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<strong>Bougainvillea</strong><br />

Kent D. Kobayashi, James McConnell l , <strong>and</strong> John Griffis<br />

Department <strong>of</strong> <strong>Tropical</strong> Plant <strong>and</strong> Soil Sciences, 1 University <strong>of</strong> Guam<br />

<strong>Bougainvillea</strong> species<br />

The genus <strong>Bougainvillea</strong>, in the<br />

Nyctinaginacea (Four-o’clock)<br />

family <strong>of</strong> plants, has 14 species,<br />

with three that are horticulturally<br />

important: B. spectabilis Willdenow,<br />

B. glabra Choisy, <strong>and</strong> B.<br />

peruviana Humboldt <strong>and</strong> Bonpl<strong>and</strong>.<br />

Many crosses among the<br />

various species have produced<br />

new hybrid species <strong>and</strong> important<br />

horticultural cultivars. <strong>Bougainvillea</strong><br />

is native to South America.<br />

The name comes from Louis<br />

Antoine de Bougainville, a French<br />

navigator <strong>and</strong> military comm<strong>and</strong>er<br />

who was the first European<br />

to take note <strong>of</strong> the plant, in Brazil, in 1768.<br />

B. spectabilis<br />

This was the first member <strong>of</strong> the genus to be identified<br />

from Brazil in 1798. German botanist Carl Ludwig<br />

Willdenow is credited with this identification. This species<br />

is noted for its hairy leaves <strong>and</strong> stems. The leaves<br />

are large <strong>and</strong> ovate, with rippling along the edges <strong>and</strong><br />

hairs on the underside. The bracts are red, dark pink, or<br />

purple, while the small flowers are cream colored. Its<br />

thorns are large <strong>and</strong> may be curved. The growth habit is<br />

dense, <strong>and</strong> the colorful bracts appear up <strong>and</strong> down the<br />

branches. The bloom cycle is seasonal, with plants<br />

blooming after the dry season or after a cool spell, which<br />

may trigger plants to bloom. B. spectabilis is Guam’s<br />

territorial flower.<br />

<strong>Bougainvillea</strong>, paper flower;<br />

pukanawila, kepalö, kiapolö, pua kepalö<br />

(Hawaiian);<br />

puti tai nobiu (Chamorro)<br />

Ornamentals <strong>and</strong> Flowers<br />

Oct. 2007<br />

OF-38<br />

B. glabra<br />

This climbing, evergreen member<br />

<strong>of</strong> the genus, also from Brazil, was<br />

first identified by Swiss botanist<br />

Jacques Denys Choisy in 1850.<br />

The elliptical leaves are green or<br />

variegated, with a glossy sheen.<br />

They are glabrous (smooth, hairless),<br />

but you may find some<br />

puberulence (presence <strong>of</strong> small<br />

hairs). Its bracts come in many<br />

sizes <strong>and</strong> shapes. Typically they are<br />

triangular <strong>and</strong> purple or mauve,<br />

although white bracts are also<br />

common. The bracts tend to appear<br />

along the branches <strong>and</strong> at the tips.<br />

The flowers are white to cream<br />

colored. The thorns are small <strong>and</strong> curved at the tips. The<br />

growth habit is spreading, <strong>and</strong> the green-leaf types are<br />

fast growing. This species blooms several times a year. B.<br />

glabra <strong>and</strong> B. spectabilis are alike in general appearance,<br />

the main differences being the bloom cycle, <strong>and</strong> B. glabra<br />

being hairless whereas B. spectabilis is hairy.<br />

B. glabra’s score under the Hawai‘i-Pacific Weed Risk<br />

Assessment program is –1, not likely to be a pest, <strong>and</strong><br />

its designation is L, meaning that it is “not currently<br />

recognized as invasive in Hawai‘i, <strong>and</strong> not likely to have<br />

major ecological or economic impacts on other Pacific<br />

Isl<strong>and</strong>s based on the HP-WRA screening process.”<br />

B. peruviana<br />

It is thought that this climbing, evergreen member from<br />

Peru was first identified by German naturalist <strong>and</strong> ex-<br />

Published by the <strong>College</strong> <strong>of</strong> <strong>Tropical</strong> <strong>Agriculture</strong> <strong>and</strong> <strong>Human</strong> Resources (CTAHR) <strong>and</strong> issued in furtherance <strong>of</strong> Cooperative Extension work, Acts <strong>of</strong> May 8 <strong>and</strong> June 30, 1914, in cooperation<br />

with the U.S. Department <strong>of</strong> <strong>Agriculture</strong>. Andrew G. Hashimoto, Director/Dean, Cooperative Extension Service/CTAHR, University <strong>of</strong> Hawai‘i at Mänoa, Honolulu, Hawai‘i 96822.<br />

An equal opportunity/affirmative action institution providing programs <strong>and</strong> services to the people <strong>of</strong> Hawai‘i without regard to race, sex, age, religion, color, national origin, ancestry, disability,<br />

marital status, arrest <strong>and</strong> court record, sexual orientation, or status as a covered veteran. CTAHR publications can be found on the Web site .


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

plorer Alex<strong>and</strong>er von Humboldt in 1810. This species is<br />

noted for its green bark. The long, thin leaves are strongly<br />

ovate <strong>and</strong> glabrous. The rounded bracts are magenta to<br />

pink <strong>and</strong> may have some distortion or wrinkling, <strong>and</strong><br />

the flowers are yellow. The thorns are short <strong>and</strong> straight.<br />

The growth habit is lanky. This species is not as vigorous<br />

as some <strong>of</strong> the hybrids <strong>and</strong> <strong>of</strong>ten requires pruning to<br />

promote substantial branching. The plants may bloom<br />

several times a year if subjected to dry periods between<br />

flushes. B. peruviana is the most stable species <strong>of</strong> the<br />

three described here, as there is very little variation in<br />

the general shape <strong>of</strong> bracts <strong>and</strong> leaves.<br />

<strong>Bougainvillea</strong> hybrids<br />

Among bougainvillea’s horticultural hybrids, B. glabra<br />

x B. peruviana is by far the most common. This cross<br />

has its own hybrid name: <strong>Bougainvillea</strong> x buttiana. The<br />

original discovery <strong>of</strong> this hybrid was made by Mrs. R.<br />

Butt in a garden in Trinidad, hence the name. The leaves<br />

are large <strong>and</strong> ovate or heart-shaped with slight hairiness<br />

on both the upper <strong>and</strong> lower sides. The bracts are usually<br />

rounded, red or dark pink, <strong>and</strong> the thorns are straight<br />

<strong>and</strong> short. The small flowers are cream colored with pink<br />

tones, although the floral tube may be the same color as<br />

the bract. The growth habit is open <strong>and</strong> requires pruning<br />

to promote a bushy appearance. B. x buttiana hybrids<br />

generally bloom several times a year.<br />

<strong>Bougainvillea</strong> x spectoperuviana is another common<br />

hybrid. The leaves are large, dark green, <strong>and</strong> ovate, <strong>and</strong><br />

these hybrids are usually hairless. The bracts are coppery<br />

red in the juvenile stage, turning to various shades<br />

<strong>of</strong> magenta or pink as they age. The thorns are straight,<br />

<strong>and</strong> the flowers are cream colored. The growth habit is<br />

large <strong>and</strong> spreading, <strong>and</strong> they generally bloom several<br />

times a year.<br />

<strong>Bougainvillea</strong> x spectoglabra is the last <strong>of</strong> the common<br />

hybrid bougainvilleas. The leaves are small <strong>and</strong><br />

dark green, with mauve or purple bracts. The thorns are<br />

numerous <strong>and</strong> curved, <strong>and</strong> the small flowers are almost<br />

white. The growth habit is well branched <strong>and</strong> thick, <strong>and</strong><br />

blooming is generally several times a year.<br />

Characteristics<br />

<strong>Bougainvillea</strong> is a tropical <strong>and</strong> subtropical woody, evergreen,<br />

shrubby vine. Typically multi-trunked or with<br />

clumping stems, it has a spreading, round plant habit<br />

with a height <strong>and</strong> spread <strong>of</strong> up to 20 feet. It climbs by<br />

sending out slender arching canes armed with stiff curved<br />

2<br />

The true, perfect flower is small, tubular, commonly white<br />

or yellow, <strong>and</strong> surrounded by showy, vibrantly colorful<br />

petaloid bracts.<br />

thorns. As they age, the stems turn from mid-green to<br />

dull green-brown. <strong>Bougainvillea</strong> is deciduous when<br />

grown in areas with a long dry season.<br />

Numerous cultivars are available, with a striking array<br />

<strong>of</strong> colors. Their colorful “flowers” are really bracts,<br />

or modified leaves, 1 ⁄2–2-inch long structures to which<br />

the true flowers are attached at the mid-rib. New growth<br />

is required for inflorescence production, which occurs<br />

on short stem axes borne laterally in the axils <strong>of</strong> leaves.<br />

From their initial emergence to full maturity, the bracts<br />

<strong>of</strong> many cultivars change color. Bracts may retain their<br />

color for several months after the flowers have finished,<br />

gradually fading to resemble the color <strong>and</strong> texture <strong>of</strong><br />

paper.<br />

Leaves are simple <strong>and</strong> alternate, with an undulate leaf<br />

margin. The leaf blade is 2–4 inches long, with much<br />

variation in shape: globular, elliptical, obivate, ovate,<br />

or cordate. Leaves are mid- to deep green, although some<br />

cultivars have variegated foliage.<br />

The true, perfect flowers are small, tubular, <strong>and</strong> surrounded<br />

by showy, colorful petaloid bracts. The vibrant<br />

colors <strong>of</strong> bougainvillea are not from the inconspicuous,<br />

commonly white or yellow, flowers but from the bracts<br />

that surround each flower.<br />

The fruit is an elongated achene less than 1 ⁄2 inch long.<br />

It is rather inconspicuous, not showy, <strong>and</strong> has a dry, hard<br />

fruit cover.


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

<strong>Bougainvillea</strong> in an urban l<strong>and</strong>scape Trained as a “st<strong>and</strong>ard”<br />

Location <strong>and</strong> l<strong>and</strong>scape uses<br />

Location<br />

Bouganvilleas grow best in full sun. High light intensity<br />

is required for good flowering. Low light <strong>and</strong> shady<br />

areas are not suitable, as the plants will drop their bracts.<br />

<strong>Bougainvillea</strong> does best at elevations from 10 to 2500<br />

feet.<br />

<strong>Bougainvillea</strong> can tolerate hot dry locations, with temperatures<br />

over 100°F. It does well in locations with a<br />

minimum <strong>of</strong> 65°F at night <strong>and</strong> 75–95°F during the day.<br />

B. glabra can tolerate slightly cooler conditions (58–<br />

64°F) than B. spectabilis (64–68°F). <strong>Bougainvillea</strong> does<br />

best with at least 25 inches <strong>of</strong> rainfall per year.<br />

<strong>Bougainvillea</strong> grows well in rich, well drained, acidic<br />

(pH 5.5–6.0) soil. It does not thrive in soil that is constantly<br />

wet. Proper soil pH is essential because it affects<br />

the availability <strong>of</strong> mineral elements. A soil pH above<br />

6.0 increases the possibility <strong>of</strong> micronutrient deficiencies,<br />

particularly iron. <strong>Bougainvillea</strong> is drought tolerant,<br />

salt tolerant, <strong>and</strong> wind resistant.<br />

<strong>Bougainvillea</strong> is very susceptible to girdling during a<br />

storm. The bark will rub <strong>of</strong>f at ground level when stems<br />

whip in high-speed winds. The plant is slow to recover<br />

from this, compared to other shrubs. If girdling is severe,<br />

the entire plant will wilt a few days after a storm.<br />

It should not be planted in extremely windy, unprotected<br />

areas.<br />

L<strong>and</strong>scape uses<br />

<strong>Bougainvillea</strong>’s growth habit <strong>and</strong> beautiful showy bracts<br />

make it a popular plant for l<strong>and</strong>scapes. It is used in mass<br />

plantings, as shrubs or bushes, <strong>and</strong> as ground cover on<br />

banks. <strong>Bougainvillea</strong> provides hedges, barriers, <strong>and</strong><br />

slope coverings. For large, difficult-to-maintain areas,<br />

bougainvillea is an excellent ground cover. It can cover<br />

a whole hillside <strong>and</strong> will choke out weed growth. Dwarf<br />

cultivars make colorful ground covers.<br />

<strong>Bougainvillea</strong> can be trained as a “st<strong>and</strong>ard,” a small<br />

flowering tree with a single trunk (photo above), over<br />

arbors, into espaliers, onto walls, or to cascade down a<br />

slope. <strong>Bougainvillea</strong> is used as an accent plant, a specimen<br />

plant, in hanging baskets, in containers, <strong>and</strong> for<br />

bonsai.<br />

Give plants enough room to grow to maturity without<br />

being crowded. Most cultivars do well 6–9 feet apart.<br />

Smaller cultivars can be planted closer together, at 3–5foot<br />

spacing. Bouganvillea should not be planted within<br />

4 feet <strong>of</strong> walkways, as the thorns could catch unsuspecting<br />

passersby.<br />

In Hawai‘i, flowering is heaviest from September to<br />

April. Flowering is promoted by short daylengths. In<br />

addition, cultural practices are important to ensure satisfactory<br />

floweringæavoid overwatering, overfertilizing<br />

with nitrogen, putting plants in heavy shade, <strong>and</strong> pruning<br />

too frequently. The flowering peak in Guam occurs<br />

during the dry months, March through May.<br />

3


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Culture<br />

Watering<br />

<strong>Bougainvillea</strong> tolerates drying <strong>and</strong> if possible irrigation<br />

should be adjusted to be a little on the dry side. They are<br />

sensitive to overwatering but should not be allowed to<br />

completely dry out.<br />

Fertilizer<br />

For best results, use organic fertilizer amendments or<br />

controlled-release fertilizers to moderate nitrogen release.<br />

At planting, amend the soil with a fertilizer high<br />

in phosphate. For long-term culture, topdressing with a<br />

controlled-release fertilizer is needed. Do not overfertilize.<br />

Too much fertilizer will promote vegetative growth<br />

<strong>and</strong> inhibit blooming.<br />

<strong>Bougainvillea</strong> needs regular fertilizing with formulations<br />

having NPK ratios <strong>of</strong> 1:1:1 or 2:1:2. Applications<br />

<strong>of</strong> soluble minor elements help prevent leaf chlorosis.<br />

Micronutrient applications can be half the recommended<br />

rate, twice a year.<br />

Pruning<br />

<strong>Bougainvillea</strong> responds well to pruning. Unless they are<br />

pruned regularly, bougainvillea grows into a tangled<br />

mass <strong>of</strong> old <strong>and</strong> new growth, <strong>and</strong> overcrowding <strong>of</strong>ten<br />

leads to pests <strong>and</strong> diseases. To prevent overcrowding,<br />

cut out any unneeded shoots. Cut all lateral shoots back<br />

to within two or three buds <strong>of</strong> the main stems. These<br />

will bear the new flowers <strong>and</strong> bracts.<br />

Regular pruning is necessary to shape the plant <strong>and</strong><br />

direct its growth because the shoots <strong>of</strong>ten grow vigorously.<br />

Flowers are borne on new growth, so pinching<br />

back <strong>and</strong> pruning is necessary to induce new growth.<br />

Pruning should be done after flowering has finished, as<br />

this encourages the new growth on which the next flush<br />

<strong>of</strong> flowers will occur.<br />

To reduce the size <strong>of</strong> plants, cut them back by about a<br />

third, removing all spindly <strong>and</strong> twiggy growth. Prune<br />

suckers from the plant’s base to encourage topgrowth.<br />

Dead wood should be removed as it appears. The long<br />

shoots can be trained in various shapes <strong>and</strong> heights—<br />

espalliers, arbors, twisted or braided trunks, or even<br />

large, fanciful animals.<br />

4<br />

Controlling timing <strong>of</strong> flowering<br />

Environment<br />

<strong>Bougainvillea</strong> will flower sooner <strong>and</strong> more pr<strong>of</strong>usely<br />

with high light intensities, moderate temperatures, <strong>and</strong><br />

longer nights. Short daylengths enhance flowering: 8–<br />

11-hour daylengths with high light intensity <strong>and</strong> temperatures<br />

above 58–64°F. Heavy shade inhibits flowering.<br />

Drought stress can stimulate flowering even under<br />

long daylengths. Growers frequently allow plants to dry<br />

just to the point <strong>of</strong> wilting to induce flowering. Use care,<br />

because excessive drying can cause leaf drop <strong>and</strong> dormancy.<br />

Cultural practices<br />

Pruning plants too frequently will inhibit flowering. Too<br />

much fertilizer will add growth but inhibit blooming.<br />

Nitrogen <strong>and</strong> phosphate are critical to flowering, but do<br />

not overfertilize. Keep soil moisture on the dry side.<br />

Pot culture<br />

Light<br />

Plants should be placed in bright light or near a window<br />

with at least 4000 foot-c<strong>and</strong>les <strong>of</strong> light. In low-light interior<br />

environments, their high light requirement leads<br />

to leaf drop. Bracts will be a lighter color than those<br />

grown in full sunshine.<br />

As a point <strong>of</strong> reference, the amount <strong>of</strong> light at noon<br />

on a clear summer day is approximately 10,000 footc<strong>and</strong>les.<br />

On an overcast winter day, it may be as low as<br />

500 foot-c<strong>and</strong>les. Indoors, on a clear summer afternoon,<br />

the direct sun entering a window may be 4000–8000<br />

foot-c<strong>and</strong>les, but the level will be only about 600 footc<strong>and</strong>les<br />

in the shade to the sides <strong>of</strong> the window. The<br />

natural light indoors on the shady side <strong>of</strong> a house may<br />

be 150–250 foot-c<strong>and</strong>les, depending on the amount <strong>of</strong><br />

window area <strong>and</strong> the presence <strong>of</strong> eaves, window blinds,<br />

or curtains.<br />

Media<br />

Any well drained potting medium is suitable for growing<br />

bougainvillea. A 1:1 (by volume) peat:perlite medium<br />

is suitable. If more weight is needed to stabilize<br />

the pot, man-made s<strong>and</strong> or soil may be added. The medium<br />

needs to be well drained to prevent a poor root<br />

system <strong>and</strong> reduced flowering. Avoid media with high<br />

peat content <strong>and</strong> water-retention levels. These types <strong>of</strong>


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

media retain too much water <strong>and</strong> will contribute to root<br />

rot. Media pH should be 5.5–6.0.<br />

Watering<br />

Plants should be watered when the medium surface becomes<br />

dry. Close attention is needed because it is easy<br />

for plants to use up the available moisture in the pots.<br />

The amount <strong>of</strong> water needed depends on the medium<br />

type, environmental conditions, size <strong>of</strong> the plant, <strong>and</strong><br />

pot size. Watering well but less frequently is better than<br />

frequent, light waterings.<br />

Fertilizing<br />

In pot cultivation, an evenly balanced controlled-release<br />

fertilizer (e.g., 8-8-8 or 10-10-10) can be applied every<br />

three months. Water-soluble fertilizer formulations can<br />

be applied weekly or bi-weekly at half-strength to provide<br />

a low nutrient concentration.<br />

Pruning<br />

Frequent pruning helps maintain the desired plant size<br />

<strong>and</strong> shape <strong>and</strong> keeps new growth s<strong>of</strong>t <strong>and</strong> thorns to a<br />

minimum. Frequent cutting back promotes constant<br />

flushing <strong>of</strong> new growth <strong>and</strong> flowers.<br />

Prune young plants to encourage a framework <strong>of</strong><br />

strong shoots emerging from the base <strong>of</strong> the plant. Prune<br />

<strong>and</strong> shape plants after they flower, keeping them about<br />

3 feet high <strong>and</strong> removing all spindly <strong>and</strong> twiggy growth.<br />

Cut out any weak or damaged growth.<br />

Growth regulators<br />

To induce lateral branching, applications <strong>of</strong> BA<br />

(benzyladenine) at 50–100 ppm can be made when the<br />

plant tips are pinched back. Apply one spray 24 hours<br />

after the first pinch <strong>and</strong> another 24 hours after the second<br />

pinch. Dikegulac sodium at 2 ounces/gallon can be<br />

used in lieu <strong>of</strong> the second pinch-plus-BA application to<br />

improve branching. Atrimmec ® , a commercial formulation<br />

<strong>of</strong> dikegulac sodium (18.5% active ingredient), has<br />

been used at 1 ounce/gallon. Sprays should be applied<br />

to unpinched shoots when they reach 3 inches or to<br />

pruned plants three days after pruning. Avoid treating<br />

plants that are under stress.<br />

To retard growth, Cycocel ® (chlormequat) has been<br />

used on potted bougainvillea as a soil drench at 0.01–<br />

0.02 ounce per pot when the axillary buds swell following<br />

the first pinch. A-Rest (ancymidol) <strong>and</strong> Bonzi ®<br />

(paclobutrazol) are also effective. Use paclobutrazol at<br />

20–40 ppm when plants are near salable size.<br />

To prevent bract abscission, use the sodium salt <strong>of</strong><br />

NAA (naphthaleneacetic acid) as a spray or dip at 10–<br />

30 ppm. At the higher rate, NAA may promote abscission<br />

<strong>of</strong> immature bracts.<br />

It is recommended that you test plant growth regulators<br />

on just a few plants before extensive use. Always follow<br />

label instructions. If label application rates or methods<br />

differ from those given above, follow the label.<br />

Propagation<br />

Cuttings<br />

S<strong>of</strong>twood terminals, maturing green wood, <strong>and</strong> matured<br />

intermediate wood stem pieces can be used for propagation.<br />

Stem cuttings should be 1 ⁄8 inch or more thick<br />

<strong>and</strong> should have at least three to five nodes. Leaves may<br />

be left on the cuttings during rooting, but remove leaves<br />

from portions <strong>of</strong> the stem that are under the surface <strong>of</strong><br />

the rooting medium.<br />

Use a well drained rooting medium such as a 1:1 (by<br />

volume) peat:perlite mixture. Other rooting media such<br />

as man-made s<strong>and</strong> <strong>and</strong> peat or coir (coconut fiber) work<br />

well. Insert cuttings 1–2 inches into the medium <strong>and</strong><br />

water thoroughly. Cuttings may be rooted directly in<br />

small pots or Jiffy-7s. Several cuttings may be rooted<br />

together in larger pots 5–6 inches in diameter. Foam<br />

propagation blocks may also be used.<br />

S<strong>of</strong>twood terminals <strong>of</strong> easy-to-root cultivars do not<br />

require a rooting hormone. With more mature wood, a<br />

rooting hormone such as IBA (3-indolebutryic acid) at<br />

2000–6000 ppm is commonly used. Higher concentrations<br />

may be needed with more difficult-to-root cultivars.<br />

Intermittent misting is commonly used to prevent<br />

desiccation during rooting. Cuttings should not be allowed<br />

to wilt.<br />

Time to rooting is about 4–12 weeks, depending on<br />

the cultivar <strong>and</strong> the use <strong>of</strong> a rooting hormone. Transplant<br />

young plants with caution to avoid damaging their<br />

brittle roots. A broad-spectrum fungicide drench applied<br />

when planting cuttings <strong>and</strong> again after transplanting<br />

helps prevent root rot.<br />

Leaf-bud cuttings<br />

Leaf-bud cuttings can be used when source material is<br />

limited. Each node can be used as a cutting. The cutting,<br />

taken from shoots that are partially matured, consists<br />

<strong>of</strong> a leaf blade <strong>and</strong> a short piece <strong>of</strong> the stem (1–1 1 ⁄2<br />

5


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

inches) with the attached axillary bud. Cut the stem section<br />

about 1 ⁄2–1 inch above <strong>and</strong> below the point <strong>of</strong> leaf<br />

attachment. Place the bud vertically in a rooting medium<br />

<strong>and</strong> lightly cover it ( 1 ⁄4 inch) so that only the leaf<br />

blade can be seen.<br />

Grafting<br />

Some cultivars that have little or no chlorophyll in their<br />

leaves are difficult to grow from cuttings <strong>and</strong> need to be<br />

grafted onto a vigorous rootstock to be propagated. Grafting<br />

is useful with delicate cultivars that have fragile root<br />

systems. It is also used when it is desired to have multiple<br />

cultivars on one plant.<br />

The scion should be free <strong>of</strong> disease. The rootstock<br />

could be a seedling or a rooted cutting <strong>of</strong> an existing,<br />

established plant. All cut surfaces should be covered with<br />

grafting wax once the join is made to prevent moisture<br />

loss. Remove shoots from the rootstock. Put the grafted<br />

plant in a humid environment so the scion does not dry<br />

out.<br />

Various kinds <strong>of</strong> grafts may be used, including wedge,<br />

whip or tongue, or approach graft.<br />

Seeds<br />

Seeds germinate readily <strong>and</strong> require no treatments to<br />

break dormancy.<br />

Pests <strong>and</strong> diseases<br />

Contact the nearest Cooperative Extension Service <strong>of</strong>fice<br />

for assistance in identifying <strong>and</strong> recommendations<br />

for control <strong>of</strong> the various pests <strong>and</strong> diseases listed in the<br />

table on p. 8–9.<br />

Use in adornment<br />

<strong>Bougainvillea</strong> bracts are commonly used in lei making.<br />

Clip clusters <strong>of</strong> bracts, preferably in the early morning,<br />

<strong>and</strong> transport them in a paper or cloth bag. Wash with a<br />

cold-water soak. Bracts can be wrapped in damp newspaper,<br />

placed in a plastic container or paper box, <strong>and</strong><br />

stored in a refrigerator at 40°F for up to 14 days.<br />

<strong>Bougainvillea</strong> bracts are also used in haku (in Guam,<br />

mwar mwar), traditional, lei-like headb<strong>and</strong>s made <strong>of</strong><br />

leaves <strong>and</strong> flowers.<br />

6<br />

Acknowledgement<br />

Thanks go to Scot Nelson <strong>and</strong> Edwin Mersino for reviewing<br />

the manuscript.<br />

References<br />

Bradley, S. 2005. The pruner’s bible: A step-by-step<br />

guide to pruning every plant in your garden. Rodale,<br />

Inc., Emmaus, PA.<br />

Criley, R.A. 1997. <strong>Bougainvillea</strong>. In: M.L. Gaston, S.A.<br />

Carver, C.A. Irwin, <strong>and</strong> R.A. Larson (eds.), Tips on<br />

growing specialty potted crops. Ohio Florists’ Association,<br />

Columbus, OH.<br />

Gilman, E.F. 1999. <strong>Bougainvillea</strong> spp. Cooperative Extension<br />

Service, University <strong>of</strong> Florida, Fact Sheet<br />

FPS-70.<br />

Growing plants for Hawaiian lei: 85 plants for gardens,<br />

conservation, <strong>and</strong> business. 2002. University <strong>of</strong><br />

Hawai‘i at Mänoa, <strong>College</strong> <strong>of</strong> <strong>Tropical</strong> <strong>Agriculture</strong><br />

<strong>and</strong> <strong>Human</strong> Resources (CTAHR).<br />

Hackett, W.P., R.M. Sachs, <strong>and</strong> J. DeBie. 1972. Growing<br />

bougainvillea as a flowering pot plant. California<br />

<strong>Agriculture</strong> 26(8):12–13.<br />

Hamilton, D.F., <strong>and</strong> J.T. Midcap. 2003. Propagation <strong>of</strong><br />

woody ornamentals by cuttings. Cooperative Extension<br />

Service, University <strong>of</strong> Florida.<br />

Iredell, J. 1994. Growing bougainvilleas. Simon &<br />

Schuster Australia. East Roseville, NSW, Australia.<br />

Neal, M.C. 1965. In gardens <strong>of</strong> Hawaii. Bishop Museum<br />

Press, Honolulu.<br />

Pertuit, A. 1999. <strong>Bougainvillea</strong>s. Cooperative Extension<br />

Service, Clemson University, Home & Garden Information<br />

Center HGIC 1553.<br />

Schoellhorn, R., <strong>and</strong> E. Alvarez. 2002. Warm climate<br />

production guidelines for bougainvillea. Cooperative<br />

Extension Service, University <strong>of</strong> Florida, ENH 874.<br />

Tavares, J., D. Hensley, J. Deputy, D. Tsuda, <strong>and</strong> A. Hara.<br />

1998. <strong>Bougainvillea</strong> looper. CTAHR publication IP-2.<br />

Teho, F. 1971. Plants <strong>of</strong> Hawaii: How to grow them.<br />

The Petroglyph Press, Hilo, HI.<br />

Watson, D.P., <strong>and</strong> R.A. Criley. 1973. <strong>Bougainvillea</strong>s.<br />

CTAHR Circular 469.


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Internode length <strong>and</strong> l<strong>and</strong>scape characteristics<br />

Internode length has a strong influence on bougainvillea<br />

growth habit <strong>and</strong> characteristics in the l<strong>and</strong>scape. Compare<br />

the distance between bract nodes on the upper two<br />

stems, from the same plant, with the stem from a different<br />

plant, below. (The tiles are 10 inches square.)<br />

Here is the plant with the shorter internodes, compact <strong>and</strong><br />

easily shaped <strong>and</strong> maintained in the l<strong>and</strong>scape.<br />

Here is the plant with the longer internodes; note its exuberant<br />

growth habit. Highly colorful at some times <strong>of</strong> year,<br />

it is capable <strong>of</strong> climbing 30–40 feet into the Norfolk Isl<strong>and</strong><br />

Pine behind it. Maintanence can be laborious.<br />

7


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Pests, diseases, <strong>and</strong> physiological problems<br />

Problem Description Symptoms Control<br />

aphids Small, rounded or pear- Aphids suck the sap, causing Insecticidal soap is the safest <strong>and</strong> most effective<br />

shaped, s<strong>of</strong>t bodied speckles on the foliage, control strategy. Controlling the ants will go a long<br />

insects. Clustered on reduced color, stunting, way in helping to control aphids, as well as<br />

stems, leaves, buds, <strong>and</strong> wilting, <strong>and</strong> deformed leaves. scales <strong>and</strong> mealybugs. Ants move these insects<br />

flowers. Usually found on In severe cases, leaf loss around <strong>and</strong> harvest the honeydew they exude.<br />

new growth. Most aphids occurs. Ants also protect these insects from predators<br />

have a pair <strong>of</strong> tube-like <strong>and</strong> parasites.<br />

cornicles on the posterior<br />

<strong>of</strong> the abdomen.<br />

bougainvillea Green or brown smooth Leaves are eaten. Severe Bacillus thuringiensis (BT) insecticide. Neemloopers<br />

skin caterpillar, one inch scalloping <strong>of</strong> leaves. As the based biological insecticide products. Most<br />

(Disclisioprocta long. Feeds primarily at population multiplies, plants synthetic insecticides with labels permitting use<br />

stellata) night. can be defoliated. against caterpillars on l<strong>and</strong>-scape ornamentals.<br />

Organic insecticide Spinosad. Spraying late in<br />

the evening, when caterpillars <strong>and</strong> adult moths<br />

are active, is recommended.<br />

grasshoppers Grasshoppers are easy to These insects occasionally Spray plants with an insecticide registered for<br />

see <strong>and</strong> should be consume large quantities <strong>of</strong> ornamental plants.<br />

controlled before they foliage. Their feeding leaves<br />

become numerous. an irregular appearance.<br />

mealybugs White, cottony masses Mealybugs suck plant Systemic insecticides are preferred. Horticultural<br />

found in leaf axils, on juices. Infested plants oil can be used, but the label should be followed<br />

underside <strong>of</strong> leaves, on become stunted, <strong>and</strong> plant to avoid burning the leaves <strong>and</strong> flowers. Burning<br />

roots, <strong>and</strong> other protected parts may die. Honey-dew may occur if the concentra-tion is too high or if<br />

areas. <strong>and</strong> sooty mold are <strong>of</strong>ten the plants are sprayed in the heat <strong>of</strong> the day.<br />

present. HDOA is releasing predator insects that should<br />

help control mealybugs in the future.<br />

mites These small pests feed on Feeding causes curling, Minimize the possibility <strong>of</strong> introducing mites into<br />

young new growth. Most puckering, deformity, the growing area on infested plant material.<br />

mites are very small <strong>and</strong> distortion, <strong>and</strong> bronzing. Eliminate weeds that are alternate hosts. Ultra<br />

difficult to see even with a Wilting may occur. fine oil, insecticidal soap, or neem may also be<br />

h<strong>and</strong> lens. used. Wash foliage with a strong spray <strong>of</strong> cold<br />

water. Wash the plants with soapy water <strong>and</strong> a<br />

s<strong>of</strong>t brush or cloth to remove insects.<br />

reniform Microscopic parasitic Poor growth, stunted, The goal is to manage their population, reducing<br />

nematodes roundworms that live in chlorotic (yellow) foliage; their numbers below damaging levels.<br />

<strong>and</strong> feed on roots, premature wilting, low vigor, Incorporate good compost or organic materials,<br />

damaging roots <strong>and</strong> thin canopy, <strong>and</strong> leaf <strong>and</strong>/or such as manure or wood shavings, into the soil<br />

preventing normal uptake bloom loss under relatively as a preplant amendment to encourage microbial<br />

<strong>of</strong> water <strong>and</strong> nutrients. mild stress; swollen, activity to depress nematode populations. Grow<br />

knotted, gnarled areas on French marigolds, tilling it in as green mulch.<br />

the roots. Solarization, the heating <strong>of</strong> soil by using clear<br />

plastic tarps to increase <strong>and</strong> trap the sun’s heat,<br />

can be an effective means <strong>of</strong> controlling<br />

nematodes in the soil.<br />

8


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Problem<br />

scales<br />

(Coccus<br />

hesperidium)<br />

whiteflies<br />

leaf spots*<br />

root rots<br />

(Rhizoctonia,<br />

Pythium, <strong>and</strong><br />

Phytophthora)<br />

chlorosis<br />

Description<br />

Scales are sedentary<br />

insects that are covered by<br />

a hard protective covering<br />

(shell) or waxy secretion.<br />

Appear on leaves, petioles,<br />

or stems.<br />

Sap-feeding insects that<br />

feed on undersides <strong>of</strong><br />

leaves. Whiteflies are tiny;<br />

the adults resemble white<br />

moths. The immature<br />

stages look like scale<br />

insects.<br />

Bacteria <strong>and</strong> fungi cause<br />

leaf spots. Can be a<br />

problem in prolonged wet<br />

conditions.<br />

Fungal diseases <strong>of</strong><br />

bougainvillea roots,<br />

resulting in their<br />

malfunction <strong>and</strong> decay.<br />

May attack cuttings during<br />

propagation. Pythium <strong>and</strong><br />

Phytophthora are classified<br />

as fungus-like organisms.<br />

A lack <strong>of</strong> normal green<br />

pigmentation in the foliage<br />

generally due to deficiency<br />

<strong>of</strong> one or more micronutrients<br />

(usually magnesium<br />

or iron) or root rot.<br />

Chlorosis could indicate a<br />

pathogenic attack <strong>of</strong> the<br />

roots or foliage, which can<br />

mimic a nutritional<br />

deficiency such as nitrogen.<br />

Symptoms<br />

Scales suck plant juices.<br />

They appear as raised or<br />

brown bumps (scales) which<br />

give stems or leaves a lumpy<br />

appearance. Feeding causes<br />

a characteristic yellow or<br />

chlorotic streak that radiates<br />

from point <strong>of</strong> attachment.<br />

Infested leaves <strong>of</strong>ten have<br />

small yellow spots where<br />

adults or immature whiteflies<br />

have fed. When populations<br />

become dense, the leaves<br />

become yellow <strong>and</strong> lower<br />

leaves are covered with<br />

black sooty mold.<br />

Small, reddish, round spots<br />

on the leaves. Spots are<br />

usually surrounded by a<br />

“halo” <strong>of</strong> pale green <strong>and</strong><br />

rapidly exp<strong>and</strong> to become<br />

irregular dark patches.<br />

Defoliation may occur.<br />

Wilting, dieback, stunting,<br />

chlorosis <strong>of</strong> foliage.<br />

Pale green leaves with dark<br />

green veins; yellowing <strong>of</strong> the<br />

foliage.<br />

Control<br />

Horticultural oils work well. A systemic insecticide<br />

may have to be used against scales. Control the<br />

ants that move the scales around <strong>and</strong> farm them.<br />

Wash the plants with soapy water <strong>and</strong> a s<strong>of</strong>t<br />

brush or cloth to remove insects. Thoroughly<br />

wash undersides <strong>of</strong> leaves where pests may also<br />

reside. Remove with alcohol-saturated cotton<br />

swab.<br />

Many insecticides are registered for <strong>and</strong> effective<br />

at controlling whiteflies. Undersides <strong>of</strong> leaves<br />

must be covered thoroughly to achieve<br />

satisfactory control. Successive sprays <strong>of</strong><br />

insecticidal soaps, oils, or synthetic pyrethrum.<br />

Various broad-spectrum fungicides may be used.<br />

Remove all dead leaves <strong>and</strong> reduce crowding<br />

between plants. Discard infected plants. Try to<br />

avoid overhead sprinkling <strong>and</strong> late afternoon<br />

watering.<br />

Remove <strong>and</strong> destroy symptomatic plants as soon<br />

as they are found. For Rhizoctonia, apply a<br />

broad-spectrum fungicide drench during<br />

transplanting or planting in the l<strong>and</strong>scape. Use a<br />

metalaxyl or phosphorus acid-based product for<br />

Pythium or Phytophthora. Avoid overwatering.<br />

Avoid planting in heavy soils.<br />

Acidify the soil with aluminum sulfate, iron<br />

sulfate, or wettable sulfur. An acidulating<br />

houseplant fertilizer can be used to lower the soil<br />

pH. An application <strong>of</strong> a complete micronutrient<br />

blend or chelated spray should help. Allow the<br />

soil to remain evenly moist but not saturated.<br />

Remove excess plants in a hedge.<br />

*(Pseudomonas stizilobii, Burkholderia <strong>and</strong>ropogonis, Cercosporidium bougainvilleae, Colletotrichum dematium forma bougainvillae,<br />

Gladosporum arthrinoides)<br />

Note: Read <strong>and</strong> follow manufacturers’ directions on labels. The pesticides mentioned are possible options for control, but mention <strong>of</strong> them<br />

should not be considered a recommendation. Licensing for sale <strong>and</strong> use <strong>of</strong> pesticide formulations varies from state to state <strong>and</strong> country to<br />

country. The pesticide label is the law. Read it before purchasing a pesticide to ensure that the product is registered for your intended use.<br />

Read the label entirely before use <strong>and</strong> follow its instructions. Mention in this publication to a company or product name does not imply<br />

recommendation <strong>of</strong> the product or company to the exclusion <strong>of</strong> others that may also be suitable.<br />

9


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Cultivars (varieties)<br />

Cultivar Bract color Comments<br />

Afterglow Yellow-orange Sparse foliage. Bracts turn a dusty rose color when old. Heavy bloomer.<br />

Barbara Karst Bright red A popular cultivar that performs well. Moderate to vigorous growth.<br />

Leaves dull, dark green, broadly ovate with pointed tips, slightly wavy<br />

margins. A constant bloomer. Flowers white <strong>and</strong> conspicuous. Thorns<br />

medium <strong>and</strong> straight.<br />

Bois-de-Rose Dusty pink Vigorous growth. Dark green furry leaves. Thorns large, slightly recurved.<br />

California Gold Gold, yellow Nice vining habit. Large long-lasting rich gold bracts; can appear yellow<br />

in some environments.<br />

Crimson Jewel Crimson, pink, orange Dwarf. Good container plant <strong>and</strong> ground cover.<br />

Double Pink Pink Huge clusters <strong>of</strong> bright pink double bracts. Repeat bloomer.<br />

Double Red Deep red Clusters <strong>of</strong> fluffy double bracts over nice green leaves.<br />

Dr. David Barry Lavender-pink Round shrubby vigorous cultivar with long elliptic leaves tapering to a<br />

(Singapore Pink, pointed tip. Very large elliptic reflex bracts. Flowers prominent, cream.<br />

Singapore Beauty) Thorns short, not prominent. Fine for containers.<br />

Elizabeth Angus Purple Vigorous growing. Leaves dark green, glossy, <strong>and</strong> tapering. Young<br />

stems puberulent. Bracts large. Flowers yellowish, large, <strong>and</strong><br />

conspicuous. Thorns stout, long, recurved.<br />

Golden Glow Bright gold, pinkish-gold Vigorous <strong>and</strong> erect. Leaves rounded <strong>and</strong> large. Bracts broadly ovate.<br />

(Millarii, Gold Queen,<br />

Hawaiian Gold)<br />

Thorns medium <strong>and</strong> straight.<br />

Golden Summers White Variegated yellow-gold <strong>and</strong> green foliage. Large bracts. Repeat<br />

(Miss Oneuse) bloomer. Thorns medium <strong>and</strong> slightly curved.<br />

Helen Johnson Reddish-purple, A dwarf compact bushy grower about three feet tall <strong>and</strong> wide. Branches<br />

(Temple Fire) opening slightly coppery freely. Leaves are small, broadly ovate, sharply pointed at apex; margins<br />

slightly wavy. Bracts sparse, broadly ovate, pointed apex. Thorns small,<br />

short, slender. Good for hanging baskets <strong>and</strong> as a ground cover.<br />

Jamaica White White with flush <strong>of</strong> pink Medium grower. Leaves large, rounded, dark green, <strong>and</strong> smooth. Bracts<br />

(Apple Blossom,<br />

Audrey Grey)<br />

medium size. Very pretty but rather shy flowering. Thorns medium.<br />

Jane Snook Pink Compact, dense. Tends to be pendulous with some vigorous canes.<br />

(Durban, President) Leaves long <strong>and</strong> light green with long petioles. Bracts large with ruffled<br />

edges. Flowers large <strong>and</strong> greenish-cream. Thorns medium <strong>and</strong> fine.<br />

John Lattin Iridescent pale lavender Erect leggy grower. Leaves long, ovate, medium green, glossy, <strong>and</strong><br />

pointed at tips. Bracts taper to a long point at apex. Flowers moderate in<br />

size, greenish-cream. Thorns long <strong>and</strong> curved.<br />

Juanita Hatten Bright red, dark pink Moderate growth habit; branches freely. Foliage may be slightly<br />

variegated in shades <strong>of</strong> green. Attractive mid-green, ovate leaves with<br />

slightly wavy margins. Bracts ovate, medium to large. Tends to be everblooming.<br />

Flowers prominent <strong>and</strong> creamy-white. Thorns moderate <strong>and</strong><br />

only slightly recurved. Good for hanging baskets.<br />

Killie Campbell Copper, red, magenta Large-growing, pendulous. Leaves dark green, long, ovate, margins<br />

wavy, tapering to long-pointed apex. Petiole long. Bracts large, thintextured,<br />

<strong>and</strong> ruffled. Flowers large, conspicuous. Thorns long, straight.<br />

Lady Hudson Pale pink Long canes <strong>of</strong>ten bear <strong>of</strong> leaves especially when young. Leaves broadly<br />

(Princess Margaret Rose) ovate with long petioles, light green, <strong>and</strong> glabrous. Bracts small, ruffled.<br />

10


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Cultivar Bract color Comments<br />

Lateritia Brick-red Dense, shrubby, slow growing. Leaves medium size, furry. Bracts<br />

(Dar-es-Salaam) medium size. Flowers conspicuous, creamy-white. Thorns medium, curved.<br />

Lady Mary Baring Bright yellow Growth erect <strong>and</strong> open. Leaves large, rounded, dark green. Bracts<br />

(Yellow Glory, medium size, rounded. Flowers not conspicuous. Thorns medium, stout,<br />

Hawaiian Yellow) <strong>and</strong> straight.<br />

Meriol Fitzpatrick Pale, shimmering Bushy. Leaves dark green <strong>and</strong> elliptic. Bracts thin, ovate, tapering, with<br />

(Dream) lavender slightly ruffled margins. Flowers medium in size, creamy-white.<br />

Thorns medium <strong>and</strong> recurved.<br />

Miami Pink Dark pink Does best in l<strong>and</strong>scapes.<br />

Miss Manila Golden-orange, Vigorous, cascading, dense habit. New growth coppery. Leaves ovate,<br />

(Tango) apricot, light pink light to medium green. Bracts large, whorled, rounded at apex, slightly<br />

ruffled. Flowers large <strong>and</strong> prominent, cream. Thorns medium to strong,<br />

straight.<br />

Mrs. Butt Red to peach Habit loose <strong>and</strong> upright, vigorous. Leaves large <strong>and</strong> rounded with a<br />

(Dame Clara Butt, short sharp point at apex. Bracts medium size, broadly ovate. Repeat<br />

Crimson Lake) bloomer. Flowers prominent, white with a tinge <strong>of</strong> pink or red. Thorns<br />

medium, stout, <strong>and</strong> straight.<br />

Mrs. McClean Bright orange Erect growth. Leaves large, rounded. Bracts medium size. Flowers not<br />

(Hawaiian Orange,<br />

Orange Glory)<br />

conspicuous, cream. Thorns medium, stout, <strong>and</strong> straight.<br />

Oo-La-La Purple Compact, small to moderate growth, bushy. Canes short. New growth<br />

(Poultoni) copper-red. Leaves broadly ovate, pointed apex, slightly wavy margins.<br />

Bracts moderately large with acute apex. Bracts open copper-red,<br />

become magenta-red, <strong>and</strong> finally have a bluish appearance. Cream<br />

flowers are conspicuous. Thorns few, small, slightly curved.<br />

Orange Fiesta Pumpkin Luminous bracts have hints <strong>of</strong> pink <strong>and</strong> salmon. Allowing plants to wilt<br />

slightly encourages more blossoms.<br />

Pink Pixie Pink, magenta-red True miniature. Small leathery leaves, internodes very short, <strong>and</strong> leaves<br />

(Hawaiian Torch, crowded. Bracts small <strong>and</strong> densely packed. Flowers prominent, small,<br />

Smartipants) <strong>and</strong> white. Thorns short, stubby, <strong>and</strong> blunt. Good for small pots.<br />

Raspberry Ice Red Compact, bushy grower. New shoots reddish. Leaves small to medium,<br />

(Hawaii, ovate, tapered to a pointed apex, margins wavy. Variegated foliage with<br />

<strong>Tropical</strong> Rainbow) cream to golden yellow margins, dark grayish-green center. Bracts<br />

medium, ovate. Repeat bloomer. Thorns short, thin, <strong>and</strong> straight.<br />

Rosa Catalina Rosy pink Dense, busy, <strong>and</strong> vigorous. Leaves ovate, furry, tapering to a point.<br />

Bracts medium, ovate with pointed apex. Flowers prominent.<br />

Thorns medium, stout.<br />

Royal Purple Dark purple Good replacement for Elizabeth Angus in humid areas.<br />

Scarlet O’Hara Orange, dark red Loose, upright, vigorous with strong canes. New growth dark red,<br />

(San Diego Red, slightly puberulent. Leaves very large, rounded, dark green, thick-<br />

Hawaiian Scarlet) textured. Bracts nearly round <strong>and</strong> very large. Flowers conspicuous,<br />

cream, large. Many thorns, medium, recurved.<br />

Summer Snow White Large bracts. Excellent for weddings. Does well in hanging baskets.<br />

Sundown Apricot Vigorous. Good bloomer. Heavy feeder.<br />

Surprise (Mary Palmer) Pink <strong>and</strong> white Grows large. Leaves large <strong>and</strong> mid-green.<br />

Sweet Dream Pale lavender Leaves completely gold. Thornless.<br />

11


UH–CTAHR <strong>Bougainvillea</strong> OF-38 — Oct. 2007<br />

Cultivar Bract color Comments<br />

Texas Dawn Light purple-pink Loose, open habit. Leaves large <strong>and</strong> rounded. Bracts rounded, medium<br />

(Purple King, size. Flowers not conspicuous, creamy-white. Thorns medium to long,<br />

Robyn’s Glory) stout, <strong>and</strong> slight recurved.<br />

<strong>Tropical</strong> Bouquet Orange, pink Compact <strong>and</strong> bushy. Bracts orange but quickly become pink.<br />

Variegata Variegated foliage.<br />

Vickie Pink, white Leaves have a large splash <strong>of</strong> gold or creamy yellow in the center.<br />

(Vicky, Vicki, Thimma) Some white bracts may be sported.<br />

Other cultivars include Carmencita, Betty Hendry, Bridal Bouquet, Buttiana, Carmencita, Closeburn, Convent, Doubloon, Dream,<br />

Dulci Dayborn, Easter Parade, Fascelles Purple, Gillian Greensmith, Gopal, Harrissi, Hugh Evans, India, Indica, Indian Flame,<br />

Jubilee, Lacquer Red, Lemner’s Special, Lilac Queen, Lyamunga No. 2, New River, Orange King, Pink Champagne, Purple<br />

Prince, Rao, Red Glory, Rhodomine, Rose Pink, San Diego Red, William Paulton, <strong>and</strong> Yellow Glory.<br />

12

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