1; Torino. 

– (1950 a): Gli isopodi terrestri della Sardegna. 2a edizione. – Boll. Ist. Mus. Zool. Univ. 

Torino 2: 77–191 and plates 1–7; Torino. 

– (1950 b): Exploration du Parc National Albert, Mission H. DAMAS, fascicule 15, Isopodi 

terrestri, 80 pp. and 121 plates; Brussels.


– (1950 c): Gli isopodi terrestri dell’estuario veneto. – Atti Mus. civ. Stor. nat. Trieste 18: 

13–59; Trieste. 

– (1951 a): Le specie di isopodi terrestri che furono erroneamente assegnate al genere Alloniscus. 

– Memorie Mus. civ. Stor. nat. Verona 6: 236–252 and plates 1–4; Verona. 

– (1952 a): Isopodi terrestri di Albania. – Boll. Ist. Mus. Zool. Univ. Torino 3: 6–38 and 

plates 1–12; Torino. 

– (1952 b): Le caratteristiche della famiglia Eubelidae. – Boll. Ist. Mus. Zool. Univ. Torino 

3: 61–80; Torino. 

– (1952 c): Considerazioni sopra la pertinenza al genere Protracheoniscus di Protracheoniscus 

pierrei. – Boll. Ist. Mus. Zool. Univ. Torino 3: 99–101; Torino. 

– (1952 d): Appunti sopra il genere Tylos. – Boll. Ist. Mus. Zool. Univ. Torino 3: 133–141; 

Torino. 

– (1952 e): Correzioni riguardanti crostacei isopodi terrestri dell’Estremo Oriente. – Archo 


– (1952 f): La evoluzione del sistema respiratorio dell’exopodite dei pleopodi nelle famiglie 

dei porcellionidi e degli eubelidi. – Boll. Zool. 19: 297–304; Naples. 

– (1952 g): Porcellio silvestrii è una buona specie e non una sottospecie di P. monticola. – 

Boll. Zool. 19: 371–376; Naples. 

– (1952 h): La fauna isopodologica terrestre della Puglia e delle isole Tremiti e la sua probabile 

origine in rapporto alla diffusione transadriatica di specie. – Memorie Biogeogr. 

adriat. 2: 109–172; Venice. 

– (1953 a): A proposito della spanadria nei crostacei isopodi terrestri. – Boll. Zool. 20: 

65–72; Naples. 

– (1953 b): Precisazioni ecologiche sui generi Ligia e Tylos. – Atti Accad. Sci. Torino 87: 


– (1953 c): Appunti sopra il genere Trachelipus (Tracheoniscus) considerato in rapporto ad 

altri generi di porcellionidi. – Boll. Ist. Mus. Zool. Univ. Torino 4: 349–358; Torino. 

– (1953 d): Buddelundiella Zangherii, nuove specie di isopodo terrestre umicolo. – Boll. 

Zool. 19: 309–316; Naples. 

– (1954 a): Considerazioni sopra la volvazione dei crostacei isopodi terrestri e sopra alcune 

coaptazioni ad essa connesse. – Boll. Ist. Mus. Zool. Univ. Torino 4: 75–93; Torino. 

– (1954 b): Contributo alle conoscenze sugli eubelidi di Asia. – Boll. Ist. Mus. Zool. Univ. 

Torino 4: 9–17 and 1 plate; Torino. 

– (1954 c): Revisione di alcune specie italiane del genere Armadillidium. – Boll. Ist. Mus. 

Zool. Univ. Torino 4: 1–31 and plates 1–16; Torino. 

– (1954 d): I rappresentanti della famiglia Porcellionidae hanno sempre il flagello delle antenne 

esterne composto di due articoli. – Boll. Ist. Mus. Zool. Univ. Torino 4: 119–122; 

Torino. 

– (1954 e): Phalloniscus pygmaeus è proprio un Phalloniscus? – Boll. Ist. Mus. Zool. Univ. 


– (1954 f): Biogeografia dell’isola di Zannone. Fauna. Isopodi. – Rc. Accad. naz. XL (Ser. 

IV) IV+V: 77–78; Rome. 

– (1955): Considerazioni sopra il cosidetto „coxopodite“ dei pereiopodi dei crostacei isopodi 

terrestri. – Atti Accad. Sci. Torino 89: 1–9; Torino. 

– (1956 a): Crostacei isopodi dei Monti Sibillini. – Memorie Mus. civ. Stor. nat. Verona 5: 

179–182; Verona. 

– (1956 b): Sul dimorfismo sessuale di due specie di Ligia. – Atti Accad. Sci. Torino 90: 


– (1956 c): Come si creano specie nuove di crostacei isopodi terrestri. – Boll. Ist. Mus. 


– (1956 d): Thermocellio congolensis subsp. Patrizii, nuova specie di isopodo terrestre per 

il Kenya. – Boll. Ist. Mus. Zool. Univ. Torino 5: 63–64; Torino. 

– (1957 a): I generi Diploexochus, Venezillo, Paramardillo [sic]. – Boll. Ist. Mus. Zool. 

Univ. Torino 5: 101–142; Torino. 

– (1957 b): Il genere Armadilloniscus e gli Scyphacidae. – Atti Accad. Sci. Torino 91: 

399–428; Torino. 

– (1957 c): Rapporti del sistema respiratorio degli exopoditi dei pleopodi del genere Tylos 

con la ecologia dello stesso. – Atti Accad. Sci. Torino 91: 1–12; Torino.


– (1957 d): Il genere Armadillo. – Boll. Ist. Mus. Zool. Univ. Torino 5: 165–191 and plates 


– (1957 e): Missione ZAVATTARI per l’esplorazione biogeografica delle Isole Pelagie. Isopodi 

terrestri. – Riv. Biol. colon. 15: 83–95; Rome. 

– (1958 a): La fauna isopodologica degli arcipelaghi di Madera e delle Canarie: la sua 

imortanza per la sistematica e la biogeografia. – Mems Estud. Mus. Zool. Univ. Coimbra 

255: 1–108; Coimbra. 

– (1958 b): Le specie di isopodi terrestri che furano erroneamente assegnate al genere Alloniscus. 

– Memorie Mus. civ. Stor. nat. Verona 6: 239–252 and plates 1–4; Verona. 

– (1960 a): Revisione del genere Alloniscus. – Boll. Ist. Mus. Zool. Univ. Torino 6: 15–79 


– (1960 b): Sulla distribuzione geografica del genere Chaetophiloscia. – Boll. Ist. Mus. 


– (1963): Precisazioni sopra il genere Nagurus (= Nagara). – Boll. Ist. Mus. Zool. Univ. 


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Int. J. Speleol. 5: 153–162; Weinheim. 

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ARGANO, R., BALDARI, F. & MANICASTRI, C. (1982): Isopodi sotterranei italiani. – Lavori Soc. 

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ARGANO, R. & PESCE, G. (1974): Un elumino mirmecofilo di Sardegna: Typhloschizidium cottarellii 

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ARGANO, R. & RAMPINI, M. (1973): Nota sulla distribuzione dei Trichoniscidae in Sardegna. 

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ARGANO, R. & UTZERI, C. (1973): Oniscoidea delle Isole Ponziane: Eluminae. – Fragm. ent. 

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ARMAS, L. DE & DAVILA, N. (1988): Isopodos terrestres de Cuba. 1. Notes sobre la familia Tylidae. 

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biol. Fr. Belg. 75: 364–368; Paris. 

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– (1948 b): Espèces nouvelles d’isopodes terrestres cavernicoles et endogés. – Notes 


– (1948 c): Les variations de Phymatoniscus tuberculatus. – Bull. Mus. natn. Hist. nat. 226: 

1468–1470; Paris. 

– (1948 d): Les isopodes volvationnels exoantennés et la genèse de leurs coaptations. – 


– (1948 e): L’origine de la ligie italique. – Bull. Inst. océanogr. 931: 13 pp.; Monaco. 

– (1948 f): La faune isopodique française. Sa répartition, ses origines et son histoire. – Revue 

fr. Ent.15: 101–139; Paris. 

– (1948 g): Mission scientifique du Fezzan (1944–1945). V. Zoologie (arthropodes). Isopodes 

terrestres récoltés par M. le professeur F. BERNARD. – Publ. Inst. Rech. sahar. 

Univ. Alger 3: 1–26; Algiers. 

– (1948 h): Les coaptations des oniscoïdes volvationnels exoantennés. – C. r. hebd. Séanc. 


– (1949): La faune nord-atlantique. – Revue fr. Ent. 16: 1–12; Paris. 

– (1950 a): Les modalités de l’évolution dans le genre Porcellio. – C. r. Séanc. Acad. Sci. 

Paris 230: 1691–1693; Paris. 

– (1950 b): Étude des caractères sexuels secondaires de Trichoniscus suivie d’un essai de 

groupement des espèces appartenant a ce genre. – Archs Zool. exp. gén. 87: 116–133; 

Paris. 

– (1950 c): Biospeologica LXXI. Campagne spéologique de C. BOLIVAR et R. JEANNEL 

dans l’Amerique du Nord (1928). Isopodes terrestres recueillis par C. BOLIVAR et R. JE- 

ANNEL (1928) et le Dr. HENROT (1946). – Archs Zool. exp. gén. 87: 183–210; Paris. 

– (1950 d): Facteurs climatiques et variants sexuels dans le genre Porcellio. – C. r. Séanc. 

Acad. Sci. Paris 230: 1691–1693. 

– (1950 e): L’évolution d’une coaptation unisexuelle et son interprétation ontogénique. – 


– (1950 f): Espèces nouvelles d’isopodes terrestres cavernicoles et endogés. II. – Notes 


– (1950 g): Sur une collection d’isopodes terrestres rassemblée par F. PIERRE, dans le sud 

algèrien, et sur la présence d’un nouvel élément irano-touranien dans la faune saharienne. 


– (1950 h): Le Canigou, massif de refuge. – Bull. Soc. Hist. nat. Toulouse 84: 197–201; 

Toulouse. 

– (1951 a): Le genre Porcellio. Évolution et systématique. – Mém. Mus. natn. Hist. nat. 

(N. S., Ser. A) 3: 81–192; Paris. 

– (1951 b): Étude des caractères sexuels secondaires de Trichoniscus suivie d’un essai de 

groupement des espèces appartenant à ce genre. – Archs Zool. exp. gén. 87: 116–133; 

Paris. 

– (1951 c): Support for the proposals relating to the generic name „Ligia“ Fabricius, 1798 

(class Crustacea, order Decapoda) and associated problems submitted by the late Miss 

M. BUITENDIJK and Dr. L. B. HOLTHUIS. – Bull. zool. Nomencl. 2: 345; London. 

– (1951 d): Objection to proposal submitted by Professor MARTIN L. ACZÉL in favour of 

the addition of the name „Tylos“ to the „Official List of generic Names in Zoology. – 

Bull. zool. Nomencl. 2: 347; London. 

– (1952 a): Biospeologica LXXIII. Isopodes terrestres (troisième série). – Archs Zool. 

exp. gén. 88: 231–362; Paris. 

– (1952 b): La répartition du complexe trichoniscoïde et les théories géologiques. – C. r. 


– (1952 c): Sur les caractères de Porcellio nigrogranulatus. – Bull. Mus. natn. Hist. nat. (2. 

Ser.) 24: 522–525; Paris. 

– (1952 d): Étude des isopodes terrestres récoltés au Vénézuela par le Dr. G. MARCUZZI. 

– Memorie Mus. civ. Stor. nat. Verona 3: 59–203; Verona. 

– (1952 e): Phalloniscus bolivianus n. sp. – Bull. Mus. natn. Hist. nat. (2. Ser.) 24: 526–529; 

Paris. 

– (1952 f): Les trichoniscides de l’hémisphère austral. – Mém. Mus. natn. Hist. nat. (N. S., 

Ser. A) 6: 1–116; Paris.


– (1952 g): Proposition en faveur du maintien du nom de „Tylos“ (Latreille MS.) Audouin 

1825 (Crustacea; Isopoda terrestria). – Bull. zool. Nomencl. 6: 174–176; London. 

– (1952 h): La répartition du complexe trichoniscoïde et la paléogéographie. – C. r. Séanc. 


– (1953 a): Remarques systématiques, morphologiques et biogéographiques sur un groupe 

de Trichoniscidae nord-atlantiques. – Bull. Mus. natn. Hist. nat. (2. Ser.) 25: 368–375; 

Paris. 

– (1953 b): La classification de la famille des Trichoniscidae. – Bull. Mus. natn. Hist. nat. 

(2. Ser.) 25: 276–278; Paris. 

– (1953 c): Sur la présence du genre Phalloniscus en Espagne, et son explication biogéographique. 

– Revue fr. Ent. 20: 68–80; Paris. 

– (1953 d): Isopodes terrestres récoltés dans les grottes de la Corse, par le Professeur P.- 

A. REMY, en 1942 et 1948. – Notes biospéol. 8: 153- 166; Paris. 

– (1953 e): La famille des Squamiferidae et l’origine des Platyarthrus. – Bull. Soc. zool. Fr. 

77: 371–388; Paris. 

– (1953 f): L’évolution considérée comme phénomène de développement. Les variations 

de Phymatoniscus tuberculatus. – Bull. biol. Fr. Belg. 87: 414–430; Paris. 

– (1953 g): Espèces nouvelles ou peu connues de Trichoniscidae cavernicoles recueillies en 

Espagne orientale. – Notes biospéol. 8: 51–66; Paris. 

– (1953 h): Les isopodes terrestres des provinces d’Almeria et de Granada. – Archivos 

Inst. Aclimat. 1: 45–75; Almeria. 

– (1953 i): A new terrestrial isopod from Oregon, Caucasonethes rothi n. sp. – Pacif. Sci. 

7: 175–178; Honolulu. 

– (1954 a): Une nouvelle espèce cavernicole appartenant a la famille des Armadillidiidae 

et provenant de l’Espagne orientale. – Notes biospéol. 9: 61–66; Paris. 

– (1954 b): Une nouvelle espèce cavernicole de Trichoniscoides, T. Drescoi n. sp., provenant 

des Asturies. – Notes biospéol. 9: 67–69; Paris. 

– (1954 c): Isopodes terrestres. Résultats biogéographiques. – In: COIFFAIT, H., CASSA- 

GNAU, P & VANDEL, A., Contribution a la connaissance de la faune cavernicole et endogée 

du Liban (Mission H. COIFFAIT au Liban, 1951). – Notes Mém. Moyen-Orient 5: 

166–169; Paris. 

– (1954 e): Note sur deux Armadillidiidae endogés et cavernicoles de la région de Grasse. 

– Notes biospéol. 9: 49–60; Paris. 

– (1954 f): Les trichoniscides endogés et cavernicoles appartenant au genre Nesiotoniscus. 

– Notes biospéol. 9: 149–156; Paris. 

– (1954 g): Le statut systématique de trois porcellions de l’Espagne orientale. – Bull. Mus. 

natn. Hist. nat. (2. Ser.) 26: 491–495; Paris. 

– (1954 h): Mission J. CADENAT aux Iles du Cap-Vert. Isopodes terrestres – Bull. Inst. fr. 

Afr. noire (Ser. A) 16: 466–478; Paris. 

– (1954 i): Sur une espèce mal connue d’isopode terrestre, Phalloniscus pygmaeus. – Bull. 

Mus. natn. Hist. nat. (2. Ser.) 26: 226–230; Paris. 

– (1954 k): Étude des isopodes terrestres recueillis aux Iles Canaries par J. MATEU en 

mars-avril 1952. – Mém. Mus. natn. Hist. nat. (N. S., Ser. A) 8: 1–60; Paris. 

– (1954 l): Nesiotoniscus delamarei n. sp. – Vie Milieu 4: 664–668; Paris. 

– (1954 m): L’évolution considérée comme phénomène de développement. – Bull. Soc. 

zool. Fr. 79: 341–356; Paris. 

– (1954 n): Le peuplement isopodique de Corse; étude biogéographique. – Revue fr. Ent. 

21:72–84; Paris. 

– (1954 o): Description d’une nouvelle espèce de Bathytropa, B. colasi n. sp. – Bull. Mus. 

natn. Hist. nat. (2. Ser.) 26: 80–84; Paris. 

– (1955 a): Isopodes terrestres récoltés dans les grottes de l’Attique par M. PIERRE STRI- 

NATI. – Notes biospéol. 10: 51–61; Paris. 

– (1955 b): Sur le statut systématique de Porcellio succinctus. – Bull. Mus. natn. Hist. nat. 

(2. Ser.) 27: 71–76; Paris. 

– (1955 c): Description de deux nouvelles espèces cavernicoles de Trichoniscus appartenant 

au groupe pygmaeus. – Notes biospéol. 10: 45–49; Paris. 

– (1955 d): Un nouvel example de répartition transadriatique (Trichoniscus Matulici). – 

Revue fr. Ent.22: 59–65; Paris.


– (1955 e): La fauna isopodique cavernicole de l’Afrique du nord (Berbérie). – Notes 


– (1955 f): Mission HENRI COIFFAIT au Liban (1951). 8. Isopodes terrestres. – Archs Zool. 

exp. gén. 91: 455–531; Paris. 

– (1956 a): Les isopodes terrestres des Açores. – Mém. Mus. natn. Hist. nat. (N. S., Ser. A) 

8: 249–264; Paris. 

– (1956 b): Sur un nouveau sous-genre de Metoponorthus et son intérêt biogéographique. 

– Revue fr. Ent.23: 21–30; Paris. 

– (1956 c): Étude d’une espèce polymorphe, Porcellio lamellatus, suivie de considérations 

sur le rôle des glandes tégumentaires chez les isopodes terrestres. – Bull. Soc. zool. Fr. 

81: 359–368; Paris. 

– (1956 d): Un nouveau porcellion provenant de l’Ile d’Ibiza (Baléares) Porcellio pityensis 

n. sp. – Vie Milieu 6: 348–353; Paris. 

– (1956 e): Une nouvelle classification du genre Porcellio. – Bull. Mus. natn. Hist. nat. (2. 

Ser.) 28: 124–128; Paris. 

– (1956 f): Remarques complémentaires et rectifications relatives a Trichorhina boliviana. 

– Bull. Mus. natn. Hist. nat. (2. Ser.) 28: 300–302; Paris. 

– (1956 g): Description d’une nouvelle espèce d’Armadillidium (A. pardoi n. sp.), provenant 

du Rif. – Bull. Mus. natn. Hist. nat. (2. Ser.) 28: 533–536; Paris. 

– (1957 a): Nouvelles espèces et sous-espèces appartenant au genre Oritoniscus. – Bull. 


– (1957 b): Sur une nouvelle espèce de Metoponorthus appartenant au sous-genre Soteriscus. 


– (1957 c): Le genre Atlantidium; sa genèse; son intérêt évolutif et biogéographique. – C. 

r. Séanc. Acad. Sci. Paris 245: 773–777; Paris. 

– (1957 d): Sur une nouvelle espèce d’Halophiloscia: Halophiloscia (Stenophiloscia) bitschi 

n. sp., et son intérêt écologique et biogéographique. – Bull. Mus. natn. Hist. nat. (2. Ser.) 

29: 231–234; Paris. 

– (1957 e): Essai de traduction des phénomènes évolutifs en termes ontogéniques. Application 

au cas des oniscoïdes. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 244: 979–984; Paris. 

– (1957 f): Sur la position systématique de Leptotrichus (Atlantotrichus n. s. g.) leptotrichoides. 


– (1957 g): Sur la constitution et la genèse des différents types d’apophyses génitales chez 

les crustacés isopodes. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 245: 2160–2163; Paris. 

– (1957 h): Nouvelles espèces ou sous-espèces d’oniscoides endogés appartenant a la 

faune française. – Revue fr. Ent. 24: 121–128; Paris. 

– (1958 a): Protracheoniscus pierrei n’est pas un porcellionide quinquétrachéate mais un 

oniscidé. – Bull. Mus. natn. Hist. nat. (2. Ser.) 30: 513–516; Paris. 

– (1958 b): Isopodes récoltés dans les grottes de la Crète par le Docteur K. LINDBERG. – 

Notes biospéol. 12: 81–101; Paris. 

– (1958 c): Sur une nouvelle espèce d’Armadillidium provenant du Rif Marocain, A. djebalensis 

n. sp. – Bull. Mus. natn. Hist. nat. (2. Ser.) 30: 291–293; Paris. 

– (1958 d): Les porcellions du groupe atlantique et du sous-groupe bético-rifain. – Bull. 

Mus. natn. Hist. nat. (2. Ser.) 30: 443–453; Paris. 

– (1958 e): Les isopodes terrestres du Rif. – Tamuda 6: 127–134; Tetuan. 

– (1958 f): Les porcellions catalans et l’origine des porcellions français. – Revue fr. Ent. 25: 

129–148; Paris. 

– (1958 g): „Porcellio pubescens“ appartient au genre Trichorhina. – Bull. Mus. natn. Hist. 

nat. (2. Ser.) 30: 360–364; Paris. 

– (1958 h): Sur un nouvel isopode terrestre provenant des grottes de Sardaigne, Nagurus 

cerrutii n. sp. – Boll. Soc. ent. ital. 88: 148–150; Genoa. 

– (1958 i): Existe-t-il des isopodes pigmentés et aveugles? – Notes biospéol. 13: 67–69; 

Paris. 

– (1959 a): Nouvelles recherches sur les isopodes volvationnels exoantennés et la genèse 

de leur coadaptions. – Bull. biol. Fr. Belg. 93: 121–139; Paris. 

– (1959 b): La faune isopodique cavernicole de la Grèce continantale. – Notes biospéol. 

13: 131–140; Paris. 

– (1959 c): Sur la présence du genre Trichorhina au Maroc et sur les affinités de ce genre. 

– Bull. Mus. natn. Hist. nat. (2. Ser.) 31: 100–104; Paris.


– (1959 d): Les Styloniscidae et les Trichoniscidae de l’Afrique du nord. – Bull. Mus. natn. 

Hist. nat. (2. Ser.) 31: 159–167; Paris. 

– (1959 e): Description d’une nouvelle espèce de l’Afrique occidentale appartenant au 

genre Niambia. – Bull. Mus. natn. Hist. nat. (2. Ser.) 31: 516–519; Paris. 

– (1960 a): Faune de France, vol. 64. Isopodes terrestres (première partie), pp. 1–416; Paris. 

– (1960 b): Les isopodes terrestres de l’Archipel Madérien. – Mém. Mus. natn. Hist. nat. 

(N. S., Ser. A) 22: 1–155; Paris. 

– (1960 c): Sur un nouveau porcellion primitif de la Gran Canaria, Porcellio strinatii n. sp.. 


– (1960 d): Les espèces d’Androniscus appartenant au sous-genre Dentigeroniscus. – Annls 

Spéléol. 15: 553–584; Paris. 

– (1961): Les isopodes terrestres de l’Ile de Minorque. – Archs Zool. exp. gén. (4. Ser.) 99: 

249–265; Paris. 

– (1962 a): Sur la présence d’éléments d’origine sud-américaine dans la faune méditerrannéenne 

terrestre. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 255: 2695–2698; Paris. 

– (1962 b): Faune de France, 66. Isopodes terrestres (deuxième partie), pp. 417–931; Paris. 

– (1963 a):Isopodes terrestres recueillis en Amérique du Sud par CLAUDE DELAMARE DE- 

BOUTTEVILLE. – In: Biologie de l’Amérique australe, vol. 2: 63–100; Paris. 

– (1963 b): Évolution morphologique et répartition géographique de Trichoniscoides arcangelii. 


– (1964 a): La place du genre Actaecia dans le sous-ordre des Oniscoidea. – Crustaceana 

6: 161–174; Leyden. 

– (1964 b): De l’emploi des appareils respiratoires pour l’établissement d’une classification 

rationnelle des isopodes terrestres. – Bull. Soc. zool. Fr. 89: 730–736; Paris. 

– (1964 c): Les isopodes cavernicoles récoltés en Grèce par le Docteur H. HENROT. – 

Annls Spéléol. 19: 729–740; Paris. 

– (1964 d): La position systématique de Leucocyphoniscus gibbosus. – Bull. Mus. natn. 


– (1965 a): La faune isopodique de l’île de Chypre. – Bull. Mus. natn. Hist. nat. (2. Ser.) 

36: 818–830; Paris. 

– (1965 b): Sur l’existence d’oniscoïdes très primitifs menant une vie aquatique et sur le 

polyphylétisme des isopodes terrestres. – Annls Spéléol. 20: 489–518; Paris. 

– (1965 c): Les isopodes terrestres et cavernicoles de la Bulgarie. – Annls Spéléol. 20: 

243–270; Paris. 

– (1965 d): Sur quelques isopodes myrmécophiles recueillis par le Docteur H. HENROT 

dans l’île d’Eubée. – Bull. Mus. natn. Hist. nat. (2. Ser.) 36: 619–621; Paris. 

– (1965 e): Les „Trichoniscidae“ cavernicoles de l’Amérique du Nord. – Annls Spéléol. 

20: 347–389; Paris. 

– (1966 a): Sur un porcellion très primitif, Porcellio laevissimus. – Bull. Mus. natn. Hist. 

nat. (2. Ser.) 38: 174–172; Paris. 

– (1966 b): Observations complémentaires sur Cantabroniscus primitivus. – Annls 


– (1967 a): Les isopodes terrestres et cavernicoles de la Bulgarie (seconde partie). – Annls 


– (1967 b): La répartition de Cantabroniscus primitivus. – Annls Spéléol. 22: 787–795; Paris. 

– (1968 a): The terrestrial Isopoda of the Azores. – Bolm Mus. munic. Funchal 22: 5–29; 

Funchal. 

– (1968 b): Les premiers isopodes terrestres et cavernicoles découverts dans l’Archipel 

Nippon. – Bull. natn. Sci. Mus. 11: 351–362; Tokyo. 

– (1968 c): Isopodes terrestres. – In: Mission zoologique belge aux îles Galapagos et en 

Ecuador (N. et J. LELEUP, 1964–65), vol. 1: 37–168; Brussels. 

– (1968 d): Les isopodes terrestres et cavernicoles de la Corse. – Annls Spéléol. 23: 

355–362; Paris. 

– (1968 e): Description d’un nouveau représentant du genre Cordioniscus suivie de considérations 

sur les voies de migration de certaines lignées d’isopodes terrestres. – Annls 

Spéléol. 23: 621–632; Paris.


– (1969 a): Results of the speleological survey in South Korea 1966. XIII. Isopodes terrestres 

récoltés dans les grottes de Corée. – Bull. natn. Sci. Mus. 12: 157–159; Tokyo. 

– (1969 b): Le mâle de Chaetophiloscia attica. – Biol. gallo-hellen. 2: 13–17; Athens. 

– (1969 c): Les isopodes terrestres de la Sicile. – Atti Accad. gioenia Sci. nat. Catania (Ser. 

7) 1: 1–59; Catania. 

– (1970 a): Les isopodes terrestres et cavernicoles de l’Archipel Nippon (second mémoire). 

– Bull. natn. Sci. Mus. 13: 373–383; Tokyo. 

– (1970 b): Un troisième oniscoïde cavernicole menant une vie aquatique: Mexiconiscus 

laevis. – Annls Spéléol. 25: 161–171; Paris. 

– (1970 c): Les isopodes terrestres des îles Rennell et Bellona. – In: The natural History of 

Rennell Island, British Solomon Islands, vol. 6, nr. 70, pp. 139–153; Copenhagen. 

– (1971 a): Évolution biologique et dérive des continents: une expérience naturelle portant 

sur cent cinquante millions d’années. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 272: 

253–256; Paris. 

– (1971 b): L’origine et l’évolution des Trachelipidae. – Bull. Soc. zool. Fr. 95: 321–328; 

Paris. 

– (1972 a): Les isopodes terrestres et cavernicoles de la Catalogne. – Miscelánea zool. 3: 

1–19; Barcelona. 

– (1972 b): La répartition des oniscoïdes dans le domaine pacifique occidental, et ses rapports 

avec la constitution de cette région du globe. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 

274: 2224–2228; Paris. 

– (1972 c): Isopodes terrestres récoltés en Thaïlande par P. JOLIVET. – Bull. Mus. natn. 


– (1972 d): La répartition des oniscoïdes et la dérive des continents. – C. r. Séanc. Acad. 

Sci. Paris (Ser. D) 275: 2069–2072; Paris. 

– (1972 e): De l’utilisation des données biogéographiques dans la reconstitution des anciens 

visages du globe terrestre. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 274: 38–41; Paris. 

– (1972 f): Sur une espèce d’isopode terrestre et cavernicole recueillie dans une grotte de 

l’Assam. – Annls Spéléol. 27: 543–546; Paris. 

– (1972 g): Les isopodes terrestres de la Colombie. – Stud. neotrop. Fauna Environm. 7: 

147–172; Lisse. 

– (1973 a): Isopodes terrestres de Népal. – Senckenb. biol. 54: 111–128; Frankfurt/Main. 

– (1973 b): Les isopodes terrestres de la Mélanésie – Zool. Verh. 125: 1–160; Leyden. 

– (1973 c): Les isopodes terrestres de l’Australie. Étude systématique et biogéographique. 

– Mém. Mus. natn. Hist. nat. (N. S., Ser. A) 82: 1–171 and one map; Paris. 

– (1973 d): Sur la présence en Catalogne du genre Libanonethes. – Miscelánea zool. 3: 

13–16; Barcelona. 

– (1973 e): L’utilisation des isopodes terrestres en tant qu’indicateurs des modalités de 

l’évolution animale. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 277: 1233–1236; Paris. 

– (1973 f):Les isopodes terrestres et cavernicoles de l’île de Cuba. – In: ORGHIDAN, T. & 

alii (eds.): Résultats des expéditions biospéologiques cubano-roumaines à Cuba, pp. 


– (1974 a): Une nouvelle espèce d’isopode terrestre provenant des Philippines: Sphaerilloides 

philippinensis n. sp. – Crustaceana 26: 61–64; Leyden. 

– (1974 b): Les distributions gondwaniennes. – Année biol 13: 393–395; Paris. 

– (1975 a): Un nouvel oniscoïde provenant du Koweit (Arabie): Koweitoniscus ahmadii n. 

g., n. sp. – Crustaceana 29: 74–78; Leyden. 

– (1975 b): Les isopodes terrestres et le Gondwana. – Mém. Mus. natn. Hist. nat. (N. S., 

Ser. A) 88: 197–133; Paris. 

– (1977 a): Les oniscoïdes de la Nouvelle-Zélande et de l’Archipel Kermadec. – Mém. 

Mus. natn. Hist. nat. (N. S., Ser. A) 102: 1–56; Paris. 

– (1977 b): Les espèces appartenant au genre Amerigoniscus. – Bull. Soc. Hist. nat. 

Toulouse 113: 303–310; Toulouse. 

– (1977 c): La faune terrestre de l’Ile de Sainte-Hélène. 4me partie. 1. Isopodes terrestres. 

– Annls Mus. r. Afr. centr. (Ser. 8vo, Sci. zool.) 220: 385–426 and plate 2; Tervuren. 

– (1980): Les isopodes terrestres recueillis en Turquie orientale et en Irak occidentale par 

le Professeur CURT KOSSWIG. – Bull. Soc. Hist. nat. Toulouse 116: 83–119; Toulouse. 

– (1981): Les isopodes terrestres et cavernicoles de l’île de Cuba (second mémoire). – In:


Résultats des Expéditions biospéologiques cubano-roumaines à Cuba, vol. 3: 35–76; 

Bucharest. 

VANDEL, A., CAILAR, J. DU & BONNET, A. (1946): Les isopodes terrestres du Gard. – Bull. 

Soc. Études Sci. nat. Nîmes 1936–1946: 1–11; Nîmes. 

VANDEL, A. & MATSAKIS, J. (1959 a): Recherche des affinités du genre Atlantidium par l’emploi 

de méthodes biométriques. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 249: 2137–2140; 

Paris. 

– & – (1959 b): Sur l’application des méthodes biométriques à la recherche des affinités 

entre groupes zoologiques. – C. r. Séanc. Acad. Sci. Paris (Ser. D) 248: 336–340; Paris. 

VARONA, A. DE & ARMAS, L. DE (1996): Nuevo género de isópodos terrestres (Isopoda: Armadillidae) 

de Cuba. – Avicennia 4/5: 95–102; Oviedo. 

VERGARA, C. (1967): Observaciones sobre la destruccion de hojas de roble por Porcellio laevis. 

– Boln r. Soc. esp. Hist. nat. (Biol.) 65: 259–269; Madrid. 

VERHOEFF, K. (1896): Ein Beitrag zur Kenntnis der Isopoda terrestria Deutschlands. – Zool. 

Anz. 19: 18–23; Leipzig. 

– (1900): Über paläarktische Isopoden (2. Aufsatz). – Zool. Anz. 23: 117–130; Leipzig. 

– (1901 a): Einige Mitteilungen über Landisopoden. – Berl. ent. Z. 46: 17–20; Berlin. 

– (1901 b): Über paläarktische Isopoden (3. Aufsatz). – Zool. Anz. 24: 33–41; Leipzig. 

– (1901 c): Über paläarktische Isopoden (4. Aufsatz). – Zool. Anz. 24: 66–79; Leipzig. 

– (1901 d): Über paläarktische Isopoden (5. Aufsatz). – Zool. Anz. 24: 135–149; Leipzig. 

– (1901 e): Über paläarktische Isopoden (6. Aufsatz). – Zool. Anz. 24: 257–260; Leipzig. 

– (1901 f): Über paläarktische Isopoden (7. Aufsatz). – Zool. Anz. 24: 403–408, 417–421; 

Leipzig. 

– (1902): Über paläarktische Isopoden. 8. Aufsatz. – Zool. Anz. 25: 241–255; Leipzig. 

– (1907 a): Über paläarktische Isopoden. 9. Aufsatz: Neuer Beitrag zur Kenntnis der Gattung 

Armadillidium. – Zool. Anz. 31: 457–505; Leipzig. 

– (1907 b): Über Isopoden, 10. Aufsatz: Zur Kenntnis der Porcellioniden. – Sber. Ges. naturforsch. 

Fr. Berlin 8: 229–281; Berlin. 

– (1908 a): Neue Isopoden-Gattungen. – Zool. Anz. 33: 520–525; Leipzig. 

– (1908 b): Über Isopoden. Androniscus n. g. – Zool. Anz. 33: 127–148; Leipzig. 

– (1908 c): Über Isopoden. Armadillidium-Arten, mit besonderer Berücksichtigung der 

in Italien und Sizilien einheimischen. – Zool. Anz. 33: 450–462, 484–492; Leipzig. 

– (1908 d): Über Chilopoden und Isopoden aus Tripolis und Barka, gesammelt von Dr. 

BRUNA KLAPTOCZ. – Zool. Jahrb. Syst. 26: 257–284 and plate 20; Jena. 

– (1908 e): Über Isopoden. 12. Aufsatz. – Arch. Naturgesch. 74: 163–198; Berlin. 

– (1908 f): Über Isopoden: 15. Aufsatz. – Arch. Biontol. 2: 335–387; Berlin. 

– (1909): Über einige Isopoden und Myriapoden aus Montenegro und Albanien. – Wiss. 

Mitt. Bosn. Herzeg. 11: 716–722; Vienna. 

– (1910): Über Isopoden, 16. Aufsatz, Armadillidium und Porcellio an der Riviera. – Jahresh. 

Ver. vaterl. Naturk. Württ. 66: 115–143; Stuttgart. 

– (1914 a): Über Mesoniscus. – Zool. Anz. 44: 425–427; Leipzig. 

– (1914 b): Zur Kenntnis der Gattung Mesoniscus. – Zool. Jahrb. Syst. 37: 493–508; Jena. 

– (1917 a): Über mediterrane Oniscoideen, namentlich Porcellioniden. – Jahresh. Ver. vaterl. 

Naturk. Württ. 73: 144–173; Stuttgart. 

– (1917 b): Zur Kenntnis der Atmung und der Atmungsorgane der Isopoda-Oniscoidea. 

– Biol. Zentralbl. 37: 113–127; Leipzig. 

– (1917 c): Über einige niederösterreichische Diplopoden und Isopoden. – Verh. zool.bot. 

Ges. Wien 67: 220–223; Vienna. 

– (1917 d): Germania zoogeographica: Die Verbreitung der Isopoda terrestria im Vergleich 

mit derjenigen der Diplopoden. – Zool. Anz. 48: 347–376; Leipzig. 

– (1917 e): Zur Kenntnis der Gattungen Trichoniscus und Mesoniscus. – Zool. Anz. 49: 


– (1917 f): Zur Kenntnis der Entwicklung der Trachealsysteme und der Untergattungen 

von Porcellio und Tracheoniscus. – Sber. Ges. naturforsch. Fr. Berlin 1917: 195–223; 

Berlin. 

– (1918 a): Zur Kenntnis der Ligiiden, Porcellioniden und Onisciden. – Arch. Naturgesch. 

82 A: 108–169; Berlin. 

– (1918 b): Ueber augenlose Armadillidien und kritische Prüfung der Familie Armadillidiidae. 

– Arch. Naturgesch. 83 A: 160–170; Berlin.


– (1919 a): Über die Larven, das Marsupium und die Bruten der Oniscoidea. – Arch. Naturgesch. 

83 A: 1–54; Berlin. 

– (1919 b). Zur Kenntnis der Gattungen Porcellium und Armadillidium in Deutschland. 

– Arch. Naturgesch. 83 A: 1–37; Berlin. 

– (1920 a): Über die Atmung der Landasseln. – Z. wiss. Zool. 118: 365–447; Leipzig. 

– (1920 b): Zur Kenntnis der Larven, des Brutsackes und der Bruten der Oniscoidea. – 


– (1920 c): Zur geographisch-biologischen Beurteilung der Land-Isopoden. – Zool. Anz. 

52: 20–31; Leipzig. 

– (1923): Zur Kenntnis der Landasseln Palästinas. – Arch. Naturgesch. 89 A: 206–231; 

Berlin. 

– (1926 a): Isopoda terrestria von Neu-Caledonien und den Loyalty-Inseln. – In: SARA- 

SIN & ROUX (eds.): Nova Caledonia, Zoologie, vol. 4: 243–366. 

– (1926 b): Über Isopoden der Balkanhalbinsel, gesammelt von Herrn Dr. BURESCH. – 

Mitt. bulg. ent. Ges. Sofia 3: 135–158; Sofia. 

– (1927 a): Über einige zoogeographisch bedeutsame Isopoden-Arten. – Zool. Anz. 73: 

323–333; Leipzig. 

– (1927 b): Über einige südosteuropäischen Trichonisciden. – Zool. Anz. 70: 200–223; 

Leipzig. 

– (1927 c): Illyrionethes n. g. eine cavernicole Trichonisciden-Gattung. – Zool. Anz. 72: 

268–274; Leipzig. 

– (1928 a): Über einige Isopoden der Zoologischen Staatssammlung in München. – Zool. 

Anz. 76: 25–36, 113–123; Leipzig. 

– (1928 b): Isopoden aus Formosa. – Mitt. zool. Mus. Berlin 14: 200–226; Berlin. 

– (1928 c): Über Stenonisciden. – Zool. Anz. 79: 58–64; Leipzig. 

– (1928 d): Über alpenländische und italienische Isopoden. – Zool. Jahrb. Syst. 56: 

93–172; Jena. 

– (1929 a): Ueber Isopoden der Balkanhalbinsel, gesammelt von Herrn Dr. I. BURESCH. 

II. Teil. – Mitt. naturwiss. Inst. Sofia 2: 129–139; Sofia. 

– (1929 b): Eine neue Diplopoden- und eine neue Isopoden-Gattung aus dem Labyrinth 

Kretas. – Mitt. Höhlen- Karstforsch. 4: 113–123; Berlin. 

– (1929 c): Arthropoden aus südostalpinen Höhlen, gesammelt von KARL STRASSER. – 

Mitt. Höhlen- Karstforsch. 4: 14–35; Berlin. 

– (1929 d): Arthropoden aus südostalpinen Höhlen, gesammelt von Herrn KARL STRAS- 

SER. 2. Aufsatz. III. Isopoda. – Mitt. Höhlen- Karstforsch. 4: 51–54; Berlin. 

– (1930 a): Über einige von Herrn Dr. C. R. BOETTGER in Italien gesammelte Diplopoden, 

Chilopoden und Isopoden. – Mitt. zool. Mus. Berlin 16: 566–573; Berlin. 

– (1930 b): Über einige norditalienische Isopoden und einen neuen Typus der Volvation. 


– (1930 c): Zur Kenntnis osteuropäischer Isopoden. – Zool. Jahrb. Syst. 59: 1–64 and plates 

1–3; Jena. 

– (1930 d): Über Isopoden aus Turkestan. – Zool. Anz. 91: 101–125; Leipzig. 

– (1931 a): Zur Kenntnis alpenländischer und mediterraner Isopoda terrestria. – Zool. 

Jahrb. Syst. 62: 1–52 and plates 1–2; Jena. 

– (1931 b): Über Isopoda terrestria aus Italien. – Zool. Jahrb. Syst. 60: 489–572 and plates 

6–8; Jena. 

– (1931 c): Vergleichend geographisch-ökologische Untersuchungen über die Isopoda 

terrestria von Deutschland, den Alpenländern und anschließenden Mediterrangebieten. 

– Z. Morph. Ökol. Tiere 22: 231–268; Berlin. 

– (1931 d): Zwei neue Illyrionethes-Arten aus Dalmatien. – Senckenbergiana 13: 122–134; 


– (1932 a): Cavernicole Oniscoideen. – Mitt. Höhlen- Karstforsch. 1932: 12–24; Berlin. 

– (1932 b): Vergleichend geographisch-ökologische Untersuchungen über die Isopoda 

terrestria, namentlich der italienischen Westalpen. – Z. Morph. Ökol. Tiere 24: 

359–393; Berlin. 

– (1933 a): Arthropoden aus südostalpinen Höhlen, gesammelt von KARL STRASSER, 

Triest. 7. Aufsatz. – Mitt. Höhlen- Karstforsch. 4: 1–21; Berlin. 

– (1933 b): Zur Systematik, Geographie und Ökologie der Isopoda terrestria Italiens und 

über einige Balkan-Isopoden. – Zool. Jahrb. Syst. 65: 1–64; Jena.


– (1933 c): Neue Isopoda terrestria aus Mexiko und dem Mediterrangebiet. – Zool. Anz. 

103: 97–119; Leipzig. 

– (1934 a): Über Isopoden aus dem Gardasee-Gebiet. – Zool. Anz. 108: 242–250; Leipzig. 

– (1934 b): Über westfälische Isopoden. – Zool. Anz. 106: 272–276; Leipzig. 

– (1934 c): Das geographische Auftreten der Isopoden in den Alpenländern im Vergleich 

zu dem der Diplopoden. – Forschn Fortschr. 10: 133; Berlin. 

– (1936 a): Ueber Isopoden der Balkanhalbinsel, gesammelt von Herrn Dr. I. BURESCH. 

III. Teil. – Mitt. koenigl. naturwiss. Inst. Sofia 9: 1–27; Sofia. 

– (1936 b): Studien über Isopoda terrestria. – Mitt. zool Mus Berlin 21: 79–163; Berlin. 

– (1936 c): Über die ersten durch Domestikation erzielten Isopoden-Rassen. – Z. Morph. 

Ökol. Tiere 31: 673–681; Berlin. 

– (1936 d): Über einige Myriapoden und einen Isopoden aus mediterranen Höhlen. – 


– (1936 e): Ueber Land-Isopoden aus Ladak. – Mem. Conn. Acad. Arts Sci. 10: 187–190; 

New Haven. 

– (1936 f): Ueber einige Isopoda aus Süd-Indien. – Rec. Indian Mus. 38: 97–102 and plate 

4; Calcutta. 

– (1936 g): Ueber einige Myriapoden und Isopoden aus Dekan, gesammelt von Herrn S. 

JONES, Madras. – Rec. Indian Mus. 38: 503–512 and plates 15–16; Calcutta. 

– (1937 a): Über einige Porcellio-Arten aus Marokko. – Zool. Anz. 119: 302–307; Leipzig. 

– (1937 b): Über eine angeblich neue Isopoden-Familie. – Zool. Anz. 118: 173–176; Leipzig. 

– (1937 c): Über einige neue und bekannte Isopoda terrestria. – Sber. Ges. naturforsch. Fr. 


– (1937 d): Die Isopoden des alemannischen Gaues und ihre geographisch-ökologische 

Bedeutung. – Beitr. naturk. Forsch. SüdwDtl. 2: 22–39 and plate 2; Karlsruhe. 

– (1938 a): Über Land-Isopoden aus Marokko. II. Teil. – Zool. Anz. 124: 61–67; Leipzig. 

– (1938 b): Weltstellung der Isopoda terrestria, neue Familien derselben und neues System. 

– Zool. Jahrb. Syst. 71: 253–264; Jena. 

– (1938 c): Morphologisch-geographisch-ökologischer Beitrag zur Kenntnis der Isopoda 

terrestria von Oberwallis und Insubrien. – Arch. Naturgesch. (N. F.) 7: 317–363; Berlin. 

– (1938 d): Zur Kenntnis der Gattung Porcellio und über Isopoda Oniscoidea der Insel 

Cherso. – Arch. Naturgesch. (N. F.) 7: 97–136; Berlin. 

– (1938 e): Über einige polynesische Oniscoideen von Prof. SIXTEN BOCKs Pazifik-Expedition 

1917–1918. – Ark. Zool. 30 A: 1–14; Uppsala. 

– (1939 a): Über einige balkanische Isopoda terrestria. – Stud. Geb. allg. Karstforsch. 

wiss. Höhlenk. (biol. Ser.) 6: 1–10; Brünn. 

– (1939 b): Land-Isopoden aus Spessart, Odenwald und Hardt. – Zool. Anz. 128: 35–47; 

Leipzig. 

– (1939 c): Zur Kenntnis der Schöbliiden. – Zool. Anz. 125: 135–137; Leipzig. 

– (1939 d): Von Dr. G. H. SCHWABE in Chile gesammelte Isopoda terrestria, Diplopoda 

aund Chilopoda. – Arch. Naturgesch. (N. F.) 8: 301–324; Berlin. 

– (1939 e): Diplopoden, Chilopoden uand Oniscoideen, hauptsächlich aus süditalienischen 

Höhlen. – Zool. Jahrb. Syst. 72: 203–224; Jena. 

– (1939 f): Die Isopoda terrestria Kärntens in ihren Beziehungen zu den Nachbarländern 

und in ihrer Abhängigkeit von den Vorzeiten. – Abh. preuss. Akad. Wiss. (math.-nat. 

Kl.) 15: 1–45; Berlin. 

– (1940 a): Über einige Diplopoden, Chilo- und Isopoden der Insel Ischia. – Zool. Anz. 

131: 271–287; Leipzig. 

– (1940 b): Strouhalius n. g., eine nordwestbalkanische cavernicole Isopodengattung. – 


– (1940 c): Über Beziehungen einiger Höhlen-Trichonisciden. – Mitt. Höhlen- Karstforsch. 

1940: 79–84; Berlin. 

– (1940 d)): Der geographische Charakter der Landisopodenfauna italienischer Mittelmeerinseln 

und über die Landisopoden der Insel Ischia. – Z. Morph. Ökol. Tiere 37: 

105–125; Berlin. 

– (1940 e): Über die Doppelhäuting der Landisopoden. – Z. Morph. Ökol. Tiere 37: 

126–143; Berlin.


– (1940 f): Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen 

Provinzen Chinas, unter Leitung von Dr. SVEN HEDIN und Prof. SÜ PING- 

CHANG. Landisopoden gesammelt von Dr. DAVID HUMMEL. – Ark. Zool. 31 A: 1–10; 

Uppsala. 

– (1941 a): Über Land-Isopoden aus der Türkei.– Istanb. Üniv. Fen Fak. Mecm. (Ser. B) 

4: 223–276. 

– (1941 b): Über eine neue südamerikanische Gattung der Isopoda terrestria. – Zool. 

Anz. 134: 169–173; Leipzig. 

– (1941 c): Untersuchungen über die Fauna des Hologäum: Chilopoden, Diplopoden 

und Landisopoden. – Zool. Anz. 135: 35–41; Leipzig. 

– (1941 d): Zur Kenntnis südamerikanischer Oniscoideen. – Zool. Anz. 133: 114–126; 

Leipzig. 

– (1941 e): Grundlagen zur Beurteilung der geographischen Verbreitung der Landisopoden 

und über deren Fauna im westnordischen Gau. – Z. Morph. Ökol. Tiere 37: 

455–490; Berlin. 

– (1941 f): Land-Isopoden. – In: TITSCHAK, E. (ed.): Beiträge zur Fauna Perus, vol. 1: 

73–80; Jena. 

– (1942 a): Sphaeromiden-Studien und Buchnerillo n. g. – Z. Morph. Ökol. Tiere 39: 

153–175; Berlin. 

– (1942 b): Land-Isopoden aus Fernando Po. – Zool. Anz. 137: 84–98; Leipzig. 

– (1942 c): Zur Kenntnis der Armadilliden und über Detonella.- Zool. Anz. 138: 

162–174; Leipzig. 

– (1942 d): Äthiopische Isopoda terrestria der Hamburger Zoologischen Museums. – 

Zool. Anz. 140: 1–26, 61–87, 149–163; Leipzig. 

– (1942 e): Land-Isopoden in einem Thermalschacht bei Badenweiler. – Zool. Anz. 139: 


– (1942 f): Grundlagen zur Beurteilung der geographischen Verbreitung von Bodentieren, 

beleuchtet an dem Beispiel des Isopoden Armadillidium vulgare. – Zoogeographica 

4: 93–110; Jena. 

– (1942 g): Landisopoden der Insel Ischia, systematisch, morphologisch, phänologisch, 

ökologisch, geographisch beurteilt. – Z. Morph. Ökol. Tiere 38: 435–482; Berlin. 

– (1942 h): Isopoda terrestria des norischen Gaues und über Grundsätze bezüglich der 

Auffassung zoogeographischer Gebiete. – Arch. Naturgesch. (N. F.) 11: 198–219; Berlin. 

– (1943 a): Cavernicole Diplopoden, Chilopoden und Land-Isopoden der Insel Capri. – 

Zeitschrift für Karst- und Höhlenkunde 1943: 153–165; Berlin. 

– (1943 b): Über Land-Isopoden aus der Türkei. 2. Aufsatz. – Istanb. Üniv. Fen Fak. 

Mecm. (Ser. B) 8: 1–29. 

– (1944): Isopoden der Insel Capri und der Sorrentinischen Halbinsel nebst ihren geographischen 

Beziehungen. – Zool. Anz. 144: 90–115; Leipzig. 

– (1946): Über Land-Isopoden der Seychellen und aus Burma. – Ark. Zool. 37 A: 1–18; 

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– (1949 a): Über Land-Isopoden aus der Türkei. III. – Istanb. Üniv. Fen Fak. Mecm. (Ser. 

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– (1949 b): Tylos, eine terrestrisch-maritime Rückwanderer-Gattung der Isopoden. – 

Arch. Hydrobiol. 42: 329–340 and plates 19–21; Stuttgart. 

– (1951 a): Diplopoda, Chilopoda und Isopoda terrestria vom Mt. Soratte in Latium. – 

Zool. Jahrb. Syst. 80: 205–255; Jena. 

– (1951 b): Land-Isopoden aus Südamerika. – In: Further zoological Results of the Swedish 

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Authors’ address: 

Dr. HELMUT SCHMALFUSS, Dr. KARIN WOLF-SCHWENNINGER, Staatliches Museum für 

Naturkunde (Museum am Löwentor), Rosenstein 1, D-70191 Stuttgart, Germany; 

e-mail: schmalfuss.smns@naturkundemuseum-bw.de. 

ISSN 0341-0145 

Schriftleitung: Dr. Wolfgang Seeger, Rosenstein 1, D-70191 Stuttgart 

Gesamtherstellung: Gulde-Druck GmbH, D-72072 Tübingen

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