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All Theses and Dissertations
2017-12-01
Laying the Foundation for a Fremont Phytolith
Typology Using Select Plant Species Native to
Utah County
Madison Natasha Pearce
Brigham Young University
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Laying the Foundation for a Fremont Phytolith Typology
Using Select Plant Species Native to Utah County
Madison Natasha Mercer Pearce
A thesis submitted to the faculty of
Brigham Young University
in partial fulfillment of the requirements for the degree of
Master of Arts
Michael T. Searcy, Chair
James R. Allison
Terry Briggs Ball
Department of Anthropology
Brigham Young University
Copyright © 2017 Madison Natasha Mercer Pearce
All Rights Reserved
ABSTRACT
Laying the Foundation for a Fremont Phytolith Typology
Using Select Plant Species Native to Utah County
Madison Natasha Mercer Pearce
Department of Anthropology, BYU
Master of Arts
Archaeobotanical evidences for the presence of wild plants at Fremont archaeological sites
are numerous. However, little can be positively argued for why those plants are present, if they
were used by site inhabitants, and how they were used. Additionally, there are likely several wild
plants that were used but that do not appear in the archaeobotanical record as pollen or
macrobotanicals, the two most commonly identified plant remains. I argue that it is possible to
provide better interpretations for how and why the Fremont used plants by researching how their
historic counterparts, the Goshute, Shoshone, Ute, and Southern Paiute, used the same plants that
are identified at prehistoric sites. I further argue that a phytolith typology for Fremont archaeology
can provide more insight into prehistoric plant use. I demonstrate its utility through a phytolith
analysis of ground stone tools from Wolf Village, a Fremont site in Utah County.
Keywords: ethnobotany, Fremont, phytoliths, Utah County, Goshute, Shoshone, Ute, Paiute, Wolf
Village, ground stone tools.
ACKNOWLEDGEMENTS
I would like to acknowledge my committee, the botanists and gardeners who helped me
identify plants, and my family and friends.
I would like thank Michael Searcy who, when I approached him in August 2013 as a very
stressed and vexed Master’s student in the Environmental Science program, cleared the way for
my application and acceptance in an unprecedented manner into the Department of
Anthropology. I’d also like to thank him for being my friend in Kanab in 2010. I’d like to thank
James Allison for letting me pursue this project; his trust and sagacious apprehension. I would
especially like to thank Terry B. Ball for being so welcoming when he first opened his office
door to a student he had never met before. I am grateful for all his guidance and positive
encouragement.
I am grateful for funds received for this research from the Charles Redd Center and the
Anthropology Department. Research funds came from the Summer Award for BYU Upper
Division and Graduate Students and the Shallit Grant. I would like to thank Casey from Central
Utah Garden, Robert Johnson from the BYU Herbarium, Heidi, Lindsey, Neal, and Fritz from
Red Butte Gardens, Tony and Esther from Thanksgiving Point, and Kim and Gail from Sego
Lily Gardens. I am grateful for Bradley Geary for allowing me to use his Nikon Optiphot 2 light
microscope, and for Mike Standing in the McDonald microscopy lab. I would also like to thank
Brooks B. Britt and Loreen Allphin for always believing in me.
I am indebted to my husband for all his support, for listening to my crazy ideas, and for
baking me cookies. I am grateful for my two children and the perspective they brought when
times were tough. I am grateful for my mother for editing all my drafts. I am also grateful for all
my babysitters: Tess, Lynn, Emily, Chloe, Jessica, Kara, Savannah, Britney, Aly, Izzy, Zoe,
Kim, Chanae, Danielle, Miranda, April, and Kari.
Finally, I am grateful for my God. I wouldn’t be in this program if He did not inspire me
to be here. I would not have made it this far without His help.
TABLE OF CONTENTS
ABSTRACT.................................................................................................................................................. ii
ACKNOWLEDGEMENTS ......................................................................................................................... iii
TABLE OF CONTENTS .............................................................................................................................. v
LIST OF FIGURES ................................................................................................................................ vii
LIST OF TABLES ................................................................................................................................. viii
Introduction ................................................................................................................................................. 1
Wolf Village .............................................................................................................................................. 2
Phytoliths .................................................................................................................................................. 3
Thesis Organization .................................................................................................................................. 6
2. Background ............................................................................................................................................. 8
The Fremont .............................................................................................................................................. 8
Ethnographies of Historic Indigenous Occupation in Utah Valley ........................................................ 17
The Shoshone .......................................................................................................................................... 19
The Goshute ............................................................................................................................................ 20
The Utes .................................................................................................................................................. 22
The Southern Paiute................................................................................................................................ 25
3. Methods.................................................................................................................................................. 28
Fremont Botanical Reports ..................................................................................................................... 28
Plant Species Native to Utah County and Documented Ethnographic Uses .......................................... 29
Plant Collection and Digestion ............................................................................................................... 31
Groundstone Slide Analysis .................................................................................................................... 36
4. Results of Typology ............................................................................................................................... 37
Calcium oxalates..................................................................................................................................... 39
Forbs ....................................................................................................................................................... 44
Trees and Shrubs..................................................................................................................................... 52
Grasses.................................................................................................................................................... 62
Discussion and Conclusion ..................................................................................................................... 71
5. Results of Ground Stone Artifact Wash Analysis .............................................................................. 74
Ground Stone Artifact 219 ...................................................................................................................... 76
Ground Stone Artifact 2357 .................................................................................................................... 76
Ground Stone Artifact 11975 .................................................................................................................. 81
v
Ground Stone Artifact 15814 .................................................................................................................. 81
Ground Stone Artifact 16494 .................................................................................................................. 82
Ground Stone Artifact 16642 .................................................................................................................. 87
Summary ................................................................................................................................................. 88
6. Discussion and Conclusion ................................................................................................................... 94
Phytolith Typology and Ground Stone Analysis ..................................................................................... 94
Indicators for Fremont Subsistence ........................................................................................................ 95
Conclusion .............................................................................................................................................. 99
References Cited ....................................................................................................................................... 101
Appendix A ............................................................................................................................................... 122
Appendix B ............................................................................................................................................... 128
Appendix C ............................................................................................................................................... 141
Appendix D ............................................................................................................................................... 153
Appendix E ............................................................................................................................................... 207
Appendix F.................................................................................................................................................209
vi
LIST OF FIGURES
Figure 2.1. Map of Utah Valley. ................................................................................................... 13
Figure 3.1. Common cross contaminates found in phytolith slides .............................................. 34
Figure 4.1. Calcium oxalates: raphides and styloids..................................................................... 42
Figure 4.2. Calcium oxalates: crystal sand, druses, prismatics, and rhombohedrals. ................... 43
Figure 4.3. Phytoliths: astrosclerids and hairs. ............................................................................. 48
Figure 4.4. Phytoliths: articulate epidermals, part one. ................................................................ 49
Figure 4.5. Phytoliths: articulate epidermals, part two. ................................................................ 50
Figure 4.6. Phytoliths: articulate epidermals part three, stomates, parenchyma, and trichomes. . 51
Figure 4.7. Phytoliths: tracheids. .................................................................................................. 59
Figure 4.8. Phytoliths: vascular tissue and papillae. ..................................................................... 60
Figure 4.9. Phytoliths: spheroids and polygonals. ........................................................................ 61
Figure 4.10. Phytoliths: elongates, part one. ................................................................................. 68
Figure 4.11. Phytoliths: elongates, part two. ................................................................................ 69
Figure 4.12. Phytoliths: grass short cell forms. ............................................................................ 70
Figure 5.1. Ground Stone FS 219, basalt metate fragment. .......................................................... 78
Figure 5.2. Ground Stone FS 2357, a worn and pecked central fragment of a vesicular basalt
mano.............................................................................................................................................. 78
Figure 5.3. Ground Stone FS 11975, a complete quartzite basin mano........................................ 83
Figure 5.4. Ground Stone FS 15814, a complete rhyolite mano. .................................................. 84
Figure 5.5. Ground Stone FS 16494, fragment of a basalt Utah-style metate. ............................. 84
Figure 5.6. Ground Stone FS 16642, a platform for mashing roots and tubers. ........................... 88
Figure 5.7. Maize phytoliths observed on ground stone artifacts. ................................................ 90
Figure 5.8. Calcium oxalate crystals, chrysophyte cysts, diatoms, sponges, and epidermals. ..... 91
Figure 5.9. Grass phytoliths. ......................................................................................................... 92
Figure 5.10. Diagnostic forms, sedge, tracheids, and others. ....................................................... 93
vii
LIST OF TABLES
Table 2.1. The Frequency of Archaeobotanical Remains at Fremonts Sites in Utah County. ..... 14
Table 3.1. Examples of Changes in Plant Nomenclature.............................................................. 31
Table 4.1. Typology Code. ........................................................................................................... 38
Table 4.2. Presence and Frequency of Calcium Oxalate Crystals. ............................................... 41
Table 4.3. Presence and Frequency of Phytoliths in Tested Forbs. .............................................. 47
Table 4.4. Presence and Frequency of Phytoliths in Tested Shrubs and Trees ............ ................ 57
Table 4.5. Presence and Frequency of Phytoliths in Tested Grasses ........................................... 66
Table 4.6. Diagnostic and Non-diagnostic Phytoliths. ................................................................. 71
Table 5.1. Wolf Village (42UT273) Ground Stone Tool Provenience and Description. ............. 75
Table 5.2. Phytolith Grass Counts. ............................................................................................... 79
Table 5.3. Diagnostic Phytolith Counts. ....................................................................................... 80
Table 5.4. Redundant Forms, Total Phytolith Count, and Other Forms. ...................................... 85
Table 5.5. Calculated Percentages for Select Phytolith Counts. ................................................... 86
viii
1. Introduction
Of the Fremont who lived in the Great Basin from about AD 300 to AD 1300, much is
known about their architecture, artifacts, and other lifeways. However, knowledge of their plant
use is lacking regarding what plants the Fremont used for food and/or medicine. Two methods
have been commonly employed to evaluate plant use in the Great Basin: macrobotanicals and
pollen analyses. Another method, phytolith analysis, has the ability to shed further light on
Fremont plant use but is rarely used because phytolith typologies for the region are scarce or
incomplete. There is also no collective database with which one can compare to identify all the
species and tissues of plants that have been tested for the presence of phytoliths.
To address this, I created a phytolith typology based on species from plant families and
genera identified at Fremont Utah Valley sites that have documented ethnographic uses. While
this typology is more specific to the Utah Valley Fremont, it can be expanded to surrounding
areas and cultural groups in the eastern Great Basin and western Colorado Plateau. Due to time
restraints, this typology is only a sample of species native to Utah County. I tested this typology
by identifying phytoliths from six ground stone artifacts from Wolf Village, a Fremont village
site located in Utah Valley. I hired a paleobotanist, Chad Yost, to analyze these samples first,
after which I compared my results to his.
Ethnographic records of the Shoshone, Goshute, Ute, and Southern Paiute were used to
provide interpretations for how and why plants identified at Utah Valley Fremont sites may have
been used. Such interpretations can aid in assessing the functionality of structures and use
surfaces, as well as help justify why botanical studies are conducted at prehistoric sites. These
groups were selected because although they are separated from the Fremont by time, they do
1
share with the Fremont similarities in architecture, artifacts, and other lifeways, in addition to
living in and around Utah Valley.
Wolf Village
I chose to study Utah Valley in Utah County because of its convenient location. To test
the viability of my phytolith typology, I selected artifacts from the Utah Valley Fremont site
Wolf Village because of the large number of unwashed ground stone artifacts available to study.
Wolf Village is a Fremont site located in the southern end of Utah Valley, a few miles
south of Utah Lake. Radiocarbon dates from the excavated structures suggest that this site was
occupied for a few decades around AD 1000-1100 (Johansson et al. 2014:33). At least seven pit
structures and two surface structures have been excavated, with the largest pit structure and
largest surface structure of the Fremont occupation of Utah Valley being found at this site. All
structures have floors of varying degrees of discernibility, post-holes, and most have hearths
(Johannson et al. 2014).
A macro- and micro-botanical study of the site from 2011 revealed that the Wolf Village
Fremont utilized, and likely farmed, Zea mays and Phaseolus sp. (Dahle 2011:33). They also
utilized several wild, native species, such as Brassica (mustard), Stipa (ricegrass), Helianthus
(sunflower), and Amelanchier (serviceberry), among others (Dahle 2011:37). Plant remains
indicative of these and several other plants came from ground stone artifact pollen analysis and
macrobotanicals collected from midden and hearth fill samples.
To understand Fremont plant use, studies using multiple methods are ideal. A review of
Utah Valley Fremont botanical reports reveal that plants remains preserve differently in the
valley. Some only preserve as pollen or as seeds, and some do not preserve at all. For example,
2
few to no plant remains have been recovered from sites such as Seamons Mound near Utah Lake
due to poor preservation because of damp soil. Furthermore, there are other plants, such as
Polygonaceae Polygonum, that are found in Utah Valley archaeological contexts that have no
records of use by historically documented groups. Several of these plants are invisible
ethnographically because either no one recorded using those plants, or no one remembers those
plants being used.
Phytoliths
Phytoliths, or plant opal silica, are the inorganic microfossil remains of plants which form
as the plant uptakes monosilicic acid, Si(OH4), through its roots and deposits that silica in
intracellular and extracellular locations (Pearsall et al. 1995:184). There the silica forms a solid
deposit that takes the shape of the cell in which it forms; in some cases, this shaped silica, or
phytolith, is taxonomically significant (Pearsall et al. 1995:184).
Phytoliths have two characteristics that in some cases make them more useful than pollen
and starch grains for archaeobotanical investigations. First, due to their inorganic nature,
phytoliths preserve better in environments that would typically destroy organic remains, such as
oxidized soils in places like Mesoamerica (Pearsall 1989:254). Second, spores, pollen, and seeds
are primarily produced during specific seasons, while phytoliths can be produced year-long and
are sometimes developed in plant structures that do not produce other forms of microfossils (Ball
et al. 2015:11).
Phytoliths are released once the plant producing them is destroyed through decay,
burning, digestion, or grinding (Ball et al. 1999:1615; Shahack-Gross et al. 2014; Wallis et al.
2014). These microfossils can then be found in several different contexts: soil, coprolites, dental
calculus, stomach contents, residue on artifacts, and lake cores (Berlin et al. 2003:115; Piperno
3
2006:81-86). They have also been found fossilized in significantly older contexts, with some
samples dating to around nine million years B.P. (Schultz et al. 2014). Phytoliths found in these
different settings have added to both prehistoric and present day environmental and
ethnobotanical reconstruction (Ball et al. 2015).
Prehistoric diet and subsistence can also be explored by analyzing phytoliths found on
ground stone tools or on ceramic vessels (Liu et al. 2014; Peto et al 2013; Power et al. 2014).
Recent phytolith research into Neanderthal diets indicates that they probably consumed just as
many plant taxa as modern humans (Henry et al. 2014). In biology, coprolites of unobservable
chimpanzee populations have been collected for phytolith analysis to determine how the
population is faring based on their diet, since heavily digested plants will only be present in feces
as phytoliths (Phillips and Lancelotti 2014).
Changes in the environment are sometimes reflected in phytoliths. Since the
photosynthetic pathways of plants affects their cell structures, researchers have been able to
distinguish between the phytoliths created by C3 versus C4 grasses (Stromberg 2002). Because of
this morphological difference, phytoliths have been used to track changes in the environment by
tracking the diversity and richness of C3 and C4 grass phytoliths in prehistoric soils (Cotton et al.
2014). Climate patterns are tracked by studying vegetation changes as manifested by different
phytoliths in lake cores (Veena et al. 2014). Phytolith analysis also adds to forestry research
because once deposited, phytoliths contribute to the silica richness in soil systems (Blecker et al.
2006). Researching the silica cycle through how plants produce phytoliths provides insight into
silica availability and the role of leaf litter in forests (Umemura and Takenaka 2014).
Any analysis using phytoliths relies upon the ability to distinguish between the different
taxa or genera the phytoliths represent. Distinguishing between phytoliths of closely related taxa
4
relies on morphometric analysis, which is the measurement of the shapes and sizes of phytoliths
(Ball et al. 1999:1615-1616). Previous morphometric analysis has provided clear means for
distinguishing between the phytoliths of closely related taxa (Ball et al. 1999; Vrydaghs et al.
2009). For example, phytoliths have been used to distinguish between wild and domestic rice,
which adds to rice domestication research (Pearsall et al. 1995; Qui et al. 2014; Zhao et al.
1998). Phytoliths have also been used to distinguish between wild and domesticated millet in
Eurasia (Zhang et al. 2011) and among banana taxa (Vrydaghs et al. 2009).
There are several kinds of phytolith forms, some being produced only by specific species
and others being more universal. Some family, genera, and species are better producers of
phytoliths than others. Typically, monocots produce more phytoliths than dicots (Piperno
2006:37).
An example of a major producer of diagnostic phytoliths is the tribe Triticeae, found in
the grass family Poaceae (Pearsall et al. 1995:184; Sangster 1970:245; Twiss et al. 1969). Every
taxa in Triticeae produces inflorescence bracts called glumes, lemmas, and paleas. Phytolith
assemblages made up of articulated dendriform wave patterns are found in these bracts, and it
has been argued that these wave patterns can be used to distinguish between Triticeae genera and
taxa (Ball et al. 2015; Piperno 2006:76-78; Rosen 1992).
Other phytolith forms include rondels from grasses, sclerids and tracheids from vascular
tissue, and hairs and hair bases (Pearsall 2015:256-264). Another common form are amorphous
silica bodies. These indicate that silica was absorbed by the plant but that either the silicification
was incomplete or it does not reflect the shape of the parent cell (Pearsall 2015:254; Piperno
2006:24). In my study, I also expected to see plants that produce no phytoliths and plants that
produce several different phytolith forms.
5
Not all plants produce phytoliths, and there may be one or two species in a genus that
produce phytoliths, but the rest of the species within that taxa do not. There is no clear pattern
regarding which plants do or do not produce phytoliths and to what degree. The biology behind
why some plants are producers and others are not is not well understood. There are also
variations among plants on when phytoliths are formed. For example, the older a grass leaf is,
the more phytoliths will likely be produced (Piperno 2006:8, 15).
While botanists can predict where in a plant phytoliths may be found, the exact locations
are unknown until the various tissues of a plant have all been tested (Piperno 2006:18). In other
words, any patterns observed in phytolith production and frequency apply only to species and
plant tissues that have so far been analyzed, and such patterns can only be tentatively applied to
species and plant tissues that have not been analyzed (Piperno 2006:18-19). Still, most phytoliths
found within a given family, genus, or species will have similar shapes (Piperno 2006:24).
My thesis fills gaps in phytolith knowledge on particular species regarding how and
where they produce phytoliths, and then applies a regionally specific typology to archaeological
samples.
Thesis Organization
In Chapter 2, I present a cursory background review of the Fremont and what is known
about their plant use, followed by a historical review of ethnographies of the historic indigenous
occupation in Utah Valley. The ethnographic literature reviewed for the Fremont are of peoples
who shared similar geography, climate, resources, technologies, and subsistence strategies. The
“use [of] ethnographic data to establish reliable correlations between archaeologically observable
phenomena and archaeologically unobservable human behavior” (Trigger 2010:33) provides the
6
validity that some researchers need when making the statement that the Fremont utilized both
domestic and wild resources. The historic groups I review are the Shoshone, the Goshute, the
Ute, and the Southern Paiute.
In Chapter 3, I describe my methods: the nature of the Utah Valley Fremont botanical
reports that I reviewed and how I narrowed my data set to only be of plants with documented
ethnographic use. I also explain the phytolith extraction methods I used to create the reference
typology as well as provide a brief overview of the ground stone tool wash methods used, and
the slide analysis methods employed.
In Chapter 4, I present the results of the phytolith typology by plant life form: forb, trees
and shrubs, and grasses. I also discuss probable diagnostic types. Detailed results are also
presented in Appendix D. In Chapter 5, I compare the results of my ground stone tool slide
identification to the results of Yost (Appendix F). I review what could be interpreted from those
results, followed by a summary of the results overall. In Chapter 6, I discuss my research and
conclusions. Specifically, I review how ethnographic data can provide interpretations for plants
identified in archaeological contexts.
7
2. Background
Before I explain my methods, I provide background information on the Fremont and the
Shoshone, Goshute, Ute, and Southern Paiute.
The Fremont
The Fremont were a geographically widespread people living in diverse environments
across most of Utah. They practiced a shared, albeit flexible, subsitence method of farming, and
possessed distinct cultural traits, likely influenced by Southwestern cultures (Jennings 1978:150,
155; Madsen 1989; Madsen and Simms 1998; Martwitt 1986:161). Early scholars of the Fremont
were Edward Palmer, Henry Montgomery, Neil Judd, Julian H. Steward, Noel Morss, and Jesse
Jennings. These researchers recognized that while the Fremont as a whole practiced farming, that
there were regionally unique variations of that farming (Janetski 2008). These regions, or
regional traditions, have been defined as macro-regions of resource abundance: northern, central,
southern, Uinta, and Snake Valley (Janetski and Talbot 2014:122; Madsen 1989). Trait lists,
which run the risk of being too restrictive or overly general, do provide a foundation for the
characterization of the Fremont as a cultural group (Talbot 2000; Searcy and Talbot 2015).
General diagnostic, material culture traits of the Fremont are the Utah metate, clay
figurines, one-rod-and-bundle basketry, the Fremont moccasin, and bone gaming pieces
(Jennings 1978; Madsen 1989; Searcy and Talbot 2015; Simms 2010). Indirect evidence of
social and ceremonial life comes from gaming pieces, large kill sites, exotic goods, rock art,
mutilated human bones, burial practices, public architecture, and villages (Janetski 2008; Searcy
and Talbot 2015; Simms 2010). The material cultural variants of the different regions and
8
borderlands have been based on ceramics, subsistence strategies, style variation in rock art and
figurines, and architecture (Janetski and Talbot 2014:122; Searcy and Talbot 2015).
An example of architectural variation includes differences between the west and east. In
western Utah, late Fremont (post-900 AD) residences were typically pit houses often with
adjoining or nearby adobe-walled storage structures, or surface residences and granaries made of
adobe blocks (Janetski 2008). In eastern Utah, residences varied more, with living quarters and
storage structures made of stone and mud masonry, or pit houses with slabs (Janetski 2008).
Overall, late Fremont villages and hamlets were often located in ideal farmland, and residential
structures may have been long-term. While several sites are located near water such as mountain
runoffs and streams, other sites are in areas where local stories tell of once extant prehistoric
ditches, suggesting that some Fremont irrigated their maize crops (Metcalfe and Larrabee
1985:244).
Maize first appears in Fremont contexts as early as AD 1, but it is not until AD 600-700
that maize seems to play a more integral role in Fremont diet (Allison 2008; Talbot 2000; Winter
1973). “The spread of agriculture north of the Colorado River was likely accomplished, at least
in part, by low- level migration of Basketmaker farmers over several centuries…This was
facilitated by relatively low populations of foragers, and current thinking is that over time some
foragers acculturated into this farming society” (Searcy and Talbot 2015:239). Evidently
subsistence did not depend on maize alone, but on several different strategies that utilized wild
resources, domesticates, cultivatable land, and trade (Janetski 2008; Search and Talbot 2015;
Talbot 2000).
These different subsistence strategies are referred to by some as Fremont adaptive
diversity, where a strategy is made up of various subsistence patterns and/or where a group
9
utilizes multiple strategies (Madsen and Simms 1998; Simms 1986). These strategies include
committed farmers who practiced varying degrees of hunting and gathering, seasonally mobile
farmers, foragers who had access to maize, and perhaps mobile hunter-gatherers who while not
wholly Fremont, extensively traded with the Fremont (Allison 2008; Simms 1986). Regarding
these last three, “it is often difficult, archaeologically, to detect a difference between farmers
occasionally acting as foragers and foragers and farmers acting interdependently, since many of
the tools and even the rock art symbols they employ are the same” (Madsen and Simms
1998:286). This adaptive strategy view of the Fremont proffered by Madsen and Simms (1998)
has been challenged more recently, especially as archaeologists work to better identify
diachronic shifts in cultural practices, including evidence that suggest farming was adopted more
widely by Fremont groups after AD 900 (Janetski and Talbot 2014; Searcy and Talbot 2015).
Searcy and Talbot (2015:252-253) include agricultural shifts and artifacts manifesting a Late
Fremont tradition as evidence for a shared heritage that was maintained by social distance from
non-Fremont groups. “Evidence for this shared heritage is embedded in stylistic commonalities
in artifacts, rock art, architecture, and farming” (Searcy and Talbot 2015:241).
At the same time, diversity in transportable, regional variants of material culture like
ceramics, architecture, and subsistence strategies suggest varied and regional adaptability to the
environment (Janetski and Talbot 2014:118). The region I explore in this thesis is Utah Valley,
roughly an 823-square mile valley, with Utah Lake comprising 148 square miles, located in Utah
County, south of Salt Lake County (Janetski 2008) (Figure 2.1).
Prior to European occupation in the 1800s, Utah Lake in Utah Valley played an
important economic role due to an abundance of native fishes, many of which are now extinct
(Janetski 1990b). Wild, edible plants such as chokecherries, and game animals such as jack
10
rabbits, were plentiful in both the uplands and lowlands of Utah Valley, providing prehistoric
peoples with many natural resources (Janetski 1990b). Three archaeological periods have been
identified in Utah Valley: the Archaic, Fremont, and Late Prehistoric. The Fremont people, the
best archaeologically documented in Utah Valley, gathered, hunted, and farmed (Janetski
1990b). Numerous Fremont sites have been found and documented since the early 1900s;
however, little attention was initially given to subsistence in this region by these researchers
(Janetski 1990b).
Evidence of farming comes from Zea mays kernels, pollen, starches, and cops found at
nine Utah Valley sites: American Fork Cave, the Hinckley Mounds, Kay’s Cabin, Seamon’s
Mound, Smoking Pipe, Spotten Cave, West Canyon, Wolf Village, and Woodard Mound (Table
2.1). Beans, also evidentiary of farming, are only found at three sites: Smoking Pipe, Spotten
Cave, and Wolf Village. Evidence of foraging include botanical remains of pinyon pine, juniper,
ephedra, and various wild grasses. Bones of fish, small and large game animals, and waterfowl
have been found at the sites previously noted.
Both horticulturalists and hunters-gatherers benefited from living near or along Utah
Lake, and it is likely that all groups utilized the wetland resources in similar ways (Janetski
1990a). While exploitation of wetland resources was much the same, Fremont living nearer the
wetlands practiced different subsistence strategies than Fremont living away from wetlands
(Janetski 1990a). The subsistence practices of the Fremont in Utah Valley are, in large part, due
to the rich wild resources available around Utah Lake.
Several types of plant remains have been found at Utah Valley Fremont sites: pollen,
phytoliths, starches, charcoal, charred and uncharred seeds, uncharred leaves and other plant
fragments (Table 2.1). There are also a few sites where no plant remains have been identified due
11
to poor preservation conditions, such as damp soil near Utah Lake. These sites include the early
excavations of Provo Mounds (Crellin 1967; Connor 1967; Madsen 1969; Miller 1969).
Between 1250 and 1400 AD, the architectural and material culture defining
characteristics of the Fremont disappear archaeologically (Madsen 1989:14). There are several
theories used to explain why the Fremont disappeared between AD 1250 and 1350 (Allison
2010:148; Madsen 1989:14; Madsen and Simms 1998:313; Martwitt 1986:171-172). One
suggestion is that there was an overall social disruption felt throughout the southwest which
included the southern neighbors of the Fremont such as the Pueblo. There may have been a
changing climate that was not conducive to farming and as such forced the Fremont to move or
change their subsistence strategies and lifeways. The Fremont may have been displaced by
Numic-speakers expanding into the area from southern California, or they left prior to this
Numic expansion. Regardless of what happened to the Fremont, what is known is that around
AD 1300, farming ended, depopulation likely occurred, and hunter-gatherers with links to the
historical Ute and Southern Paiute moved into the Fremont and Virgin areas (Allison 2010:150;
Jennings 1978:235).
12
Figure 2.1. Map of Utah Valley. Showing Spotten Cave, Woodard Mound, Wolf Village, Utah
Lake, and Utah Valley (Museum of Peoples and Cultures, Provo. Courtesy Scott Ure).
13
Table 2.1. The Frequency of Archaeobotanical Remains at Fremonts Sites in Utah County.
14
Family
Subfamily, genus
Adoxaceae
Amaranthacae
Amaranthaceae
Amaranthaceae
Amaranthaceae
Amaranthaceae
Apiaceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Betulaceae
Betulaceae
Boraginaceae
Boraginaceae
Brassicaceae
Brassicaceae
Brassicaceae
Caryophyllaceae
Caryophyllaceae
Cleomaceae
Cucurbitaceae
Cupressaceae
Cupressaceae
Cyperaceae
Cyperaceae
Ephedraceae
Euphorbiaceae
Fabaceae
Sambucus
Suaeda
Amaranthus
Atriplex
Cheno-ams
Chenopodium
none specified
Ambrosia
Artemisia
Chenopod. berlandieri
Cirsium
Helianthus
High spine
Iva axillaris
Liguliflorae
Low spine
none specified
Taraxacum
Alnus
none specified
Amsinckia
Cryptantha
Brassica
Lepidium
none specified
none specified
Silene
Cleome
Cucurbita
Juniperus
Juniperus monosperma
none specified
Scirpus
Ephedra nevadensis
Euphorbia prostrate
none specified
American
Fork
Cave
-
Hinckley
Mounds
Kay's
Cabin
Seamon’s
Mounds
Smoking
Pipe
Spotten
Cave
West
Canyon
Wolf
Village
Woodard
Mound
Sum
S
S, P
S
P
P
P
S
P
P
P
S, P
P
S
-
S
S
S
S
-
T
-
P
O, P
P
P
P
P
P
P
P
P
P
O
-
S
S
S
O
S
-
-
S
S
S, P
S
P
S
S
P
P
P
P
S
S
S
P
S
S
S, P
P
S
P
S
P
S
S
S
P
P
S
S
S, P
P
S
S
1
1
2
2
5
3
2
1
3
1
1
3
3
1
2
3
2
1
2
1
1
1
1
1
3
1
1
2
1
5
2
3
3
1
1
2
Table 2.1. contd., The Frequency of Archaeobotanical Remains at Fremonts Sites in Utah County.
15
Family
Subfamily, genus
Fabaceae
Malvaceae
Nyctaginaceae
Papaveraceae
Pinaceae
Pinaceae
Plantaginaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Polygonaceae
Polygonaceae
Polygonaceae
Polygonaceae
Polygonaceae
Portulacacea
Quercus
Ranunculus
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Salicaeae
Sapindaceae
Sarcobataceae
Solanaceae
Solanaceae
Phaseolous
Sphaeralcea
Boerhaavia
Argemone
Abies
Pinus
Plantago
Eragrostis
Hordeum/Elymus
none specified
Panicum
Phragmites
Sporobolus
Stipa hymenoides
Zea mays
Eriogonum
Polygonum
Polygonum bistortoides
Poly. lapathifolium
Rumex
Portulaca
Quercus
none specified
Ameliancher
none specified
Prunus
Prunus virginiana
Rosa
Rubus
Salix
Acer
Sarcobatus
none specified
Physalis
American
Fork Cave
S
-
Hinckley
Mounds
p
S
S, P
S
S
S
P
P
P
P
P
-
Kay's
Cabin
S
S
S
-
Seamon’s
Mounds
T
Y
T, Y
-
Smoking
Pipe
S
P
P
P
P
S, P
P
P
P
P
P
P
-
Spotten
Cave
S
S
O
S
S
West
Canyon
O
-
Wolf
Village
S
S
S
P
P
S, P
P, T
S
S
S
S, P
S, P
P
S
S
S
P
S
S
P
P
S
Woodard
Mound
P
S
S
S, P
S
P
P
-
Sum
3
1
1
1
2
4
1
1
1
5
1
1
1
2
9
2
3
1
1
2
1
1
1
0
3
1
1
1
1
3
3
3
1
2
Table 2.1. contd., The Frequency of Archaeobotanical Remains at Fremonts Sites in Utah County.
Family
Solanaceae
Typhaceae
Typhaceae
Total
Subfamily, genus
Solanum jamesii-type
Typha
Typha latifolia
American
Fork Cave
1
Hinckley
Mounds
S, P
P
26
Kay's
Cabin
P
8
Seamon’s
Mounds
T
5
Smoking
Pipe
P
25
Spotten
Cave
10
West
Canyon
1
Wolf
Village
S
P
47
Woodard
Mound
P
19
Sum
1
3
4
142
Key: O = fragments, other; P = pollen; S = seed, kernel, both whole and fragment; T = starch; Y = phytolith
Note: Table created from American Fork Cave (Hansen 1941), the Hinckley Mounds (Peterson 2016; Puseman 2016), Kay’s Cabin (Puseman
and Cummings 2001), the Seamon’s Mound burials (Yost 2009), Smoking Pipe (Billat 1985; Forsyth 1984; Scott 1984), West Canyon
42UT119 (Wheeler 1968), Wolf Village (Dahle 2011, Cummings 2011), and Woodard Mound (Richens 1983).
16
Ethnographies of Historic Indigenous Occupation in Utah Valley
Statements by researchers that the Fremont utilized domestic and wild plant resources in
their diets are tenuous when only based on archaeological findings. For example, the presence of
charred seeds from wild plants found in the fill of sites are often used as evidence for dietary and
other plant use. These seeds, though, could be charred because they were part of kindling, there
was a nearby natural fire, or they were charred for ceremonial reasons. Another example is
pollen of wild plants found on groundstone and other vessels. Archaeologists have yet to develop
infallible methods that differentiate between pollen rain and pollen from plants intentionally
brought to a site. Wild plant starches and phytoliths in teeth calculus could come from an
individual using their teeth as a tool to process plant materials. The presence of plant material in
coprolites and of carbon isotope ratios indicative of wild plants found in bone collagen are more
difficult to dismiss as being caused by natural processes. Unfortunately, coprolites are far and
few between and carbon isotopes do little more than define if the carbon was from a C3 or a C4
source. The coupling of archaeobotanical findings with ethnographic sources strengthens
statements regarding the utilization of domestic and wild plant resources in prehistoric diets.
There is no known group of prehistoric peoples in the New World that relied entirely
upon domesticates. All prehistoric groups utilized both wild plants and domesticates. The degree
to which they relied on these plants varies group to group. To survive, prehistoric groups had to
rely upon wild resources as much as, and in some cases more than, domesticated resources.
Knowing what these resources were is difficult to tease out of the archaeological record because
of the poor preservation of organic material. Understanding how and why wild plant resources
were used can be just as difficult.
17
I reviewed numerous ethnographic sources to determine plant use by more recent
indigenous groups to identify ways in which plants may have been used for medicine and food. I
applied my findings to the results of my ground stone phytolith analysis to better understand if,
how, and why the Fremont may have used plants (see Chapter 5 and 6).
For my ethnographic analogy, I focused on the Shoshone, Goshute, Ute, and Southern
Paiute, because they lived in Utah Valley, nearby, or in similar geographical and environmental
terrains. Sources on these historic groups come from Spanish exploration in the late 1700s, fur
trappers in the early 1800s, followed by government explorers, immigrants and settlers in the
mid-1800s, Indian agents in the later 1800s, and ethnographic studies in the early 1900s (Janetski
1983:28). The first to observe and record plant use by the indigenous groups living in the Great
Basin include Silvestre Velez de Escalante in 1776, John C. Fremont in 1844 (Fowler 2000:100,
102); and in Utah, Jedidiah S. Smith, Daniel T. Potts, and William Clayton.
Silvestre Velez de Escalante explored the Great Basin in 1776. He witnessed the
consumption of black Manzanita berries by the Utes. He also observed the Timpanogotzis Ute,
or Fish Eaters, in Utah Valley gather seeds that they turned into gruel (Velez de Escalante
1995:27, 72).
John C. Fremont (1846:172), while traveling through Utah Valley in May of 1844,
observed that there were two kinds of people which inhabited the valley: the Diggers and the
Fish Eaters. The Diggers were “miserable and sparsely peopled,” eating seeds and roots, living in
single family units (Fremont 1846:172), whereas the Fish Eaters were of a seemingly higher
social status. Wild sage was the only wood in the valley, and Fremont (1846:172) thought the
valley ideal for grazing. Fremont found yampah roots to be the most agreeably flavored for
18
eating, and noted that Convallaria stellata (false Solomon’s seal) was considered the best
remedial plant among Indians, but he does not specify for what (Fremont 1846:87, 170).
On his expedition through Utah from 1826-1827, Jedidiah S. Smith noted plants and
animals used by the Ute surrounding Utah lake. The Ute harvested service berries and, when
there was no game to hunt, dug roots for food (Brooks 1977:44, 45). Smith later encountered the
Paiute and remarked that they seemed to subsist entirely on roots, which were dried and mashed
into cakes (Brooks 1977:49). He also interacted with the Goshute, whom he called the “children
of nature… [because of their] connecting link between the animal and intellectual
creation…quite in keeping with the country in which they are located” (Brooks 1977:185). He
did not mention, though, how they used plants.
An acquaintance of Smith, Daniel T. Potts, in a letter dated July 8, 1827, said that those
around and especially south of “Utaw Lake” lived in structures of bulrushes as there were no
trees, and that their diet consisted of roots, grass seeds, and grass (Bagley 1964:136-137). Some
of these groups even called themselves “Pie-Utaws” (Bagley 1964:137). An early Utah pioneer,
William Clayton, noted in his journal in 1847 that the Indians south of Utah Lake “raise corn,
wheat, and other kinds of grain and produce in abundance,” including pumpkins (Clayton
1921:278).
The Shoshone
Western Shoshone territory was vast, ranging from Death Valley, California, to Tooele
Valley in northwestern Utah. The environment and resource availability of this territory was
diverse, and boundaries between different tribes were often fluid (Thomas et al. 1986). Dietary
and medicinal plant use among the Western Shoshone was perhaps more diverse than that of the
19
Ute (Thomas et al. 1986). They relied heavily on foraged plants, but also hunted and
occasionally farmed (Lowie 1924; Thomas et al. 1986). Some of the first frontiersmen to interact
with the Shoshone were Jedidiah Smith in 1827 and John C. Fremont in 1845. During early
Euro-American contact, 43 different Shoshone subgroups were said to have existed at one time
(Thomas et al. 1986).
Percy Train, James R. Henrichs, and W. Andrew Archer visited with several Nevada
Shoshone informants about medicinal plant use from 1937-1941 (Train et al. 1941). They
recorded diverse uses of 200 plants, relying on the knowledge of older Shoshone, especially
those credited with extensive medicinal plant knowledge (Train et al. 1941). They estimated that
around 1,700 Shoshones then lived throughout Nevada in various colonies and reservations
(Train et al. 1941). The love of and close contact to mountains and mountain flora likely led to
the Shoshone possessing a wider knowledge of medicinal plants than the other contemporary
Indian groups (Train et al. 1941).
In the 1900s, Robert Lowie was one of the first to study the Northern Shoshone (Fowler
2000). The Wind River Shoshone of Wyoming did not farm, but were largely dependent on
vegetable foods (Lowie 1924). They hunted bison, mountain sheep, groundhogs, and other small
and large game (Lowie 1924). Women gathered roots, carrots, chokecherries, and other berries,
and used metates to grind their seeds (Lowie 1924). They raised tobacco which they smoked
along with substitutes such as kinnikinnick (Lowie 1924).
The Goshute
The Goshute, formerly spelled Gosiute, while a separate landholding entity, have been
considered the impoverished cultural and ecological cousins of the Western Shoshone (Thomas
20
et al. 1986:262). Bands that are known to have traveled and/or lived in Utah are the Tooele
Valley Goshute, Rush Valley Goshute, Cedar Valley Goshute, Skunk Valley Goshute, and Trout
Creek Goshute (Thomas et al. 1986). Their lands and lifeways were deeply impacted by the
Mormon settlers through displacement and the introduction of diseases (Thomas et al. 1986:263).
Ralph V. Chamberlin studied the Goshute in the spring of 1901, and later in 1905
(Chamberlin 1909; Fowler 2000). He is credited as being one of the first to attempt “a thorough
treatment of the uses of plants by a single Great Basin group” (Fowler 2000:103). One of his
informants in 1901 was an Uintah Ute man named Tungaip who was living with the Goshute and
who taught him several Ute names and Ute uses of plants. Chamberlin (1909:27) remarks that
“[t]heir dependence upon the vegetable kingdom was, naturally, less intimate than with such
tribes as the desert-dwelling Goshute.” Much of the Ute terminology corresponded to Shoshone
and Goshute vocabulary, yet Chamberlin notes that Tungaip was the only Ute he consulted
(Chamberlin 1909). In his later work in 1905, he more intimately studied Shoshonean plant use
and catalogued over 300 plant species (Fowler 2000:103).
The Goshute once lived in all the desert territory along the southern and western land
bordering the Great Salt Lake (Chamberlin 1911). The environment and geography of this area
includes mountain ranges interspersed with valleys of alkali flats, playas, and grasses
(Chamberlin 1911). Plants often found in these terrains included the common greasewood,
cheno-ams of various types, as well as junipers, pinyons, and many herbaceous and fruit bearing
plants (Chamberlin 1911). Prior to the arrival of Mormon pioneers, Goshute numbers were said
to have been in the thousands, but the introduction of foreign diseases, such as measles, reduced
these numbers (Chamberlin 1908). When Chamberlin visited with them in 1901, he found that
they had been living in Skull Creek and Toole County, Utah, for many years and that their
21
numbers had dwindled so much that they were no longer considered a tribe but a band
(Chamberlin 1913).
Despite their low numbers, Chamberlin (1913:2) believed the Goshute to be essentially
self-sustaining. The Goshute hunted a wide range of animals, such as antelope, deer, ground
squirrel, crickets, and rabbit (Chamberlin 1911). The large game animals were used for food,
blankets, and clothing. However, the main Goshute food source was plants, as their nickname
“Root Diggers” suggests (Chamberlin 1911:337). Medicines for ailments such as bruises, burns,
and colds, materials for household supplies, and so forth, were all primarily derived from plants
(Chamberlin 1911).
The Utes
The Utes were rapidly displaced in the 1850s, and so any records after the 1850s are
tainted by displacement and poor memory (Janetski 1991). Sources on the Ute come from
Spanish explorers, fur trappers, government explorers, immigrants and settlers, Indian agents,
and ethnographic studies, ranging from 1775-1940 (Janetski 1991). Escalante was one of the first
to acknowledge cultural differences among the Ute tribes (Smith 1974). Ute origins stem from
the Numic-speaking people, which includes the Shoshone, and spans a geographical area from
California to the Rockies (Smith 1974).
In general, the Ute primarily inhabited areas in central and northeast Utah and western
Colorado; they hunted and fished, as well as utilized wild plant resources (Smith 1974). The Ute,
though, are not a homogenous people. They share similar traits, such as beliefs regarding
marriage, death, and kinship, but ultimately there are distinct differences among the various
tribes, such as the Uintah, Pahvant, Sanpits, Moanunts, Seuvarits, and Timpanogots of Utah
22
Valley (Janetski 1991; Smith 1974). For example, Western Ute bands, such as the Timpanogots,
had access to more roots, nuts, lilies, and berries than their Eastern counterparts who had access
to more grasses (Callaway et al. 1986:337). Differences in geography, geology, and ecology, has
led to differences and diversity among the Ute bands. Boundaries of these groups were not static,
but shifted with different events, history, and climate (Janetski 1991).
The Timpanogot Ute were defined by their proximity to and use of Utah Lake, also
referred to as Timpanogot Lake (Janetski 1983). They were often referred to as “Fish Eaters”
because of their heavy reliance on and exploitation of lacustrine resources from Utah Lake
(Janetski 1991). They were territorial regarding usufruct rights to resources within a geographic
region. They cured ailments through shamans, songs, and smoke (Janetski 1991). The
Timpanogots crafted objects out of wood, bone, antler, clay, stone, and animal hide, including
beads, baskets, buckskin shirts, ceramics, and arrowheads (Janetski 1991).
The Timpanogots subsisted by fishing, hunting, and gathering a variety of foods, berries,
nuts, seeds, roots, and greens, including tobacco, several of which they stored for the winter
(Janetski 1991). Waterfowl, sage grouse, ground squirrel, new shoots and roots were spring
foods, and fish were both a spring and summer food (Janetski 1983). In the summer they
consumed waterfowl, grasses, weeds, sunflowers, bulrushes, berries, and insects (Janetski 1983).
During the fall, migrant waterfowl, pine nuts, large game, and rabbits were part of their diets
(Janetski 1983). Then in winter, the Timpanogots ate cached foods, elk, deer, and bison (Janetski
1983).
They appear to have lived in “numerous, small, essentially permanent villages located
along the lower reaches of [Utah Lake] feeder streams and the eastern shores of Utah Lake”
(Janetski 1983:67). In the smaller, seasonal camps located throughout the valley, dwellings were
23
domed wikiups or willow houses. During the winter, they were along river bottoms and in the
spring, they were south of the lake (Janetski 1983). The archaeological presence of Ute in Utah
Valley are manifest in American Fork Cave, the Beely Site, Spotten Cave, and the Spencer Site
(Janetski 1983).
Another tribe, the Ute of Navajo Springs, of Ignacio, Colorado, and of Whiterocks, Utah,
hunted large and small game, including eagles and rabbits, and fished (Lowie 1924). They
gathered grass seeds, berries, and chokecherries, which were dried and cached for winter. They
used metates to grind their seeds, and often employed different types of metates (Lowie 1924).
Another example of Ute dependence and use of plant foods come from the northern Ute
group, the Uintah, and also the Uncompaghre and White River of northwest Colorado. Details of
their lives come from oral histories of individuals living during the 1850’s-1880’s (Smith 1974).
At this time, the northern Utes were no longer living on their native lands, having been driven
out by Mormon settlers, and eked out a day-to-day living (Smith 1974). These Utes had lived on
what may be considered the most productive land, where they practiced traditional hunting,
fishing, and gathering.
Regarding plant use, roots were collected, berries, leafy tops, and greens were gathered,
pine nuts were popular, and quaking aspen tree sap was a delicacy (Smith 1974). Specifics on
plant species names does not exist because Smith’s plant collection was destroyed in a car crash
(Smith 1974). What plants she does identify are by their common name: blackberry, blueberry,
buffalo berry, chokecherry, currant, gooseberry, juniper berry, raspberry, service berry, squaw
berry, strawberry, rose hips, wintergreen, pinyon nuts, wild onion, Indian potatoes, edible roots,
yampa, sego lily, and various seeds (Smith 1974). Plants also held cultural importance. For
example, food taboos were common for pregnant women, such as yampa, which was said to
24
cause miscarriages, and eating beaver would prevent the water from breaking during labor.
Another example are menstrual huts, which were made of willow in summer and cedar in winter
(Smith 1974). Plants were also commonly used medicinally (Smith 1974). Other ways plants
were used were for baskets, cordage, and in pipes for smoking (Smith 1974:118).
The Southern Paiute
Some speculate that the Paiute-Shoshone arrived in southern Utah between AD 1100 and
1200 (Holt 2006). The first record of European contact with the Southern Paiutes was by
Escalante and Dominguez in October of 1776 (Holt 2006). In some instances, these records did
not clearly distinguish between where the Ute ended and the Southern Paiute began; this is
partially due to close cultural similarities (Euler 1966; Kelly and Fowler 1986).
Early explorers found the Paiute to be peaceful foragers and horticulturalists (Holt 2006).
Slave trade was a concern and fear among the Paiute. Despite their apprehensions, the Paiutes
welcomed the first Mormons to Utah, providing them with food. Regrettably, these relations
soured and many Mormons turned hostile or ambivalent to the Southern Paiute (Holt 2006).
Many Mormon settlements, such as St. George, were on Paiute campsites and displaced the
Paiute. By the 1860s the Paiute were “destitute and hungry. Whites were pouring into their land,
and they could do nothing to stop them” (Holt 2006:34). While Mormon and federal government
rhetoric was that of creating Paiute self-sufficiency, the opposite resulted from policies made by
both groups (Holt 2006:xvii). Their ability to feed and care for themselves faltered as traditional
resources were used up by the Mormons. Some traditional resources were still available, such as
pine nuts, jackrabbits, and other wild plants (Holt 2006).
25
The climatic variation and varied ecologies of southern Utah initially led to Southern
Paiute adaptive diversity: seminomadic mobility and seasonal migration, winter base camps,
wild and domestic resources, windbreaks and brush shelters, and so forth (Euler 1966:13-14;
Holt 2006; Kelly and Fowler 1986:371). They would travel by foot and often camped adjacent to
water and juniper stands. To maintain mobility, tools and other items were often not complex in
design or construction (Holt 2006). Clothes were made predominately from animal hides, both
twined and coiled baskets were made, but not all groups made pottery (Kelly and Fowler 1986).
The Southern Paiute subsisted by hunting, gathering numerous plant foods and
cultivating native plants such as corn, squash, beans, and sunflower (Euler 1966:33; Kelly and
Fowler 1986). They hunted small game, birds, insects, and large game, and fished where possible
(Kelly and Fowler 1986:370). Prickly pear, potatoes, reeds, berries, melons, and wild grapes
were also important food items (Euler 1966).
Botanical explorer Edward Palmer collected several seed samples from the Southern
Paiute in the 1870s; however, his samples were lost for some time and only a portion have
resurfaced (Bye 1972). There are at least sixty plants in the relocated portion of the collection,
each reportedly to be from plants used as food and/or medicinally. These plants include
domesticates such as beans and squash, and wild plants such as amaranths, grasses, yucca,
sunflower, and cliff rose (Bye 1972).
Conclusion
The presence of a plant in archaeological contexts does not always mean that the plant
was used or consumed, unless found in a coprolite. Some plants, such as corn, were clearly
cultivated to be consumed by prehistoric peoples. Yet the use of native plants by prehistoric
26
peoples is more difficult to interpret (Barlow and Metcalfe 1996; Doebly 1984). The study of
archaeobotanical remains can indicate what plants were available at a site, and a review of
ethnographic reports can provide interpretations for how and why those plants may have been
used.
To provide some interpretation for the presence of plants in archaeological contexts, I
have created a geographically-specific ethnographic comparison between the prehistoric Utah
Valley groups and historically documented peoples who lived near, around, or in Utah Valley. I
wanted a narrow data set where there would be multiple shared traits between the groups because
the fewer shared traits between the “ethnographic source and the prehistoric subject,” the greater
the inability to expect them to have other traits in common (Wylie 1985:94, 98).
I believe my interpretations are viable because some plants, such as corn, pinyon pine,
and sunflowers, have inherent attributes that make their use easier to identify (Bye 1985:376).
These plants would likely be used for food regardless of scarcity or abundance (Barlow and
Metcalfe 1996; Bye 1985; Doebly 1984; Heiser 1951). Moreover, the study’s geographically
narrow focus suitable considering the focus on Utah Valley Fremont. The Fremont, for example,
are archaeologically unique and distinct from the Hohokam, the Patayan, and the Mississippian
mound builders, and the same can be said of the Ute, the Apache, and the Sioux. It is likely,
therefore, that while the interpretations presented are weakened by the separation of time, they
are of value because of the shared geography, climate, and resources, and similar technologies,
and subsistence strategies between prehistoric Utah Valley peoples and historic Utah Native
Americans.
27
3. Methods
The methods for this thesis included building a phytolith data base from plants with
documented ethnographic uses identified from Fremont sites. I then tested the phytolith data base
by analyzing groundstone from Wolf Village. The steps involved in the creation of my phytolith
typology began with deciding parameters, specifically what kinds of plants would I sample and
why. Utah Valley is home to hundreds of native plant species. To study all the plants in the
Valley would require several years of full-time research. Such a study, in some ways, would also
not be entirely productive because it is likely that not all plant species possessed the same
economic, dietary, medicinal, or spiritual value for the Fremont.
The plants I chose at the beginning of my research were those that likely had dietary and
medicinal value. These plants were determined by reviewing ethnographic reports of historical
groups who lived in and around Utah Valley after AD 1300 to the late 1800s. By choosing such
plants and these groups, my goal was to create a bridge “to establish reliable correlations
between archaeologically observable phenomena and archaeologically unobservable human
behavior” (Trigger 2010:33). The unobserved human behavior was Fremont plant use, and the
observed phenomena is evidence of these plant remains at Fremont sites.
Fremont Botanical Reports
To begin, I reviewed Utah Valley Fremont botanical reports to create a data set of all
identified plants from archaeological sites. These reports are associated with the following sites:
American Fork Cave (Hansen 1941), Kay’s Cabin (Puseman and Cummings 2001), Hinckley
Mounds (Peterson 2016; Puseman 2016), the Seamon’s Mound burial (Yost 2009), Smoking
28
Pipe (Billat 1985; Forsyth 1984; Scott 1984), Spotten Cave (Pearce 2012), West Canyon
42UT119 (Wheeler 1968), Wolf Village (Dahle 2011, Cummings 2011), and Woodard Mound
(Richens 1983) (Figure 2.1). With a few exceptions, all analytical reports of macrobotanicals,
pollen, phytolith, and starches were included in the data set (Appendix A). These exceptions are
as follows.
Uncharred macrobotanicals were not included because unless preservation conditions are
favorable, such seeds are more likely modern than prehistoric (Minnis 1981:147).
Uncharred seeds from the Spotten Cave coprolites, however, were included because the
coprolites are known to be prehistoric.
Pollen from floors, hearths, and middens were not included because these samples are not
a clear indicators of plant consumption (Bryant and Holloway 1983; Dahle 2011:26;
Pearsall 1989).
Pollen from the fill of Smoking Pipe was included because of the few botanical remains
from this site.
Charcoal was excluded because it is unclear if the plant source was used for food or
medicine.
All phytolith and starch reports were included because they are few in number.
Plant Species Native to Utah County and Documented Ethnographic Uses
I used Welsh et al. (1987, 2008) to identify the species within the genera and families I
had recorded and evaluated which of those species were native to Utah Valley. The reason
genera and families were expanded to include species was to better identify which plants to
collect for the phytolith typology. The creation and uniqueness of phytoliths can vary from
29
species to species within a genera or family. Therefore, individual species of a genus or family
should be analyzed separately.
I did not include species that were used for ornamentation or that were represented in the
valley by one specimen. Species that were adventive from outside the Americas were also not
included because they likely come from European colonization. Adventive species from Central
and South America were included, though, because the northward introduction of plants like
corn, beans, and squash into the Great Basin likely included other plants.
To narrow my data set further, I examined how peoples living in similar ecological and
environmental terrains used the plants on my list for food and medicine. I also researched what
part of the plant they used. I researched documented Native American groups that would have
lived in, nearby, or traveled through Utah Valley, and had access to the same plant resources
(D’Azevedo 1986). These groups, the Shoshone, Goshute, Ute, and Southern Paiute, also
possessed similar technologies to the prehistoric Utah Valley inhabitants. In addition to the
ethnographies, Fowler (1986), Palmer (1878), and Yanovsky (1936) provide further insight into
how plants were used. My methods are like those employed by Rainey and Adams (2004), who
created an American Southwest ethnobotanical database used for interpreting archaeological
sites excavated by the Crow Canyon Archaeological Center. Regarding their work, they state:
The purpose of this work is to summarize information from published and unpublished
ethnographies that document how Native peoples of the American Southwest used—and,
in some cases, continue to use—selected plant resources. The data contained herein have
been used to suggest and support interpretations of archaeobotanical remains recovered
from [archaeological] sites[.] [Rainey and Adams 2004]
30
All species in Appendix A that are native to Utah Valley, whether they have documented
ethnographic uses or not, are included in Appendix B. A detailed list of how and why specific
plants were used, and by whom, is in Appendix C. These data are incomplete because of the
possibility that some plants listed in the ethnographic sources were not included due to plants
having multiple scientific and common names (Table 3.1). In addition, there are many plants for
which there are no ethnographic records, and there are ethnographic records I have not yet
reviewed.
Table 3.1. Examples of Changes in Plant Nomenclature.
Previous or Alternative Family Name
Family Name Used
Chenopodiaceae
Amaranthaceae
Sarcobatus was in Chenopodiaceae
now in Sarcobataceae
Umbelliferae
Apiaceae
Compositae
Asteraceae
Graminae
Poaceae
Capparaceae
Cleomaceae
Leguminosae
Fabaceae
Plant Collection and Digestion
After I compiled the lists of plants that had been identified at Utah Valley Fremont sites,
which species were available in Utah Valley, and of those which had documented ethnographic
use, I collected samples. I collected plant specimens from winter 2014 through the summer of
2015. This involved several visits to the BYU Herbarium, managed by the BYU Monte L. Bean
Life Museum, and with permission from the curator, Robert Johnson. I was only allowed to
sample material from the herbarium if it was loose, meaning the plant material had broken off
the original sample. The summer 2015 sampling involved visits to several nurseries, gardens, and
native locales in Utah and Salt Lake Counties. I called ahead and acquired permission before my
31
sample trips. At these centers, I contacted botanists to assist in the identification of plants. I
collected 53 of the plants listed on my data set. I was unable to collect more due to time
restraints.
I collected plant specimens from Red Butte Gardens in Salt Lake, Sego Lily Gardens in
Draper, Water Wise Nursery in Salt Lake City, Thanksgiving Point Gardens in Lehi, Central
Utah Gardens in Orem, the BYU Herbarium, and from the following wildlife and recreation
lands: Nine Mile Canyon, Spanish Fork Canyon, and Provo Canyon.
In fall 2015, Dr. Terry Ball trained me in the digestion of plant materials (see Portillo et
al 2006). Individual plant tissues or parts were treated separately through the entire digestive
process. For example, the berries and leaves of Shepherdia canadensis (buffalo berry) have
documented ethnographic uses, and as such the berries and the leaves were both analyzed, but
separately. However, due to two problems—hardy plant material and diatoms—and access to
different resources, my methods varied slightly.
Plant tissues were sonicated in a Mettler Electronics Ultrasonic Cleaner for five minutes
with Micro, a sodium ammonium triethanol ammonium laboratory cleaning solution. Afterward,
the plant material was placed in a new beaker and sonicated for an additional five minutes in
only distilled water. Sonicating the plant material removes any terrestrial diatoms, dust, and other
debris attached to the outer surfaces.
When possible, the plant tissues were then ground or cut into smaller pieces to create
more surface area which enabled easier digestion. While wet plant tissue is not always conducive
to being cut or ground, I did notice that they were easier to digest than non-sonicated, nonground, or -cut plant tissue.
32
Each plant tissue sample was placed in a glass beaker to which around 40 ml of chromic
acid was added. This acid broke down and dissolved the organic matter, leaving behind only
inorganic material, such as phytoliths. These beakers were stirred using a glass stir rod and were
heated on a Corning Glass Works PC-35 hot plate for several minutes or until the acid started to
foam, suggesting a chemical reaction was taking place. The beakers were then set aside for at
least 24 hours, with a petri dish atop to protect the samples from contamination in the shared lab.
All stages of the digestion involving acid, except centrifuging, were conducted under a fume
hood.
After at least 24 hours, the beakers were stirred once more and the acid-phytolith mixture
was poured into 15 ml centrifuge tubes that were then placed in a swinging head bench-model
centrifuge. Tubes were centrifuged for three to five minutes at 2400-2800 rpm. The supernatant
was then removed with a disposable pipette. Distilled water was then added to the vials and the
phytoliths and remaining organic precipitate matter thatcoalesced at the bottom of the vials were
agitated with a new disposable pipette. Once resuspended in the water, the vials were centrifuged
once more. This process of resuspension and centrifugation was often repeated four or five times
until the samples were clean. On a few occasions with very clouded samples, I found that I had
to resuspend and centrifuge samples seven or eight times. Clean samples were then stored in
vials of 70% ethanol.
I encountered two problems in the process. First, some plant material was hardier than
others. In several cases, after plant material had been digesting in acid for over 24 hours, organic
matter still remained. This suggested that not all possible phytoliths for that sample had been
released. I found three solutions to this problem and performed them as I saw fit. First, I used a
ceramic mortar and pestle to grind the plant material into smaller pieces as a preliminary break
33
down of organic matter. Second, I dried the material in a drying oven to remove excess water
that would interfere with digestion. Third, I extracted the undigested material and placed it in a
beaker with fresh chromic acid for an additional digestion. Some plants, such as the flower tops
of Achillea millefolium, took two digestions, and others such as Abies concolor, Crataegus
douglasii, and Shepherdia canadensis took three digestions.
The second problem was the presence of diatoms. In some samples, the presence of
diatoms was negligible. However, in several samples the presence of diatoms made it difficult to
locate and identify the phytoliths. To counter this, I added sonication to my methods. Two
diatom genera were frequently present: Fragilariaceae Fragilaria (sp.), and Aulacoseiraceae
Aulacoseira (sp.) (Diatoms of the United States 2016) (Figure 3.1).
Figure 3.1. Common cross contaminates found in phytolith slides: A. diatoms, B. spores
The residual material following digestion was mounted using permount diluted with
xylene, glass slides, and glass cover slips. As many phytoliths and residual material as possible
were placed on a slide, dried, and then a few drops of a permount mixture were added, and the
cover slip placed atop. I used two mounting mediums: one was of toluene and beta-pinene
polymer and the second was of toluene and butyl benzyl phthalate. For both mediums, a few
drops of xylene were added. Glass weights were used to ensure that the cover slip adhered flat.
34
Slides were left to set for at least 24 hours. If any permount mixture escaped from under the
cover slip and dirtied the slide, xylene and a scalpel were used to remove excess permount
mixture.
Slides were reviewed at 10x, 20x, and 40x for phytoliths and other residual material, such
as pollen, vascular tissue, and diatoms, and were imaged at magnification using a Zeiss Axiovert
135 light microscope and Nikon Optiphot-2 with an attached Infinity 2 camera. Images were
often examined by Dr. Ball, and phytoliths were reimaged if needed.
When searching for probable diagnostic phytoliths, I sought after common shapes. I also
researched all the plants I digested to see if other botanists and archaeobotanists had also found
phytoliths for the same plant (Appendix D). In several occasions, I used these reports and
compared them to my images.
To test the effectiveness of the typology, six pieces of groundstone from Wolf Village
were sent to Chad Yost of Paleoscapes in Arizona (Table 5.1). These pieces were wrapped in
aluminum foil when they were extracted to protect the ground surfaces from contamination.
Yost’s methods for washing the ground stone artifacts and then analyzing the washed
material is as follows. Loose dirt was brushed off the utilized surface of the groundstone, which
was then washed in distilled water. Once as much dirt as possible was off the stone, a surfactant
such as tepol was added to the clean, utilized surface. A sonicating toothbrush was then used to
further clean the surface. Any material removed in this final cleaning was collected and set aside
for analysis. The residue was centrifuged into a pellet, all water was removed, and the dry pellet
stored in alcohol. To remove microfossils from residue, the sample was suspended in various
heavy liquids for several hours at a time. After Yost analyzed the prepared slides for phytoliths,
35
he shipped the slides to me. This way I could analyze groundstone samples and compare my
results to his. I did not review his results before I conducted my analysis.
Groundstone Slide Analysis
Slide analysis was conducted using the Nikon Optiphot 2 at 10x, 20x, and 40x
magnifications. I created an analysis sheet based on the forms I observed in my typology, and I
used Pearsall’s typology examples in Paleoethnobotany (2015), as well as traditional commonly
identified phytoliths, such as the Panicoid cross (Appendix E).
I made several passes across the slides, which were prepared by Chad Yost, until I had
counted between 200 and 300 phytoliths. In some instances, I reviewed multiple slides from the
same sample. The counting procedures resemble pollen counting. When counting pollen,
botanists suggest higher counts around 1,000 grains for complex and diverse samples, and counts
of 150-200 grains for samples of twenty taxa or less. A 200-grain count is estimated to provide
“about 75-85 percent accuracy for common taxa”, which is considered sufficient (Pearsall
2015:222). This logic and counting process has been utilized by other phytolith analysts (Piperno
2006:115). After reaching the 200-300 phytolith count, I would make additional scans to identify
forms from rare types and add those to my counts, following methods common among with other
phytolith analysts (Piperno 2006:115).
36
4. Results of Typology
Of the 52 plants that I processed, 23 produced discernable phytoliths, 22 primarily
produced vascular tissue, seven produced no phytoliths, and 19 produced calcium oxalate
crystals. After discussing calcium oxalate crystals, I present the results of my phytolith typology
according to plant life form: forbs, trees and shrubs, and grasses (Granite Seed 2017; Utah State
University 2017). I had no sedges or rushes in my samples.
My results chapter is patterned after the results of Morris (2008), McCune (2014), and
McNamee (2013). While these researchers only described phytoliths that were common or
abundant, I have chosen to describe all forms observed because some forms they found in
abundance I found to be uncommon, and some forms I found in abundance they did not record.
This may be because they used the ashed material method which involves removing organic
material with extreme heat, whereas I removed organic material with acid.
All phytoliths, including tracheids and the like, were photographed and described using
the International Code for Phytolith Nomenclature (ICPN) (Madella et al., 2005) and ICPN 2.0
(Terry Ball, personal communication January 2017) (Table 4.1). I also reviewed phytoliths on
the PhytCore DB Images DB (2017). Slides were analyzed using 100x, 200x, and 400x
magnifications. The abundance or production index (PI) of each phytolith form was assessed
using a variation of the coding methods used by Wallis (2003) and McCune (2014):
1) Non-producer (NP): no phytoliths observed
2) Rare (R): one or two phytoliths observed for the entire slide
3) Uncommon (U): phytoliths sparse, about 3-30 per slide, with most fields empty
4) Common (C): a few phytoliths observed in most of the fields, about 30-100 per slide
5) Abundant (A): several phytoliths were observed in all the fields, >100 per entire slide
37
C. Sclerids (Figure 4.3)
C1. Astrosclerid
Table 4.1. Typology Code.
E. Epidermal, Articulate (Figure 4.4,
4.5, 4.6)
E1. Sinuate epidermal, psilate texture
E1a. striate texture
E1b. heavy or light striations
E2. Polygonal epidermal, psilate texture
E2a. granulate texture
E3. Ligulate epidermal, psilate texture
E3a. ligulate to collumnate
E4. Entire epidermal, psilate texture.
E4a. striate texture
E5. Favose epidermal
E6. Crenate epidermal
E7. Blocky epidermal, lateral striations.
Grass-type
H. Hairs (Figure 4.3)
H1. Lancelote hair
H1a.striate texture, unsegmen.
H1b. psilate texture, unsegmen.
H1c. psilate texture, segmented
H1d.granulate texture, unseg.
H2. Acicular hair
H2a. striate texture, unsegmen.
H2b. psilate texture, unseg.
H2c. ovoid base with
tuberculate processes.
Elymus type
H2d. needle/rod like.
O. Calcium Oxalates (Figure 4.1, 4.2)
O1. Raphide, single: needle shaped w/
one end pointed
O1a. bundle, same orientation
O1b. bundle, differ. orientation
O1c. small raphide-types
connected together and of differ
orientations
O1d. “dunmbbell” bundle
O2. Styloid, single
O2a. cluster
O3. Crystal sand
O4. Druse: several facets radiating from
a central core
O4a. druse-like
O5. Prismatic, rectangular, single
O5a. rectangular cluster
O5b. hexagon
O5c. hexagon cluster
O6. Rhombohedrals
G. Grass Short Cell Forms (Figure
4.12)
G1. Pooideae types
G1a. round/oblong
G1b. square/rectangular
G1c. keeled
G1d. pyramidal
G1d1. aculeated
G1e. trapeziform, sinuate
G1f. reniform shape
G2. Chloridoid types
G2a. saddle
G3. Stipa types
G3a. bilobate
G4. Panicoideae types
G4a. cross
G4b. bilobate
G5. Zea mays types
G5a. wavy top
G5b. ruffle top
G5c. IRP, rectangular
G6. Trichomes
G6a. lancelote style hairs
G6b. hair bases
G6c. bulliform
G7. Rondel, general.
G7a. circular/ovoid
G7b. polylobes/bilobes
V. Vascular Tissue
V1. Tracheids (Figure 4.7)
V2. Vascular tissue indeterminate
(Figure 4.8)
V3. Stomata (Figure 4.6)
V4. Trichome (Figure 4.6)
V4a. umbrella peltate
Y. Parenchyma (Figure 4.6)
Y1. Parenchyma
38
P. Papillae (Figure 4.8)
P1. Papillae with ligulate margins
P1a. tuberculated
P1b. pitted edges.
Grass-type.
S. Spheroid, Polygonal (Figure
4.9)
S1. Irregular sub-spheroid forms
S1a. ruminate texture
S1b. ruminate to facetate
S1c. granulate texture
S2. Blocky, psilate to facetate
texture
S2a. psilate to granulate
S2b. facetate texture
S3. Spheroids
S3a. granulate texture
S3b. ruminate texture
S4. “crescents”, or half nuclei
S5. Ellipsoids with tuberculate
processes. Elymus type.
L. Elongates (Figure 4.10, 4.11)
L1. Elongate with pilate margins.
Grass-type.
L1a. Achillea type
L1b. pilate to clavate
margins. Grass-type.
L2. Elongate with entire margins.
Grass-type.
L2a. granulate texture.
L2b. psilate texture,
needle like. Grass-type.
L3. Elongate, dendritic margins.
Grass-type
L4. Elongate, crenate margins.
Grass-type
L5. Elongate, aculeate margins.
Grass-type
L5a. curled. Grass-type
L5b. non-grass type.
L5c. granulate texture
Grass-type.
L6. Elongate, sinuate margins.
Grass-type
Calcium oxalates
Calcium oxalate crystals (CaOx) occur as monohydrated whewellites (CaC2O4-H2O) or
as dihydrated weddellites (CaC2O4-(2+X)H2O) (Franceschi and Horner 1980). CaOx crystals are
often formed in the idioblasts of plants of many taxonomic levels, from algae to giant
gymnosperms (Franceschi and Nakata 2005). CaOx crystals, akin to phytoliths, serve as
protection against herbivory and as calcium regulators (Franceschi and Nakata 2005). In some
cases, the crystals are a substitute for sclerenchyma (Schneider 1901). Crystals that are formed in
a cell wall are often either rhombohedral or prismatic, whereas crystals that form within a cell
can take any form (Franceschi and Nakata 2005). No crystals have been reported in pollen,
although the crystals can be found mixed with pollen (Anitha and Sandhiya 2014; Cote 2009).
The size of these crystals vary greatly due to cell type, function, and environmental factors such
as calcium availability (Franceschi and Nakata 2005).
Several have reported that a plant species will only form just one crystal type or a subset
of crystal forms (Franceschi and Nakata 2005; Monje and Baran 2002). CaOx crystals are
abundantly produced in angiosperms and gymnosperms, but not all plants in these groups
produce the crystals (Franceschi and Nakata 2005). Like silica phytoliths, then, the distribution
and production of CaOx crystals is species dependent (Franceschi and Nakata 2005). There does
seem to be greater production in leaves than in stems of most plants (Anitha and Sandhiya 2014).
The development of CaOx crystals appears to be correlated with the development and
maturity of a plant (Chairiyah et al. 2016; Cote 2009; Franceschi and Nakata 2005). The most
common forms are druse, raphide, prismatic, styloid, and sand.
Variations in crystals related to physiological maturity can include three druse states:
solid, semi-solid, and loosely druse (Chairiyah et al. 2016). Raphides can be observed as single
39
needles or as bundles of needles, and can vary in length from 15 μm to 260 μm (Chairiyah et al.
2016). Single raphide crystals are rarely attached to each other and are transparent in color,
whereas bundles can vary from black to reddish-brown in color (Chairiyah et al. 2016). Bundles
of raphides can be of similar or different orientations (He et al. 2012). Prismatic crystals can be
rectangular, square, or hexagonal (Chairiyah et al. 2016). Styloids can be observed in groups of
transparent, irregularly shaped, small rectangles in bundles or in star-like clusters (Chairiyah et
al. 2016). Crystal sand often appear as masses of small angular crystals (Franceschi and Nakata
2005). Other forms include raphide-style bundles and platy aggregations (He et al. 2016).
CaOx crystals have been observed in species of Prunus, Cactus, Opuntia, Crataegus,
Solanum, and in Typha latifolia (Anitha and Sandhiya 2014; Borrellii et al. 2010; Monje and
Baran 2002; Schneider 1901). Plants that are known producers of silica phytoliths may not be
producers of CaOx crystals and vice-versa.
I observed CaOx crystals in nineteen plants (Table 4.2). Eight produced raphides, seven
produced styloids, ten produced crystal sand, two produced druses, eight produced prismatic
shapes, and one produced rhombohedrals (Figure 4.1 and 4.2). Diagnostic types were observed
in Prunus virginiana and Opuntia polycantha, and probably Pinus edulis and Solanum jamesii.
Only four were forbs, one was grass-like, and fourteen were shrubs and/or trees. Six species had
crystals formed in their berries, nine in their leaves, one in its twigs, one in its flower head, one
in its nuts, and the last were in the pads, bud, and spines of Opuntia. Of the nineteen species that
produced CaOx crystals, two, Fragaria vesca and Sambucus cerulea, were not silica phytolith
producers. No CaOx crystals were observed in the grasses I tested, and they have rarely been
observed in other grass species (Prychid and Rudall 1999). Arctostaphylos patula may produce
rectangular pristmatic forms. However, more observations are needed.
40
Species
Amelanchier alnifolia
Shrub/tree
Amelanchier utahensis
Shrub/tree
Artemisia dracunculus
Forb
Artemisia ludoviciana
Forb
Artemisia tridentata
Shrub
41
Cercocarpus ledifolius
Shrub/tree
Ephedra viridis
Shrub
Fragaria vesca*
Forb
Holodiscus dumosa
Shrub
Table 4.2. Presence and Frequency of Calcium Oxalate Crystals.
Plant Tissue
Berry
Berry
Berry
Berry
Berry
Leaves
Leaves
Leaves
Phytolith
O2a
O3
O1c
O2
O5
O1c
O3
O3
PI
U
U
U
U
U
U
R
U/R
Figure
4.1.R
4.2.A
4.1.G
4.1.L
4.2.K
4.1.H
none
none
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Berry
Berry
Berry
Leaves
Leaves
Leaves
Leaves
Berry
O1a
O1b
O1c
O3
O5
O5b
O1b
O1c
O5
O1
O1c
O2
O2a
O5
O2
O2a
O5
O5b
O3
U
U
U
U
U
U
U
U
U
U
U
U
U
U
U
U
U
U
U/R
4.1.C
4.1.E
4.1.I
4.2.B
4.2.L
4.2.R
none
4.1.F
4.2.M
4.1.A
4.1.J
4.1.M
4.1.S
4.2.N
4.1.N
4.1.T
4.2.O
4.2.S
none
Juniperus osteosperma
Tree
*Denotes plant tissue that was not sonicated.
Species
Opuntia polycantha
Shrub
Pinus edulis
Tree
Pinus flexilis
Tree
Prunus virginia
Shrub/tree
Rosa woodsii
Shrub
Sambucus cerulea
Tree
Solanum jamesii
Forb
Typha latifolia
Plant Tissue
Pads
Pads
Buds
Spines and hairs
Twigs
Phytolith
O4
O4a
O4a
O4a
O5a
PI
C
A
C
C
C
Figure
4.2.H
4.2.H
4.2.H
4.2.H
4.2.Q
Nuts
Nuts
Nuts
Nuts
Nuts
leaves
Leaves
O2
O2a
O4a?
O3
O5
O1
O6
O5b
C
C
U/R
R
R
R
C
U
4.1.O
4.1.U
4.2.I
4.2.C
4.2.P
none
4.2.V
4.2.T
Berry
O3
U
4.2.D
Leaves
Leaves
Leaves
Leaves
Leaves
Pollen head
Pollen head
Pollen head
Pollen head
Pollen head
Pollen head
O2
O2a
O1d
O3
O4
O1
O1a
O2
O2a
O3
O5b
C
C
C
U
R
U
U
U
U
U
U
4.1.P
4.1.V
4.1.K
4.2.F
4.2.J
4.1.B
4.1.D
4.1.Q
4.1.W
4.2.G
none
Figure 4.1. Calcium oxalates: raphides and styloids.
A-B: Single raphides (O1). A. Ephedra viridis, leafy green. B. Typha latifolia pollen head. C-D: Raphide
bundle of same orientations (O1a). C. Artemisia tridentata leaves. D. Typha latifolia pollen head. E-F:
Raphide bundles of different orientations (O1b). E. Artemisia tridentata leaves. F. Cercocarpus ledifolius
leaves. G-J: Small raphide-types connected together and of different orientations (O1c). G. Amelanchier
utahensis berry. H. Artemisia dracunculus leaves. I. Artemisia tridentata leaves. J. Ephedra viridis leaves.
K: Raphide “dumbbell” bundle (Old). K: Solanum jamesii leaves. L-Q: Single styloid (O2). L.
Amelanchier utahensis berries. M. Fragaria vesca berries. N. Holodiscus dumosa leaves. O. Pinus
flexulis nuts. P. Solanum jamesii leaves. Q. Typha latifolia pollen head. R-W: Styloid cluster (O2a). R.
Amelanchier alnifolia berry. S. Fragaria vesca berry. T. Holodiscus dumosa leaves. U. Pinus flexulis
nuts. V. Solanum jamesii leaves. W. Typha latifolia pollen head.
42
Figure 4.2. Calcium oxalates: crystal sand, druses, prismatics, and rhombohedrals.
A-G: Crystal sand (O3). A. Amelanchier alnifolia berry. B. Artemisia tridentata leaves. C. Pinus flexulis
nuts. D. Sambucus cerulea berry. E. Sherpherdia argentea berry. F. Solanum jamesii leaves. G. Typha
latifolia pollen head. H-J: Druse and druse-like (O4 and O4a). H. Opuntia polycantha pads. I. Pinus
flexulis nuts. J. Solanum jamesii leaves. K-P: Rectangular, single prismatic (O5). K. Amelanchier
utahensis berry. L. Artemisia tridentata leaves. M. Cercocarpus ledifolius leaves. N. Fragaria vesca
berry. O. Holodiscus dumosa leaves. P. Pinus flexulis nuts. Q: Rectangular, prismatic cluster (O5a). Q.
Pinus edulis twigs. R-T: Hexagon prismatic (O5b). R. Artemisia tridentata leaves. S. Holodiscus dumosa
leaves. T. Rosa woodsii leaves. U: Rhombohedrals (O6). V. Prunus virginiana leaves.
43
Forbs
Fifteen forbs were analyzed for this typology (Table 4.3). One, Nicotiana attenuata, will
not be included because I was unable to collect or grow an acceptable sample to digest and had
little success in trying to digest a few of the seeds. Two others, Fragaria vesca and Typha
latifolia, were non-producers of silica phytoliths. The phytolith forms I observed most frequently
were epidermals, spheroids, and hairs. No forms appeared to be diagnostic.
Asteraceae
Seven Asteraceae species were forbs, all seven produced tracheid phytoliths. Five were
common producers of at least one phytolith form. Two were primarily uncommon producers of
phytoliths.
Achillea millefolium was a common producer of phytoliths. The leaves produced four
phytoliths: sinuate striate epidermal (E1a), lanceolate unsegmented psilate hairs (H1b), irregular
ruminate sub-spheroid phytoliths (S1a), and tracheids (V1). All four phytoliths were commonly
found. The hair and epidermal phytoliths were often found articulated. It appears that the
articulate epidermal converge to a point, the ultimate terminus of which is the hair. The
inflorescence produced one common form: long thin pilate elongate phytoliths (L1a); and three
phytoliths that were uncommon: papillae phytoliths with ligulate margins and occasional
tuberculate processes (P1a), S1a, and V1.
The Artemisia species I sampled all produced epidermal phytoliths and tracheid
phytoliths, but there was variation in the epidermals between species (Figure 4.4 and 4.5). In the
seeds and associated leaves, A. biennis commonly produced Ela, and rarely produced astrosclerid
phytoliths (C1) (Figure 4.3), ligulate epidermal phytoliths with psilate texture (E3), ruminate
spheroid phytoliths (S3b), and V1. The leaves of A. dracunculus commonly produced sinuate
44
epidermal that were either heavily or lightly striated (E1b) and V1. The leaves uncommonly
produced E3, crenate epidermal phytoliths (E6), acicular unsegmented striated hair phytoliths
(H2a), S1a, psilate to facetate texture blocky phytoliths (S2) that appear to be disarticulated
psilate polygonal phytoliths, and stomata (V3) in the articulate epidermals. A. ludoviciana leaves
commonly produced psilate sinuate epidermal phytoliths (E1), blocky phytoliths that were either
granulate or psilate (S2a), andV1. The leaves also uncommonly produced parenchyma (Y1). The
inflorescence tops commonly produced V1. This forb also produced some CaOx crystals.
Balsamorhiza sagittata was a common producer of one phytolith form and two
uncommon forms in the leaves and an uncommon producer of two forms in the inflorescence.
Lancelote psilate segmented hairs (H1c) were commonly produced in the leaves, and irregular
sub-spheroid phytoliths with facetate to ruminate texture (S1b), facetate blocky phytolithss
(S2b), and Y1 were uncommonly produced. Irregular sub-spheroid phytoliths with granulate
texture (Slc) and V1were uncommon in the inflorescence.
Solidago canadensis was an uncommon producer of S1c, Hlc, V1 and indeterminate
vascular tissue (V2).
Vigueiera multiflora uncommonly produced entire epidermal phytoliths with psilate
texture (E4), H1c, and V1.
Fabaceae
Entire epidermal phytoliths with striated texture (E4a) were rarely observed in the roots
of Hedysarum boreale.
45
Malvaceae
Sphaeralcea munroana leaves uncommonly produced polygonal epidermal phytoliths
with granulate texture (E2a), entire epidermal phytoliths with striate texture (E4a), V1, V3, and
Y1.
Polygonaceae
S1c were commonly observed in the leaves of Eriogonum ovalifolium. V2 was
uncommonly observed in the roots of E. ovalifolium. S1a was commonly observed in the leaves
of Eriogonum umbellatum. Uncommonly observed in the leaves were fragments of E1, favose
epidermal (E5), and V1.
Rosaceae:
For Fragaria vesca, see the section on calcium oxalate crystals.
Solanaceae
V1 and V2 were commonly observed in the leaves of Solanum jamesii. Spheroids with
granulate texture (S3a) and S1b were uncommonly observed in the leaves. V1 was rarely
observed in the tubers. See the section on calcium oxalate crystals for additional forms.
Typhaceae
Typha latifolia was found to be a non-producer of silica phytoliths. It did produce CaOx
crystals.
46
Species
Achillea millefolium
Artemisia biennis*
Artemisia dracunculus
47
Artemisia ludoviciana
Basalmhoriza sagittata
Table 4.3. Presence and Frequency of Phytoliths in Tested Forbs.
Plant Tissue
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Inflorescence
Inflorescence
Leafy seeds
Leafy seeds
Leafy seeds
Leafy seeds
Leafy seeds
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
New growth
Leaves
Leaves
Leaves
Leaves
Inflorescence
Leafy tops
Leafy tops
Leafy tops
Leafy tops
Inflorescence
Inflorescence
Phytolith
E1a
H1b
S1a
V1
L1a
P1a
S1a
V1
E1a
C1
E3
S3b
V1
E1b
V1
E3
E6
H2a
S1a
S2
V3
E1b
E1
S2a
V1
Y1
V1
H1c
S1b
S2b
Y1
S1c
V1
*Denotes plant tissue that was not sonicated.
PI
C
C
C
C
C
U
U
U
C
U
U
U
U
C
C
U
U
U
U
U
U
U
C
C
C
U
C
C
U
U
U
U
U
Figures
4.4.F
4.3.C
4.9.A
4.7.A
4.10.F
4.8.J
4.9.A
4.7.A
4.4.G
4.3.A
none
4.9.bb
none
4.4.J
4.7.C
none
4.5.M
4.3.I
4.9.B
4.9.Q
4.6.C
4.4.J
4.4.A
4.9.S
4.7.D
4.6.E
4.7.D
4.3.E
none
none
4.6.G
4.9.G
4.7.F
Species
Eriogonum ovalifolium*
Eriogonum umbellatum*
Fragaria vesca*
Hedysarum boreale*
Solanum jamesii
Solidago canadensis
Sphaeralcea munroano
Typha latifolia
Viguiera multiflora
Plant Tissue
Roots
Leaves
Leaves
Leaves
Leaves
Leaves
Berries
Roots
Tubers
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Inflorescence
Inflorescence
Leaves
Leaves
Leaves
Leaves
Leaves
Entire stalk
Inflorescence
Inflorescence
Inflorescence
Phytolith
V2
S1c
S1a
E1
E5
V1
E4a
V1
S1b
S3a
V1
V2
H1c
S1c
V1
V2
E2a
E4a
V1
V3
Y1
E4
H1c
V1
PI
U
U
C
U
U
U
NP
R
R
U
U
U
U
U
U
U
U
U
U
U
U
U
NP
U
U
U
Figures
4.8.C
4.9.I
4.9.C
4.4.C
4.5.L
4.7.L
4.5.I
4.7.U
none
4.9.aa
4.7.U
4.8.G
4.3.F
4.9.P
4.7.V
none
4.4.M
4.5.K
none
4.6.D
4.6.O
4.5.H
4.3.G
4.7.W
Figure 4.3. Phytoliths: astrosclerids and hairs.
A-B: Astrosclerid (C1). A. Artemisia biennis seeds with leaves. B. Gutierezia sarothrae tops with leaves.
C: Lancelote hair with psilate texture, unsegmented (H1b). C. Achillea millefolium leaves. D-G:
Lancelote hair with psilate texture, segmented (H1c). D. Amelanchier alnifolia berries. E. Basalmhoriza
sagittata leafy tops. F. Solidago candensis inflorescence. G. Viguiera multiflora inflorescence. H:
Lancelote hair with granulate texture, unsegmented (H1d). H. Poa fendleriana florets. I: Acicular hair
with striate texture, unsegmented (H2a). I: Artemisia dracunculus leaves. J-K: Acicular hair with psilate
texture, unsegmented (H2b). J. Elymus cinereus florets. K. Holodiscus dumosa inflorescence and leaves.
L-M: Acicular hair with an ovoid base with tuberculate processes (H2c). L. Elymus cinereus florets. M.
Elymus glaucus florets. N: Acicular hair that is needle-like (H2d). N. Festuca ovina florets.
48
Figure 4.4. Phytoliths: articulate epidermals, part one.
A-E: Sinuate epidermal with psilate texture (E1). A. Artemisia ludovicianna leaves. B. Artemisia
tridentata leaves. C. Erigonum umbellatum leaves. D. Rosa woodsii leaves. E. Sporobolus airoides
florets. F-I: Sinuate epidermal with striate texture (E1a). F. Achillea millefolium leaves. G. Artemisia
biennis seeds with leaves. H. Gutierezia sarothrae tops with leaves. I. Holodiscus dumosa leaves. J:
Sinuate epidermal with heavy or light striations (E1b). J. Artemisia dracunculus leaves and new growth
plant tops. K-L: Polygonal epidermal with psilate texture (E2). K. Rosa woodsii leaves. L. Shepherdia
argentea leaves. M: Polygonal epidermal with granulate texture (E2a). M. Sphaeralcea munroano leaves.
N: Ligulate epidermal with psilate texture (E3). N. Prunus virginiana leaves. O-P: Ligulate epidermal
with ligulate to collumnate margins (E3b). O. Elymus cinereus florets. P. Elymus glaucus florets.
49
Figure 4.5. Phytoliths: articulate epidermals, part two.
A-H: Entire epidermal with psilate texture (E4). A. Deschampia cespitosa florets. B. Elymus cinereus
florets. C. Elymus glaucus florets. D. Holodiscus dumosa leaves. E. Juniperus communis young shoots. F.
Shepherdia argentea berries. G. Stipa hymenoides florets. H. Viguiera multiflora inflorescence. I-K:
Entire epidermal with striate texture (E4a). I. Hedysarum boreale roots. J. Rhus aromatica leaves. K.
Sphaeralcea munroano leaves. L: Favose epidermal (E5). L. Erigonum umbellatum leaves. M-O: Crenate
epidermal (E6). M. Artemisia dracunculus leaves. N. Holodiscus dumosa leaves. O. Sporobolus airoides
florets
50
Figure 4.6. Phytoliths: articulate epidermals part three, stomates, parenchyma, and trichomes.
A-B: Blocky epidermal, lateral striations (E7). A. Stipa hymenoides florets. B. Sporobolus airoides
florets. C-D: Stomata (V3). C. Artemisia dracunculus leaves and inflorescence. D. Sphaeralcea
munroano leaves. E-O: Parenchyma (Y1). E. Artemisia ludoviciana leaves. F. Artemisia tridentata
leaves. G. Basalmhoriza sagittata leafy tops. H. Deschampia cespitosa florets. I. Festuca ovina florets. J.
Gutierezia sarothrae tops with leaves. K. Holodiscus dumosa leaves. L. Prunus virginiana leaves. M.
Rhus aromatic berries and leaves. N. Rosa woodsii leaves. O. Sphaeralcea munroano leaves. P: Umbrella
peltate trichome (V4a). P. Shepherdia argentea leaves and berries.
51
Trees and Shrubs
Thirty-three of the plants sampled for this study were trees and shrubs (Table 4.4). Four
species produced no phytoliths, twenty-two produced at least one form that was uncommon or
rarely produced, and fifteen commonly produced at least one phytolith form, often tracheids. The
phytoliths that were produced most frequently observed were tracheids and spheroids. A lack in
silificiation in woody species has been noted by others (Morris 2008).
Adoxaceae
No phytoliths of any kind were observed in the berries of Sambucus cerulea or in the
berries of Sambucus racemosa. See the section on calcium oxalate crystals for the berries of
Sambucus cerulea.
Amaranthaceae
V1 were commonly observed in the inflorescence of Atriplex truncata. Uncommonly
produced S2 amd S2b were also observed.
Anacardiaceae
E4a, S1c, V1, and Y1 were uncommonly observed in the leaves of Rhus aromatica. V2
and Y1 were uncommonly observed in the berries of R. aromatica.
Asteraceae
While other Artemisia species are forbs, Artemisia tridentata is a shrub. Unlike other
Artemisia species, I found that A. tridentata uncommonly produced epidermal phytoliths in the
52
leaves. These took the form of E1, and a few were in the process of breaking apart. These broken
epidermal phytoliths are likely the source of the S2b phytoliths that were uncommonly observed
in the leaves. S1b were uncommonly produced in the leaves and were commonly produced in the
inflorescence. V1 were uncommon in the inflorescence, and V2 was common in the wood. Y1
was rarely observed in the leaves.
In the inflorescence and leaves of Chrysothamnus nauseosus, Slc, S2a, S2b, and V1 were
commonly observed.
Ela, entire elongate phytoliths with granulate texture (L2a), and Slc, V1, Y1 were
uncommonly observed in the leafy tops of Gutierrezia sarothrae. There is also a probable C1
phytolith that was rarely observed.
Cactaceae
V1 was uncommon in the pads of Opuntia polycantha. V1 was common and V2 was
uncommon in the bud, and S1b was uncommon in the spines and hairs. For additional forms, see
the section on calcium oxalate crystals.
Cupressaceae
I observed no phytoliths in Juniperus scopulorum. E4 and V1 were rare in the young
shoots of J. communis. S3a and V1 were uncommonly observed in, respectively, the berries and
leaves of J. osteosperma.
53
Elaeagnaceae
E4 were rarely observed in the berries of Shepherdia argentea. Umbrella shaped peltate
trichomes (V4a) were commonly observed in the berries. For additional forms, see the section
on calcium oxalate crystals. Polygonal epidermal phytoliths with psilate texture (E2), V1, and
V4a, were uncommon in the leaves of Shepherdia argentea. In the berries of Shepherdia
canadensis, S2b were uncommon. In the leaves, S1c and V1 were uncommon.
Ericaceae
I found no phytoliths in Arctostaphylos patula.
Ephedraceae
V1 were commonly observed in the leafy greens of Ephedra nevadensis. S2b were
uncommonly observed in the leafy greens. V1 were commonly observed in the leafy greens of
Ephedra viridis¸ and S1a and S2b were uncommonly observed in the woody twigs. For
additional forms, see the section on calcium oxalate crystals.
Pinaceae
The needles of Abies concolor are non-producers of phytoliths. Other Abies species have
been known to produce phytoliths (Appendix D).
No phytoliths were found in the needles or sap of Pinus edulis. Thick crescent phytoliths,
or half spherical granulated outlines (S4) were uncommonly observed in the shells of the nuts of
P. edulis. This phytolith may be diagnostic of pinyon pine nuts. For additional forms, see the
section on calcium oxalate crystals. V2 was uncommonly observed in the twigs.
54
Pinus flexulis produced V1 uncommonly in the nuts. See the section on calcium oxalate
crystals for more forms.
I found no phytolithss in the sap of Pinus monophylla. I did observe that elongates with
aculeate margins (L5b), and V1 were uncommon in the needles.
Rosaceae
Of the nine Rosaceae shrub and tree species, only two were common producers of
phytoliths: Prunus virginiana and Rosa woodsii.
I observed uncommon H1c in the berries of Amelanchier alnifolia. For additional forms,
see the section on calcium oxalate crystals. In the berries of Amelanchier utahensis, V1 are
uncommon. In the wood of A. utahensis, V1 was uncommon, and V2 that compares favorably to
E2 were common. For additional forms, see the section on calcium oxalate crystals.
Cercocarpus ledifolius produced uncommon V2 that compares favorably to E2 in the
wood. For additional forms, see the section on calcium oxalate crystals. S2 and V2 were
uncommonly observed in the berries of Crataegus douglasii. V2 was uncommon and H2b was
commonly observed in the flowers of Holodiscus dumosus. E1a, E3, E4, E6, H2b, V1 and Y1
were uncommonly observed in the leaves. For additional forms, see the section on calcium
oxalate crystals.
Prunus virginiana commonly produced E3, V1, V2, and Y1 in the leaves and V2 were
commonly observed in the berries of P. virginiana. S1c and V2 were commonly observed in the
roots. See calcium oxalate crystals for rhombohedrals.
V2 was uncommon and S1c rare in the leaves of Purshia mexicana. No phytoliths were
observed in the inflorescence. S1c and S2b were uncommonly observed in the leaves of Purshia
55
tridentata. S3b were uncommonly observed in the berries of Rosa woodsii. Y1 were
uncommonly observed in the leaves, and E1, E2, and V1 were commonly produced. CaOx
crystals were also observed.
Sarcobataceae
S2b and V1 were uncommonly observed in the leaves of Sarcobatus vermiculatus.
Saxifragaceae
S1a were uncommonly produced in the berries of Ribes aureum.
56
Table 4.4. Presence and Frequency of Phytoliths in Tested Shrubs and Trees, Part I.
Species
Abies concolor
Tree
Amelanchier alnifolia*
Shrub/tree
Amelanchier utahensis
Shrub/tree
Arctostaphylos patula
Shrub
Artemisia tridentata*
Shrub
57
Atriplex truncata*
Shrub
Cercocarpus ledifolius
Shrub/tree
Chrysothamnus nauseous
Shrub
Craetaegus douglasii*
Tree
Ephedra nevadensis
Shrub
Plant Tissue
Needles
Phytolith
PI
NP
Figures
Berries
H1c
U
4.3.D
Berries
Wood
Wood
Leaves
V1
V1
V2
U
U
C
NP
4.7.B
4.7.B
4.8.A
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Twigs
Inflorescence
Inflorescence
Inflorescence
Leaves
Wood
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Inflorescence
Inflorescence
Berry
Berry
Green stems
Green stems
E1
S1b
S2b
Y1
S1b
V1
V2
S2
S2b
V1
U
U
U
R
C
U
C
U
U
C
NP
U
C
C
C
C
C
C
C
C
U
U
U
C
4.4.B
4.9.E
none
4.6.F
4.9.E
4.7.E
none
none
4.9.U
none
V2
S1c
S2a
S2b
V1
S1c
S2a
S2b
V1
S2
V2
S2b
V1
*Denotes plant tissue that was not sonicated.
4.8.B
4.9.H
4.9.T
4.9.V
4.7.G
4.9.H
4.9.T
4.9.V
4.7.G
4.9.R
none
4.9.W
4.7.J
Species
Ephedra nevadensis
Ephedra viridis
Shrub
Gutierezia sarothrae*
Shrub
Holodiscus dumosa
Shrub
Juniperus communis
Shrub/tree
Juniperus osteosperma
Tree
Juniperus scoporulum
Shrub/tree
Opuntia polycantha
Shrub
Pinus edulis*
Tree
Plant Tissue
Wooden twigs
Green stems
Wooden twigs
Wooden twigs
Leafy tops
Leafy tops
Leafy tops
Leafy tops
Leafy tops
Leafy tops
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Twigs
Young growth
Berries
Leaves
Twigs
Bud
Bud
Pad
Spines/hairs
Nuts
Sap
Needles
Twigs
Phytolith
V1
S1a
S2b
E1a
L2a
S1c
V1
Y1
C1
E1a
E4
E6
H2b
V1
Y1
H2b
V2
E4
V1
V1
S3a
V1
V2
V1
S1b
S4
V2
PI
NP
C
U
U
U
U
U
U
U
R
U
U
U
U
U
U
C
U
NP
R
R
U
U
NP
Figures
C
U
U
U
U
NP
NP
U
4.7.O
none
4.7.O
4.9.F
4.9.cc
4.7.K
none
none
4.4.H
4.10.N
4.9.J
4.7.M
4.6.J
4.3.B
4.4.I
4.5.D
4.5.N
4.3.K
4.7.N
4.6.K
4.3.K
4.8.D
4.5.E
none
none
4.9.Z
4.8.E
Species
Pinus flexulis
Tree
Pinus monophylla
Tree
Table 4.4 Presence and Frequency of Phytoliths in Tested Shrubs and Trees, Part II.
Prunus virginiana
Shrub/tree
Purshia mexicana
Shrub/tree
58
Purshia tridenta
Shrub
Rhus aromatic*
Shrub/tree
Ribes aureum
Shrub
Rosa woodsii
Shrub
Sambucus caerulea*
Shrub
Plant Tissue
Seeds
Resin
Needles
Needles
Berries
Leaves
Leaves
Leaves
Leaves
Roots
roots
Inflorescence
Leaves
Leaves
Leaves
Leaves
Berries
Berries
Leaves
Leaves
Leaves
Leaves
Berries
Berries
Leaves
Leaves
Leaves
Leaves
Berries
Phytolith
V1
L5b
V1
V2
E3
V1
V2
Y1
S1c
V2
S1c
V2
S1c
S2b
V2
Y1
E4a
S1c
V1
Y1
S1a
S3b
E1
E2
V1
Y1
PI
U
NP
U
U
C
C
C
C
C
C
C
NP
R
U
U
U
U
U
U
U
U
U
U
U
C
C
C
U
NP
*Denotes plant tissue that was not sonicated.
Figures
none
4.11.N
none
none
4.4.N
4.7.P
none
4.6.L
4.9.K
none
4.9.L
none
4.9.M
4.9.X
4.8.F
4.6.M
4.5.J
4.9.N
4.7.Q
4.6.M
4.9.D
none
4.4.D
4.4.K
4.7.R
4.6.N
Species
Sambucus racemosa
Shrub
Sarcobatus vermiculatus
shrub
Shepherdia argentea
Tree
Shepherdia canadensis
Shrub
Plant Tissue
Berries
Phytolith
PI
NP
Figures
Leaves
Leaves
Berries
Berries
Leaves
Leaves
Leaves
Berries
Leaves
Leaves
S2b
V1
E4
V4a
E2
V1
V4a
S2b
S1c
V1
U
U
R
C
U
U
U
U
U
U
none
4.7.S
4.5.F
4.6.P
4.4.L
4.7.T
4.6.P
4.9.Y
4.9.O
none
Figure 4.7. Phytoliths: tracheids.
A-W: Tracheids (V1). A. Achillea millefolium leaves and inflorescence. B. Amelanchier utahensus
berries and wood. C. Artemisia dracunculus leaves. D. Artemisia ludoviciana leaves and inflorescence. E.
Artemisia tridentata inflorescence. F. Basalmhoriza sagittata inflorescence. G. Chrysothamnus nauseous
leaves and inflorescence. H. Deschampia cespitosa florets. I. Elymus cinereus florets. J. Ephedra
nevadensis green stems. K. Ephedra viridis green stems. L. Eriogonum umbellatum leaves. M. Gutierezia
sarothrae tops with leaves. N. Holodiscus dumosa leaves. O. Opuntia polycantha bud and pad. P. Prunus
virginiana leaves. Q. Rhus aromatic leaves. R. Rosa woodsii leaves. S. Sarcobatus vermiculatus leaves.
T. Shepherdia argentea leaves. U. Solanum jamesii tubers and leaves. V. Solidago canadensis
inflorescence. W. Viguiera multiflora inflorescence.
59
Figure 4.8. Phytoliths: vascular tissue and papillae.
A-G: Vascular tissue, unknown (V2): A. Amelanchier utahensis wood. B. Cercocarpus ledifolius wood.
C. Eriogonum ovalifolium roots. D. Holodiscus dumosa inflorescence. E. Pinus edulis twigs. F. Rhus
aromatic berries. G. Solanum jamesii leaves. H-I: Papillae with ligulate margins (P1). H. Elymus
cinereus. I. Poa fendleriana. J: Papillae with ligulate margins, tuberculated (P1a). J. Achillea millefolium
inflorescence. K-L: Papillae with pitted edges. Grass-type. (P1b). K. Elymus cinereus florets. L. Elymus
glaucus florets.
60
Figure 4.9. Phytoliths: spheroids and polygonals.
A-D: Irregular sub-spheroid forms with ruminate texture (S1a). A. Achillea millefolium leaves and
inflorescence. B. Artemisia dracunculus leaves. C. Eriogonum umbellatum leaves. D. Ribes aureum
berries. E-F: Irregular sub-spheroid forms with ruminate to facetate texture (S1b). E. Artemisia tridentata
leaves and inflorescence. F. Opuntia polycantha spines and hairs. G-P: Irregular sub-spheroid forms with
granulate texture (S1c). G. Basalmhoriza sagittata inflorescence. H. Chrysothamnus nauseous leaves and
inflorescence. I. Eriogonum ovalifolium leaves. J. Gutierezia sarothrae tops with leaves. K. Prunus
virginiana roots. L. Purshia Mexicana leaves. M. Purshia tridenta leaves. N. Rhus aromatic leaves and
berries. O. Shepherdia canadensis leaves. P. Solidago canadensis inflorescence. Q-R: Blocky with psilate
to facetate texture (S2). Q. Artemisia dracunculus leaves. R. Craetaegus douglasii berries. S-T: Blocky
with psilate to granulate textures (S2a). S. Artemisia ludoviciana leaves. T.Chrysothamnus nauseous
inflorescence and leaves. U-Y: Blocky with facetate texture (S2b). U. Atriplex truncata inflorescence
with seeds. V. Chrysothamnus nauseous leaves and inflorescence. W. Ephedra nevadensis green stems.
X. Purshia tridenta leaves. Y. Shepherdia canadensis berries. Z-aa: Spheroids with granulate texture
(S3a). Z. Juniperus osteosperma leaves. aa. Solanum jamesii leaves. bb: Spheroids with ruminate texture
(S3b). bb. Artemisia biennis seeds with leaves. cc: “crescents”, or half nuclei (S4). cc. Pinus edulis nuts.
dd-ee: Ellipsoids with tuberculate processes (S5). dd. Elymus cinerus florets. ee. Elymus glaucus florets.
61
Grasses
Seven grasses were analyzed for the typology. The seven grasses were uncommon,
common and abundant phytolith producers in the florets (Table 4.5).
Grasses produce short-cell and long-cell phytoliths (Figure 4.9-4.12). The three major
types of grass short-cell phytoliths include pooidoid, chloridoid, and panicoid. The short cell
phytoliths include saddles, trapeziforms, and rondels. In descriptions of these short cells, the base
of short cells is the largest and flattest surface, and the top is the side opposite (Mulholland
1989). Long-cells include elongates, epidermals, and tracheary elements. Other forms often
observed are trichomes, such as papillae, hair cells, and buliforms (Piperno 2006:34-35).
Pooideae, such as Festuca, Poa, Bromus, Elymus, to name a few, produce rondels, and
pyramids (Brown 1984; Kearns 2001:286). These cool-season grasses produce rondel shapes
such as circular, rectangular, elliptical, crescent, and oblong (Twiss et al. 1969). They also
sometimes produce bilobates and polylobates.
The warm season grasses found in arid to semi-arid areas in the subfamily Chlorideae
typically produce saddles (Kearns 2001:286). There are two saddle types, one being the thinner
version of the other (Twiss et al. 1969). Of all the grasses in the southwest, Chloridoid grasses
are most abundant (Gould and Shaw 1983:120). However, some in this subfamily also produce
bilobate and pooideae phytoliths (Twiss 1987:181).
Panicoids are warm-season and/or tall grasses that grow in many habitats. Panicoideae,
such as Panicum and Schizayrium produce panicoid lobates (Kearns 2001:286), as well as
bilobates and polylobates. Panicoid phytolith shapes include: bilobate, cross, body oblong with
oblong platform, small bilobate, Zea mays rondel, and Zea mays saddle.
62
Long cell phytoliths are found in all grasses and while diagnostic of the grass family,
these phytoliths are not often diagnostic of a subfamily, tribe, and so forth (Twiss et al. 1969).
Diagnostic phytoliths do exist, in particular the elongate with dendritic margins. Additionally,
current studies on the elongate with dendritic margins found in the Triticeae tribe suggest that it
may be possible to differentiate between the different genera of this tribe based on morphometry
(Ball et al. 1999).
Pooideae
Five of the grass species I tested are in the Pooideae subfamily. Compared to the two
other tested subfamilies, there were more long cell types observed in the the Stipea and
Chloridoid subfamilies than in the Pooideae subfamily.
In Deschampia cespitosa I observed 17 phytoliths: eight long cells, and nine short cells.
The long cells were E1, E4, elongates with pilate margins (L1), elongates with entire margins
(L2), elongates with aculeate margins (L5), elongates with aculeate margins that are curled
(L5a), V1, and blocky epidermal with lateral striations (E7). The short cells were round to
oblong rondels (G1a), square to rectangular rondels (G1b), keeled rondels (G1c), and pyramidal
rondels (Gld), some of which had aculeate processes (G1d1), and polylobes (G7b). I also
observed were Y1, and trichomes, including lancelote-style hairs and hair bases (G6a, G6b).
None of the forms were diagnostic.
There were 17 phytoliths in Elymus cinereus: seven long cell types, and ten short cell
types. Long cell phytoliths include E4, ligulate to collumnate epidermal (E3a), dendritic
elongates (L3), elongate with crenate margins (L4), L5, aculeate elongates with granulate texture
(L5c), and V1. Short cell phytoliths include G1b, G1c, G1d, P1, some with pitted edges (P1b),
63
G6a, G6b, an acicular hair with an ovoid base with tuberculate processes (H2c), and ellipsoids
with tuberculate processes (S5) that were likely hair bases for the acicular hairs with psilate
texture (H2b). L3 and H2c are diagnostic.
In Elymus glaucus, there were 15 phyoliths: six long cell types, and nine short cell typess.
The long cells were E3a, E4, E7, L3, L4, and L5. The short cells were G1c, G1d, G1d1, reniform
shaped rondels (G1f), G6a, H2c, P1, P1b, and S5. A few of the H2c phytoliths had granulate
texture. L3 and H2c are diagnostic.
There were 16 phytoliths in Festuca ovina: eight long cell types and eight short cell
types. Long cell types were L1, elongates with clavate to pilate processes (L1b), L2, elongates
with entire margins and granulate texture (L2a), L5, L5a, L5c, and V1. Short cells were G1a,
G1c, G1d, G1d1, G6a, G6b, needle-like acicular hairs (H2d), and Y1.
I observed 15 phytoliths in Poa fendleriana: six long cell types, and nine short cell types.
The long cells were L1, L2, L4, articulate L4, L5, L5c, articulate L5 with P1, and V1. The short
cells were G1a, G1b, G1c, trapeziforms (Gle), G7b, P1, P1b, G6a, and H1d.
In an intensive study of nine Poa species, 31 morphotypes were identified and biogenic
silica content was found to average 16.8% of dry weight per species. No correlation, though, was
found between the number of phytoliths identified and the biogenic silica content (Lisztes-Szabo
et al. 2015:371). Elongate long cells and short cells were common in each of the species, but
there was variation between the species with respect to texture and ornamentation, frequency,
and number of types. Psilate and sinuate textures were observed in all species. Rondeltrapeziforms and lancelote trichomes were also observed in all species (Lisztes-Szabo et al.
2015:371). Papillae were only observed in one of the species, and the trigonal pyramid type was
also rarely observed (Lisztes-Szabo et al. 2015:371). Statistical analysis revealed that the various
64
Poa species clustered together in different groups, and that there are interspecific variations
between the different species (Lisztes-Szabo et al. 2015:377). Yet there were three types that
characterize this genus: elongate psilate, rondel trapeziform, and lancelote or scuitform types
(Lisztes-Szabo et al. 2015:377).
In the Poa species I tested, I did observe the psilate elongates and what I believe were the
lancelote types. Trapeziform rondels were uncommon.
Stipea
While in the Pooideae sub-family, the only species in this tribe that I analyzed was Stipa
hymenoides. Species in the Stipea tribe produce a diagnostic phytolith: the Stipa-type bilobe
(G3a). I observed this short cell type, as well as seven other short cells, and eleven long cells,
totaling 19 phyoliths total. Short cells were G3a, G1a, G1e, H1d with a prickly-base, H2d, G6a,
G6b, and probable bulliform phytoliths (G6c). Long cells were E4, E7, L1, L2, L2a, elongates
with entire margins that were needle-like (L2b), L4, L5, L5c, elongates with sinuate margins that
were articulate (L6).
Chloridoid
I only tested one species in the Chloridoid sub-family: Sporobolus airoides. I observed 13
phytoliths: ten long cell types, and three short cell types. Short cells include chloridoid type
saddles (G2a), polylobes (G7b), and circular to ovoid shaped rondels (G7a). Long cells were L1,
L2, L2a, L2b, L5, L5c, and articulate L5, E1, E6, E7, and V1.
65
Table 4.5. Presence and Frequency of Phytoliths in Tested Grasses, Part I.
66
Species
Subfamily/Tribe
Cell Type
Phytolith
PI
Figure
Species
Subfamily/Tribe
Deschampia
cespitosa*
Pooideae
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Short
E1
E4
E7
L1
L2
L5
L5a
V1
G1a
G1b
G1c
G1d
G1d1
G6a
G6b
G7b
Y1
E3a
E4
L3
L4
L5
L5c
V1
G1b
G1c
G1d
G6a
G6b
H2b
H2c
P1
P1b
S5
U
C
U
C
C
C
C
U
C
C
C
C
C
U
U
U
U
U
U
A
U
C
C
U
C
C
C
U
U
U
U
C
C
U
none
4.5.A
none
4.10.A
4.10.H
4.11.E
4.11.L
4.7.H
4.12.A
4.12.E
4.12.H
4.12.M
4.12.P
4.12.Y
none
4.12.jj
4.6.H
4.4.O
4.5.B
4.10.S
4.11.A
4.11.F
4.11.O
4.7.I
4.12.F
4.12.I
4.12.Q
4.12.Z
none
4.3.J
4.3.L
4.8.H
4.8.K
4.9.dd
Elymus
glaucus
Pooideae
Festuca
ovina*
Pooideae
Elymus
cinereus
Pooideae
*Denotes plant tissue that was not sonicated.
Cell
type
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
PI
Figure
E3a
E4
E7
L3
L4
L5
G1c
G1d
G1d1
G1f
G6a
H2c
P1
P1b
S5
L1
L1b
L2
L2a
L5
L5a
L5c
V1
G1a
G1c
G1d
G1d1
G6a
G6b
H2d
Y1
U
U
U
A
C
C
C
C
C
C
C
C
C
C
C
C
U
C
C
A
A
C
U
C
U
U
U
C
U
U
U
4.4.P
4.5.C
none
4.10.T
4.11.B
4.11.G
4.12.J
4.12.N
4.12.R
4.12.V
4.12.aa
4.3.M
none
4.8.L
4.9.ee
4.10.B
4.10.G
4.10.I
4.10.M
4.11.H
4.11.M
4.11.P
none
4.12.B
4.12.K
4.12.O
4.12.S
4.12.bb
none
4.3.N
4.6.I
Table 4.5 Presence and Frequency of Phytoliths in Tested Grasses, Part II.
Species
Poa
fendleriana*
Subfamily/Tribe
Pooideae
Sporobolus
airoides*
Chloridoid
67
Cell Type
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Long
Long
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Phytolith
L1
L2
L4
L5
L5c
V1
G1a
G1b
G1c
G1e
G6a
G7b
H1d
P1
P1b
E1
E6
E7
L1
L2
L2a
L2b
L5
L5c
V1
G2a
G7a
G7b
*Denotes plant tissue that was not sonicated.
PI
C
C
C
C
C
U
C
C
C
C
C
U
C
U
U
C
C
C
A
A
A
A
A
A
U
C
C
C
Figure
4.10.C
4.10.J
4.11.C
4.11.I
4.11.Q
none
4.11.C
4.11.G
4.11.L
4.11.T
4.11.cc
4.11.jj
4.3.H
4.8.I
none
4.4.E
4.5.O
4.6.B
4.10.D
4.10.K
4.10.O
4.10.Q
4.11.J
4.11.R
none
4.12.W
4.12.ff
4.12ii
Species
Stipa
hymenoides*
Subfamily/Tribe
Stipea
Cell type
Long
Long
Long
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
E4
E7
L1
L2
L2a
L2b
L4
L5
L5c
L6
V1
G1a
G1e
G3a
G6a
G6b
G6c
H1d
H2d
PI
U
U
C
C
C
U
C
C
C
C
U
A
A
A
C
C
C
C
C
Figure
4.5.G
4.6.A
4.10.E
4.10.L
4.10.P
4.10.R
4.11.D
4.11.K
4.11.S
4.11.T
none
4.12.D
4.12.U
4.12.X
4.12.dd
none
4.12.ee
none
none
Figure 4.10. Phytoliths: elongates, part one.
A-E: Elongate with pilate margins. Grass-type. (L1). A. Deschampia cespitosa florets. B. Festuca ovina
florets. C. Poa fendleriana florets. D. Sporobolus airoides florets. E. Stipa hymenoides florets. F:
Elongate with pilate margins. Achillea type (L1a). F. Achillea millefolium inflorescence. G: Elongate with
pilate to clavate margins. Grass-type. (L1b). G. Festuca ovina florets. H-L: Elongate with entire margins.
Grass-type. (L2). H. Deschampia cespitosa florets. I. Festuca ovina florets. J. Poa fendleriana florets. K.
Sporobolus airoides florets. L. Stipa hymenoides florets. M-P: Elongate with entire margins, granulate
texture. (L2a). M. Festuca ovina florets. N. Gutierezia sarothrae tops with leaves O. Sporobolus airoides
florets. P. Stipa hymenoides floret. Q-R: Elongate with entire margins, psilate texture, rod-needle like.
Grass-type (L2b). Q. Sporobolus airoides florets. R. Stipa hymenoides florets. S-T: Elongate, dendritic
margins. Grass-type (L3). S. Elymus cinereus florets. T. Elymus glaucus florets.
68
Figure 4.11. Phytoliths: elongates, part two.
A-D: Elongate, crenate margins. Grass-type (L4). A. Elymus cinereus florets. B. Elymus glaucus florets.
C. Poa fendleriana florets. D. Stipa hymenoides florets. E-K: Elongate, aculeate margins. Grass-type
(L5). E. Deschampia cespitosa, florets. F. Elymus cinereus florets. G. Elymus glaucus florets. H. Festuca
ovina florets. I. Poa fendleriana florets. J. Sporobolus airoides florets. K. Stipa hymenoides florets. L-M:
Elongate, aculeate margin, curled. Grass-type (L5a). L. Deschampia cespitosa florets. M. Festuca ovina
florets. N: Elongate, aculeate margin, non-grass type (L5b). N. Pinus monophylla needles. O-S: Elongate,
aculeate margin, granulate texture. Grass-type (L5c). O. Elymus cinereus florets. P. Festuca ovina florets.
Q. Poa fendleriana florets. R. Sporobolus airoides florets. S. Stipa hymenoides florets. T: Elongate,
sinuate margins. Grass-type (L6). T. Stipa hymenoides florets.
69
Figure 4.12. Phytoliths: grass short cell forms.
A-D: Pooideae type- round/oblong (G1a). A. Deschampia cespitosa florets. B. Festuca ovina florets. C.
Poa fendleriana florets. D. Stipa hymenoides florets. E-G: Square/rectangular (G1b). E. Deschampia
cespitosa florets. F. Elymus cinereus florets. G. Poa fendleriana florets. H-L: keeled (G1c). H.
Deschampia cespitosa florets. I. Elymus cinereus florets. J. Elymus glaucus florets. K. Festuca ovina
florets. L. Poa fendleriana florets. M-O: Pyramidal (G1d). M. Deschampia cespitosa florets. N. Elymus
glaucus florets. O. Festuca ovina florets. P-S: Aculeated (G1d1). P. Deschampia cespitosa florets. Q.
Elymus cinereus florets. R. Elymus glaucus florets. S. Festuca ovina florets. T-U: trapeziform, sinuate
(compares favorably) (G1e). T. Poa fendleriana florets. U. Stipa hymenoides florets. V. Reniform shape
(G1f). V. Elymus glaucus florets. W: Chloridoid types- saddle (G2a). W. Sporobolus airoides florets. X:
Stipa types-bilobate (G3a). X. Stipa hymenoides florets. Y-dd: lancelote style hairs (G6a). Y.
Deschampia cespitosa florets. Z. Elymus cinereus florets. aa. Elymus glaucus florets. bb. Festuca ovina
florets. cc. Poa fendleriana florets. dd. Stipa hymenoides florets. ee: bulliform cf (G6c). ee. Stipa
hymenoides florets. ff: Rondel, general- circular/ovoid (G7a). ff. Sporobolus airoides florets. jj-ll:
polylobes/bilobes (G7b). jj. Deschampia cespitosa florets. kk. Poa fendleriana florets. ll. Sporobolus
airoides florets.
70
Discussion and Conclusion
The species that were the most common and abundant producers of phytoliths were
grasses and forbs. Shrubs and trees were the most frequent non-producers or rare and uncommon
producers of phytoliths. Common and abundant phytoliths were most observed in the leaves of
these species. Of the tested species and plant material, root and wood samples produced close to
no phytoliths. Nine produced probable diagnostic types, and ten forms are non-indicators and
non-diagnostic (Table 4.6). I believe it can be safely stated that the diagnostic types may be
specific to the genus-level, but more testing is needed.
Table 4.6. Diagnostic and Non-diagnostic Phytoliths.
PhytolithForm
Elongate with psilate margins
Umbrella peltate trichome
Crescent, half-nuclei
Elongate with dendritic margins
Druse
Rhombohedrals
Prismatic clusters
Raphide “dumbbell” bundle
Unsegmented acicular hair, psilate texture, knobby base
Saddle
Stipa bilobate
Pooideae rondel types
Panicoideae rondel types
Zea mays rondel types
Astrosclerid
Hairs
Articulate epidermals
Styloids, raphides, sand, prismatic
Spheroids
Trichomes
Stomates, Papillae
Parenchyma
Elongates
Tracheids
Diagnostic level
Achillea genus
Shepherdia genus
Pinus genus
Triticea tribe
Opuntia genus
Prunus genus
Pinus genus
Solanum genus
Elymus genus
Chloridoid tribe
Stipa
Pooidoids
Panicoids
Zea mays
Uncertain
none
none
Non-grasses, tree and shrub
None
Grasses
None
None
Grasses
Non-grasses
Plant Part
Inflorescence
Berries, leaves
Nut shells
florets
Pad, bud
Leaves
Twigs
leaves
Florets
Florets
Florets
Florets
Florets
Cob
uncertain
Florets
Florets
-
Hairs were observed in five of the forbs, four of which are from the Asteraceae family.
Hairs were only observed in three of the tested shrubs and trees, with their occurrence only being
71
common in the inflorescence of Holodiscus dumosa. The other two species were uncommon
producers. Only one observed hair morphotype could be diagnostic, and that was the
unsegmented acicular hair with psilate texture and a knobby base produced by Elymus glaucus.
Articulated epidermals were observed in all Artemisia species tested regardless of plant
form, although frequency and style varied. These phytoliths were also observed in five other
forbs and seven trees and shrubs, and in four grasses. None of the species tested produced
diagnostic articulated epidermals.
Tracheids were observed in ten forbs and twenty trees and shrubs. Tracheids are perhaps
the least diagnostic of all the phytoliths observed since so many plants produce them, including
grasses. However, McNamee (2013:120) observed that when tracheids are found in the soil
record that they are an indicator of non-grasses.
Indicators of grass species are elongates, most of which were produced only by grasses
save for the Achillea millefolium type. While elongates are primarily produced by grasses,
individual grasses cannot yet be identified by this phytolith form except for grasses from the
Triticeae tribe. Elymus cinereus and Elymus glaucus both produced elongates with dendritic
margins. This form is diagnostic of the Triticeae tribe.
A probable indicator of trees and shrubs may be calcium oxalate crystals. Of the
seventeen plants that produced these crystals, thirteen were trees and shrubs. Morris (2008:127160) also noted that trees and shrubs more often produced calcium oxalates than their forb and
grass counterparts. Of the phytoliths I observed, there were two species that produced diagnostic
forms and two with a probable diagnostic form. These were the druses from Opuntia polycantha,
the rhombohedrals from Prunus virginia, and the prismatic clusters of Pinus edulis and dumbbell
72
raphide bundle from Solanum jamesii are the probable diagnostic forms. No CaOx crystals were
observed in the grasses.
Clear non-diagnostic forms include papillae, stomates, indeterminate vascular tissue, and
spheroids. In contrast, the umbrella peltate trichomes may be diagnostic of Shepherdia species,
and the crescent, or half-nuclei spheroid may be diagnostic of Pinus edulis.
Regarding grass types, other than the elongates with dendritic margins and Zea mays
rondel-types, no other grass phytoliths are diagnostic to a genus or species level. Some
phytolithss, such as the Stipa-type bilobate, are indicative of subfamilies, but nothing more.
73
5. Results of Ground Stone Artifact Wash Analysis
Six ground stone artifacts (Table 5.1) from Wolf Village (42UT273) were selected for
analysis and for the testing of the typology I created for the Utah Valley area. In this chapter, I
discuss the ground stone artifacts and the phytoliths identified thereon. For Field Specimens (FS)
219, 2357, 15814, and 16494, I counted the same slides as Yost; however, I also scanned all
additional slides. For FS 11975 and 16642, I counted and scanned both slides, whereas Yost only
counted and scanned the first slide.
I employed Yost to identify phytoliths on the ground stones to test my accuracy in
counting and the viability of the typology I created. If I found the same types at a similar
frequency as Yost, then that would suggest that my counting was accurate and the typology
functional. What I found was that despite minor differences in specific phytolith counts, the
overall story of what plants were used on what ground stone artifacts was similar between Yost
and myself. What this demonstrates is that the regional typology created for the Utah Valley
Fremont, as based on plants with documented medicinal and dietary ethnographic uses, is
reliable and valid, and that my counting was sufficient.
Tables 5.2-5.4 present phytolith counts and Table 5.5 calculated percentages, with Yost’s
counts and percentages being denoted with a Y. The terms used by both Yost and myself were
standardized for greater consistency in the tables. For example, all grass bilobates and crosses are
included in the Zea mays counts and not in the C4 grass counts. Zea mays and C4 grasses are
panicoids, all of which make crosses and bilobates (Terry Ball, personal communication, May
13, 2017). However, I found the transitional forms among panicoids between crosses and
bilobates difficult to differentiate into separate Zea mays and C4 categories. Therefore, I have
74
included all such forms under evidences for Zea mays (Figure 5.7). I also included the scanned
observations from Yost’s in his overall counts since I had included scanned observations in my
totals. Not included was the Commelina domed cylinder Yost observed in FS 219, the Cucurbita
scalloped hemispherical in FS 2357, and the Tradescantia pyramidal in FS 16494 (Appendix F).
Table 5.1. Wolf Village (42UT273) Ground Stone Tool Provenience and Description.
Sample
No.
219
2357
11975
Feature
Size
assoc.
Description
F48 in 15cm by 10
F5
cm, 7.5cm
thick, 1362.4g
F111 in 7 by 7.25,
F130 in 4.5cm thick,
F110
534.3g
F866 in 18 by 6.25,
F864
8 cm thick,
1715.7g
15814
F1119
in F923
17.5 by 9.5,
5 cm thick,
1059.2g
16494
F1158
in F522
25 by 9,
11.5 cm thick,
12360.4 g
16642
F882 in
F864
17 by 11, 7cm
thick,
2778.25g
Artifact Description
A fragment of a flat metate made of
basalt. One side is clearly
worked/shaped.
This appears to be a central fragment
of a vesicular basalt trough mano. The
edges of the surface are well curved.
A complete, quartzite basin mano. The
distal ends could have been used for
pounding. Although pecking was used
to shape the stone and to sharpen
grinding surfaces, only one side has
clear evidence of grinding.
Complete ovoid-shaped rhyolite mano.
There is no ground surface. The edges
have been shaped, and one surface is
flat and another is more convex.
Likely a mano preform.
This fragmented, proximal end of a
basalt metate has what appears to be a
shelf on the ventral side that is often
associated with Utah style metates.
The sides seem to have been shaped,
and the bottom near the break appears
to have been used for grinding. There
is faint evidence of a red stain. The
entire stone is smooth.
Quartzite slab used as a surface to
mash fruits and roots.
75
Wear
Form
Worn,
pecked,
ground
Worn,
pecked
Slab without
formed
trough
Loaf shaped
Worn,
pecked,
ground
Loaf shaped
Ground
One handed,
flat/tabular
loaf
Worn,
pecked,
ground
Trough with
secondary
shelf
Minor
ground
wear,
mashed
Slab
Ground Stone Artifact 219
FS 219 is a basalt metate fragment, with a discernible ground and pecked use surface
(Figure 5.1). It was located in the fill outside of Structure 3, a pit house with well-preserved
floor, side benches, and post holes (Johansson et al 2014:39).
Two slides were prepared from this sample; both Yost and I counted and scanned the first
slide, additionally I also scanned the second slide. Minor variations in some of the counts may be
attributed to my scanning of the second slide and Yost not. However, we both found similar
forms at a similar frequency. Compared to the other ground stone artifacts, FS 219 has a high
maize and sedge phytolith recovery.
This artifact’s phytolith assemblage is characterized by 55.1-57.8% grasses, which
include rondels from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and Phalaris
grasses, and elongates and ellipsoids indicative of Elymus. Phragmites was also observed.
Sedges accounted for 21.3-27.1% of the sample, and Zea mays types, such as double-walled and
wavy-top rondels, made up 15.1-21.3% of the sample. No other diagnostic phytoliths were
observed. Non-phytolith forms observed were calcium oxalate crystals, diatoms, and sponge
(Figure 5.8). Additionally, FS 219 was characterized by a low starch granule recovery, with
sedges, grasses, Zea mays, Elymus, Typha, and Calochortus/Fritillaria (Liliaceae) represented
(Appendix F:7).
Ground Stone Artifact 2357
FS 2357 is a worn and pecked central fragment of a vesicular basalt mano that was likely
used on a trough metate (Figure 5.2). It was found in a stratum of Structure 5, which was a subrectangular pit house.
76
Eight slides were prepared from this sample. Yost and I both counted and scanned slide
one. Yost then scanned slides two and three, whereas I scanned all the remains slides, two
through eight. I believe that variations in some of the counts can be attributed to this.
Additionally, these were the first slides that I chose to analyze using my typology. Variations
between counts, though, were never greater than the 13.1% difference observed in the total
C3/C4 grass count. Compared to the other ground stone artifacts, FS 2357 is characterized by
having a high grass phytolith recovery.
The phytolith assemblages observed are made up of 77.3-89.6% grasses, including
rondels from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and Phalaris grasses,
and elongates and ellipsoids indicative of Elymus. Phragmites was also observed. Sedges
accounted for 2.3-5.8% of the sample, and Zea mays types, such as double-walled and wavy-top
rondels, made up 7.8-9.7% of the sample. Other diagnostic types observed include epidermal
indicative of Prunus, Pinus crescents, and a microfossil that closely resembled Cucurbita forms
(Figure 5.10). Diatoms and sponge microfossils were also observed. FS 2357 had the second
highest starch granule recovery, with grasses, sedge, maize, Elymus-type, Typha, Apiaceae,
Calochortus/Fritillaria (Liliaceae) represented. Yost suggests that the straight-line brakes
observed on several of the grass elongates and epidermis are indicative of anthropomorphic
cutting and/or rolling pressure from being ground (Appendix F:8). The dark coloring of several
of the grass microfossils is likely indicative of exposure to fire, for example, for parching.
77
Figure 5.1. Ground Stone FS 219, basalt metate fragment.
Figure 5.2. Ground Stone FS 2357, a worn and pecked central fragment of a vesicular basalt
mano.
78
Table 5.2. Phytolith Grass Counts.
79
SAMPLE (FS #)
219 219-Y 2357 2357-Y 11975 11975-Y 15814 15814-Y 16494 16494-Y 16642 16642-Y
Trapeziform sinuate: Pooideae
82
91
173
140
125
92
177
145
152
84
163
100
Rondel-keeled: Pooideae
2
8
9
16
3
40
15
23
3
39
2
33
Rondel-square/rectangular: Pooideae
5
0
9
0
4
0
9
0
5
0
7
0
Rondel-angular keel: Phalaris
0
1
0
3
0
1
0
1
0
1
0
1
Rondel-plateau saddle: Phragmites
0
4
0
4
0
1
0
0
0
0
0
1
Bilobate-Stipa type (cf): Stipea
0
8
5
4
0
5
2
14
0
8
0
0
TOTAL C3 Short Cells
89
112
196
167
132
139
203
183
160
132
172
135
TOTAL C4 Chloridoid Saddle
2
6
16
2
0
17
1
3
0
4
0
9
Rondel-all forms: C3 & C4 Grasses
21
25
23
39
29
37
25
47
33
42
57
41
Grass epidermis, general
0
1
1
4
0
0
0
0
0
0
0
0
Epidermal-crenate/sinuate margins
3
0
1
0
0
0
0
0
1
0
0
0
Bulliform (cf)
8
4
12
3
5
4
0
0
4
3
5
0
Trichome, general
1
13
2
24
0
32
0
8
1
27
1
19
Trichome base (hair)
8
1
2
0
9
2
7
1
5
1
12
0
Elongate-entire margins,
11
9
16
41
17
38
4
29
7
43
18
32
Elongate-crenate margins
2
0
4
0
0
0
1
0
0
0
0
0
Elongate-aculeate margins
2
13
22
20
8
25
6
11
5
14
6
24
Elongate-charred
1
0
0
0
0
0
1
0
0
0
0
0
Elongates-articulate
8
0
20
0
0
0
0
0
5
0
2
0
TOTAL redundant Grass types
65
66
103
131
68
138
44
96
61
130
101
116
156
184
315
300
200
294
248
282
221
266
273
260
Total
(cf) = compares favorably.
Samples Yost identified are listed directly after the samples I identified. My counts are demarcated with the ground stone artifact (FS) number, and Yost’s
(Appendix F) are demarcated by a Y.
See Figure 5.9.
Table 5.3. Diagnostic Phytolith Counts.
SAMPLE (FS #)
219
219-Y
2357
2357-Y
11975
11975-Y
15814
15814-Y
16494
16494-Y
16642
16642-Y
Rondel-Wavy top: cf. Maize
21
14
13
5
9
4
7
5
21
21
10
12
Rondel-ruffle top: cf. Maize
3
0
1
1
4
0
3
1
1
1
0
0
Rondel-half decorated: cf. Maize
0
0
2
0
0
0
0
0
0
0
0
0
Rondel-charred/double wall (cf)
11
10
8
14
2
5
5
12
14
9
6
7
Rondel cluster/epidermis: Maize
0
0
3
2
0
0
1
4
2
1
0
0
Cross: Panicoideae/Maize (cf)
18
9
5
2
3
1
4
2
23
10
7
9
Rondel-cross/bilobe: cf. Maize
8
17
8
2
3
11
2
2
9
14
12
17
IRP-type: Maize glume
0
0
0
1
0
0
0
0
0
1
0
1
TOTAL Maize types observed
61
50
40
27
21
21
22
26
70
57
35
46
Elongate-dendritic
0
8
0
9
0
1
1
2
1
4
2
2
Ellipsoid Elymus type
2
0
4
0
2
0
1
0
4
0
6
0
80
TOTAL Elymus types
2
8
4
9
2
1
2
2
5
4
8
2
Thin w/ridges: Sedge stem
61
88
24
7
52
30
7
4
38
12
56
42
Sedge: root and cone cell
0
2
0
1
0
3
0
0
0
1
0
3
Epidermal-Prunus type
0
0
1
0
0
0
0
0
0
0
0
0
Crescent/Nuclei-Pinus type
0
0
5
0
0
0
0
0
3
0
2
0
124
148
74
44
75
55
31
32
116
74
101
93
Total diagnostic types
(cf) = compares favorably.
Samples Yost identified are listed directly after the samples I identified. My counts are demarcated with the ground stone artifact (FS) number, and Yost’s are
demarcated by a Y.
See Figure 5.7.
Ground Stone Artifact 11975
FS 11975 is a complete quartzite basin mano that has been pecked and ground (Figure
5.3). The mano was likely shaped before use, and was found in a midden associated with
Structure 9 and its associated vent tunnel.
Two slides were prepared from this sample. Yost counted and scanned slide one, and I
counted and scanned both slides one and two. I believe variations in our counts may be attributed
to this. The greatest variations were observed between redundant grass form totals and the totals
for all grasses combined. Compared to the other ground stones, FS 11975 is characterized by a
low Zea mays phytolith recovery.
This stone’s phytolith assemblage was made up of 70.1-83.1% grasses, including rondels
from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and Phalaris grasses, and
elongates and ellipsoids indicative of Elymus. Phragmites was also observed. Sedges accounted
for 9.3-18.1% of the sample, and Zea mays types, such as double-walled and wavy-top rondels,
made up 5.9-7.3% of the sample. No other diagnostic types were observed. Diatoms and sponge
microfossils were present. FS 11975 had a relatively low starch granule recovery, with grasses,
maize, Elymus, and Calochortus/Fritillaria (Liliaceae) represented.
Ground Stone Artifact 15814
FS 15814 is another complete mano (Figure 5.4). This one was made of rhyolite and
seems to be one-handed. It was found in the vent tunnel associated with Structure 9.
Only one slide was prepared for this sample, which was counted and scanned by both
Yost and myself. The phytolith counts for this stone have some of the least variation between
81
counts when compared to the other ground stones. Additionally, this ground stone is
characterized by a high grass phytolith and a low sedge and Zea mays phytolith recovery.
The phytolith assemblage observed for FS 15814 is made up of 85.9-88.7% grasses,
including rondels from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and
Phalaris grasses, and elongates and ellipsoids indicative of Elymus. Sedges accounted for 1.32.4% of the sample, and Zea mays types, such as double-walled and wavy-top rondels, made up
7.6-8.1% of the sample. No other diagnostic types were observed. Sponge microfossils, diatoms,
Chrysophyte cysts, and CaOx crystals were also present. FS 15814 had the third highest starch
grain count, with grasses, maize, Elymus, sedges, and Liliaceae represented.
Ground Stone Artifact 16494
FS 16494 is a fragment of what appears to be a Utah-style metate (Figure 5.5). This
artifact as been shaped, worn, pecked, and ground. It was found in the vent shaft fill associated
with Structure 8.
Two slides were prepared from this sample; Yost counted and scanned just the first slide,
I did the same but also scanned the second slide. I do not believe that the additional scanned slide
adequately accounts for the variations between counts. The greatest variation was observed
between the redundant grass forms. When compared to the other stones, FS 16494 is
characterized by a high Chrysophyte cyst and a high Zea mays phytolith recovery.
The phytolith assemblages observed are made up of 65.7-78.3% grasses, including
rondels from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and Phalaris grasses,
and elongates and ellipsoids indicative of Elymus. Sedges accounted for 3.8-11.0% of the
sample, and Zea mays types, such as double-walled and wavy-top rondels, made up 16.5-20.3%
82
of the sample. Other diagnostic types observed include the Pinus crescent. Diatoms, sponge
microfossils, CaOx crystals, and Chrysophyte cysts were also observed. FS 16494 had the
second lowest starch granule recovery, with most coming from grasses and sedge, with maize
and Elymus being diagnostic. Corn smut spores were recovered as well.
Yost speculates that the high presence of Chrysophyte cysts may be attributed to the
processing of fibrous roots from aquatic plants (Appendix F:9). Chrysophyte cysts are algae
much like diatoms, but are different in that they are more ecologically restricted. They are often
found in freshwater habitats, such as montane lakes, ephemeral ponds, and snowmelts, and can
survive some winter freezing (Appendix F:2). I did not note cysts on other groundstones because
I did not yet understand what to look for. I noted them for FS 16494 and FS 16642 because I had
learned what the cysts looked like.
Figure 5.3. Ground Stone FS 11975, a complete quartzite basin mano.
83
Figure 5.4. Ground Stone FS 15814, a complete rhyolite mano.
Figure 5.5. Ground Stone FS 16494, fragment of a basalt Utah-style metate.
84
Table 5.4. Redundant Forms, Total Phytolith Count, and Other Forms.
SAMPLE (FS #)
85
219
219-Y
2357
2357-Y
11975
11975-Y
15814
15814-Y
16494
16494-Y
16642
16642-Y
Epidermal-ligulate margins
0
0
1
0
0
0
0
0
1
0
3
0
Epidermal-sinuate margins
1
0
1
0
0
0
0
0
0
0
0
0
Epidermal-lattice
0
0
0
0
9
0
0
0
0
0
0
0
Epidermal-entire margins
1
0
0
0
0
0
0
0
0
0
1
0
Parenchyma
2
0
0
0
0
0
0
0
0
0
0
0
Spheroids, general-ruminate texture
0
0
0
1
1
6
4
6
3
5
7
2
Tracheids
0
0
2
0
0
0
0
0
2
0
4
0
Astrosclerid
0
0
0
0
0
0
0
0
1
0
1
0
Stomates
2
0
16
0
0
0
0
0
0
0
2
0
Hair, segmented & acicular
1
0
1
0
0
0
0
0
0
0
1
0
Hair, general
0
0
1
0
0
0
0
0
0
0
2
0
Hair bases, general
0
0
2
0
3
0
8
0
0
0
0
0
Total general types
7
0
24
1
13
6
12
6
7
5
21
2
TOTAL Phytoliths
287
332
413
345
288
355
291
320
344
345
395
355
Chrysophyte cysts
0
1
0
2
0
2
0
13
200
137
60
13
CaOx-raphide, single (cf)
3
-
0
-
0
-
0
-
0
-
2
-
CaOx-druse cf
0
-
0
-
0
-
0
-
0
-
6
-
1
-
1
-
2
-
4
-
2
-
0
-
Sponge
3
6
0
4
11
20
3
1
12
8
23
8
Diatoms (observed)
x
x
x
x
x
x
x
x
x
x
x
x
CaOx-rectangular/
hexagon prismatic (cf)
(cf) = compares favorably.
Samples Yost (Appendix F) identified are listed directly after the samples I identified. My counts are demarcated with the ground stone artifact
(FS) number, and Yost’s are demarcated by a Y.
See Figure 5.8.
Table 5.5. Calculated Percentages for Select Phytolith Counts.
SAMPLE (FS #)
219
219-Y
2357
2357-Y
11975
11975-Y
15814
15814-Y
16494
16494-Y
16642
16642-Y
Total C3 grasses
31.0
33.7
47.5
48.4
45.8
39.2
69.8
57.2
46.5
38.3
43.5
38.0
Chloridoid saddle
0.7
1.8
3.9
0.6
0.0
4.8
0.3
0.9
0.0
1.2
0.0
2.5
Redundant grass forms
22.6
19.9
24.9
38.0
23.6
38.9
15.1
30.0
17.7
37.7
25.6
32.7
Total C3/C4 grass types*
54.4
55.4
76.3
87.0
69.4
82.8
85.2
88.1
64.2
77.1
69.1
73.2
Total Zea mays types
21.3
15.1
9.7
7.8
7.3
5.9
7.6
8.1
20.3
16.5
8.9
13.0
Total Elymus
0.7
2.4
1.0
2.6
0.7
0.3
0.7
0.6
1.5
1.2
2.0
0.6
Total sedge
21.3
27.1
5.8
2.3
18.1
9.3
2.4
1.3
11.0
3.8
14.2
12.7
All diagnostic
43.2
44.6
17.9
12.8
26.0
15.5
10.7
10.0
33.7
21.4
25.6
26.2
All general
2.4
0.0
5.8
0.3
4.5
1.7
4.1
1.9
2.0
1.4
5.3
0.6
*does not include Elymus or Zea mays.
Samples Yost (Appendix F) identified are listed directly after the samples I identified. My counts are demarcated with the ground
stone artifact (FS) number, and Yost’s are demarcated by a Y.
86
Ground Stone Artifact 16642
The function of this artifact was not wholly apparent until after microfossil analysis.
Initial observations of the stone were that there was no discernable ground surface, but further
investigation coupled with what the microfossils revealed suggest that this stone was used as a
platform upon which tubers, fruits, and the like, were mashed. FS 16642 was found in a midden
associated with Structure 9 (Figure 5.6).
Two slides were prepared from this stone. Yost only counted and scanned the first, I
counted and scanned both slides. The variation between counts never exceeded 7.1%. Compared
to the other ground stone artifacts, FS 16642 is characterized by having a high Chrysophyte cyst
and high CaOx crystal recovery.
The phytolith assemblages observed are made up of 72.1-73.8% grasses, including
rondels from C3 and C4 grasses, particularly Pooideae, Stipea, Chloridoid, and Phalaris grasses,
and elongates and ellipsoids indicative of Elymus. Phragmites was also observed. Sedges
accounted for 12.7-14.2% of the sample, and Zea mays types, such as double-walled and wavytop rondels, made up 8.9-13.0% of the sample. Other diagnostic types observed include the Pinus
crescent. Diatoms and sponge microfossils were also observed. Maize starch counts were higher
from FS 16642 than from all the other artifacts combined. Additional starch grains came from
grasses, sedges, Elymus, Apiaceae, Triteleia, and Calochortus/Fritillaria (Liliaceae) were all
represented. A corn smut spore was also recovered.
The CaOx crystals observed seem to be damaged druses (Figure 5.8). This crystal form is
commonly observed in cacti and opuntia species. In the plants sampled for the typology, the only
species to abundantly and commonly produce druses was Opuntia polycantha. Given that the
87
interpretation of the function of this stone artifact is a surface that fruits and roots were mashed
upon, it may be likely that cacti may have also been mashed upon the stone.
Figure 5.6. Ground Stone FS 16642, a platform for mashing roots and tubers.
Summary
The Fremont of Wolf Village used their ground stone artifacts, in particular manos and
metates, to process C3 and C4 grasses such as Stipa hymenoides, Elymus glaucus, and several
88
other species. They also processed maize, sedges, and also probably pine nuts, chokecherry
fruits, and cacti. Based on the starches recovered, several roots and tubers, especially those
belonging to the Apiaceae and Liliaceae families, were also used. Several of these plants may
have been previously processed through parching or baking prior to being ground on the stones.
Evidence of this is derived from the presence of burnt grass and maize phytoliths.
89
Figure 5.7. Maize phytoliths observed on ground stone artifacts.
1. Wavy top rondel. 2. Ruffle top rondel. 3. Double-wall rondel. 4. Panicoid cross. 5. Panicoid bilobe.
6. Maize rondel cluster. 7. Half-decorated rondel. 8. Probable IRP.
a. FS 219. b. FS 2357. c. FS 11975. d. FS 15814. e. FS 16494. f. FS16642.
90
Figure 5.8. Calcium oxalate crystals, chrysophyte cysts, diatoms, sponges, and epidermals.
Calcium Oxalates: A. FS 219 calcium oxalate raphide. B. FS 219 calcium oxalate prism. C. FS 11975
calcium oxalate prismatic. D. FS 16494 calcium oxalate prism. E. FS 16642 calcium oxalate druse. Cysts:
F. FS 16494 chrysophyte cyst. Diatoms: G. FS 2357 diatoms. H. FS 16642 diatom. Sponge: I. FS 219
sponge microfossil. J. FS 16494 sponge microfossil. K. FS 16642 sponge microfossil. Epidermal
phytoliths: L. 11975, lattice epidermal. M. 219, epidermal with entire margins. N. 16642, epidermal with
entire margins. O. 2357, epidermal with sinuate margins. (219 not pictured). P. 16642, epidermal with
ligulate margins (2357 and 16494 not pictured). Hairs and hair bases: Q. 2357, hair base. R. 15814, hair
base (11975 not pictured). S. 219, segmented hair. T. 2357, segmented hair. U. 2357, hair. V. 16642 hair.
W. 16642, segmented hair.
91
Figure 5.9. Grass phytoliths.
A. FS 219, bulliform. B. FS 2357, bulliform. C. FS 11975, bulliform. D. FS 16642, bulliform. E. 219, trichome. F.
16494 trichome. G. 16642, trichome. H. 2357, articulate elongates with aculeate margins. I. 16642, elongates with
aculeate margins. J. 219, elongate with entire margins. K. 2357, elongate with entire margins. L. 16642, elongate
with entire margins. M. 2357, Elongate with crenate margins. N. 2357, sinuate Pooideae trapeziform. O. 15814,
sinuate Pooideae trapeziform. P. 16494, sinuate Pooideae trapeziform. Q. 16642, sinuate Pooideae trapeziform.
Rondels: R. 2357, keeled rondel. S. 15814, keeled rondel. T. 16642, keeled rondel. U. 2357, square rondel. V.
15814, square rondel. W. 16642, square rondel. X. 2357, stipea rondel. Y. 15814, stipea rondel. Z. 219, chloridoid
saddle. aa. 2357, chloridoid saddle. bb. 15814, chloridoid saddle. cc. 2357, round rondel. dd. 15814, round rondel.
ee. 16642 round rondel. Other forms: ff. 219, polylobe. gg. 2357, probable bulliform. hh. 2357, probable bulliform.
ii. 16642, rondel cluster. jj. 16642, polylobe.
92
Figure 5.10. Diagnostic forms, sedge, tracheids, and others.
A. 2357, Prunus epidermal. B. 2357, Pinus crescent. C. 16494, Pinus crescent. D. 16642, Pinus crescent.
E. 2357, Elymus ellipsoid. F. 15814, Elymus ellipsoid. G. 16642, Elymus ellipsoid. H. 16642, dendritic
elongates. I. 219, sedge. J. 2357, sedge. K. 11975, sedge. L. 15814, sedge. M. 16494, sedge. N. 16642,
sedge. O. 16494, astrosclerid. P. 16642, astrosclerid. Q. 15814, spheroids. R. 16642, spheroid. S. 2357,
stomates. T. 219, parenchyma. U. 2357, tracheids. V. 16494, tracheids. W. 16642, tracheids. X.
219, burned articulates. Y. 2357, unclassified epidermal. Z. 2357, unclassified hair.
93
6. Discussion and Conclusion
This thesis research has sought to fills gaps in phytolith knowledge on particular species
regarding how and where they produce phytoliths, and then apply a regionally specific typology
to archaeological samples. I conclude by discussing the results of the typology, what the
groundstone analysis adds to our understanding of Utah Valley Fremont plant use, and how
ethnographic resources can provide further interpretation.
Phytolith Typology and Ground Stone Analysis
I employed Yost to identify phytoliths on the ground stones to test the viability of the
typology I created and my accuracy in counting. Despite minor differences in specific phytolith
counts, the overall story of what plants were used on what ground stone artifacts was similar
between Yost and myself. This demonstrates that the regional typology created for the Utah
Valley Fremont based on plants with documented medicinal and dietary ethnographic uses is
reliable and valid, and that my counting was sufficient.
Regarding the typology, I believe that there may be benefits in creating regionallyspecific typologies. For example, if it can be concluded that in the Utah Valley region the only
species to produce tuberculated ovoid forms are those in the Elymus genus, then when such a
form is found it can be concluded that it is from Elymus.
Regarding counting, differences greater than five-percent between counts were most
commonly observed among the grass counts. This difference never exceeded 15.3 percent. I
believe the counts vary because I found it difficult to differentiate effectively between the Stipa-
94
bilobe, the panicoid cross, the maize cross, chloridoid saddles, and notched bilobates, as well as
between rondel forms such as pyramid, keeled, and forms indicative of C3 and C4 grasses.
Some types while uncommon to find are not indicators of any specific plant. These
phytolith forms include stomates, articulate epidermal, astrosclerids, hairs, spheroids, and
parenchyma. Several forms were more commonly observed, but were ultimately just indicators
of general types. For example, tracheids, which were observed on three of the stones, are
indicators of non-grass plants. Other examples include trichomes (hairs, hair bases, and
buliforms) and elongates as indicators of grasses, and calcium-oxalate crystals as indicators of
non-grasses, particularly trees and shrubs.
Indicators for Fremont Subsistence
The phytolith and starch analysis of six ground stone artifacts from Wolf Village has
added the following family to Utah Valley Fremont plant use: the Liliaceae family (specifically
Calochortus and Fritillaria). Liliaceae has previously not been found in archaeological contexts
in Utah Valley. If we accept the validity of regionally specific typologies, the following specific
plants can be added to current understanding of Utah Valley Fremont plant use: Poaceae Elymus
grasses, Rosaceae Prunus berries, Pinyon pine nuts and Cactaceae. Additional evidence
supporting the use of Cucurbita in Utah Valley was also found. Previously, there was only one
recording of Cucurbita from a rind at Spotten Cave, of one Prunus from a seed at Wolf Village,
and of one Elymus starch from Seamons Mounds teeth tartar. The presence of the Chrysophyte
cysts on ground stones FS 16494 and 16642 suggests the use of lakeside vegetation as well.
These plants are suggestive of diets that included foraging and farming, or at least of wild
and domestic plant use. When interpreting how these plants may have been used, I focus on
95
plants that would have likely been processed on a ground stone. I would not expect to find plants
that have no ethnographic documentation for being ground or crushed on ground stones. This is
likely why plants, such as Achillea millefolium, that have diagnostic phytoliths were not found in
the ground stone samples. Achillea millefolium was primarily prepared by boiling, not grinding.
Specifics regarding suggested interpretations for how the Fremont may have used these
plants, as well as what species would have been available are as follows. Apiaceae starch was
found on FS 2357 and FS 16642. Prior to this, Apiacea starch had been found in teeth tartar from
Seamon’s mound (Yost 2009:6), and as pollen on ground stone from Hinckley Mounds (Peterson
2016). I did not test any Apiaceae species for my phytolith typology. There are 24 Apiaceae
species in the valley (Welsh et al. 1987:613-637), and ten have documented ethnographic uses.
Of these ten, seven have preparation processes that involve grinding: Angelica pinnata, Carum
gairdneri, Cymopterus globosus, Ferula multifida, Heracleum maximum, Ligusticum filicinum,
and Osmorhiza occidentalis (see Appendix C). The use of these plants was primarily medicinal,
and often the roots were used. Three Apiaceae species were used as food. It may be that the
Fremont may have ground roots of Apiaceae species on their ground stones medicinally for
ailments such as rheumatism, swellings, sore throats, and coughs (Appendix C).
Cyperacae sedge phytoliths and starches were found on all ground stones tested. Sedge
pollen has been found on ground stones from Woodard Mound (Richens 1983) and Hinckley
Mounds (Peterson 2016). Additionally, seeds indicative of the Cyperaceae family have been
found in the fill of Hinckley Mounds, Wolf Village, and Woodard Mound. I did not test any
Cyperaceae species for my typology. There are 48 Cyperaceae species found in Utah Valley
(Welsh et al. 1987:653-684), three of which of have documented ethnographic uses. Microfossils
likely coming from sedge roots were observed by Yost, and the roots of Scirpus acutus and
96
Scirpus maritimus were both consumed by the Goshute. It may be that the Fremont were also
processing Scirpus roots for food. Carex (sp) is a common sedge in the Utah Valley area, but
there is no documented evidence of its use. This does not mean that it may not have been used
prehistorically by the Fremont, only that none of the more historic groups who lived in the valley
and surrounding areas have any recollection of using this plant.
Evidence of Liliaceae was recovered as starch from all stones except for FS 16494.
Specifically, these starches may likely be from Calochortus or Fritillaria. There have been no
previous findings of this family in the valley from Fremont archaeological contexts. I did not test
any Liliaceae species for phytoliths. In the valley, there are 15 species that may have been
available to prehistoric and historic peoples (Welsh et al. 1987:800-811). Ten of these have
documented uses. In particular, the processing of the bulbs of Smilacina stellata, Veratrum
californicum, Zigadenus paniculatus, and Zigadenus venenosus were reported as being crushed,
often for medicinal purposes. It may be that the Fremont were crushing bulbs of Liliaceae
species on their stones for medicinal purposes, to cure ailment such as sprains, burns, and
rheumatism (Appendix C).
Pinaceae Pinus phytoliths were identified on FS 2357, 16494, and 16642. Pinus pollen
has been found on both ground stones and fill, coming from the sites Hinckley Mounds,
Smoking Pipe, Wolf Village, and Woodard Mound (see Appendix A). There are five species
found in the valley, three with documented ethnographic uses. I tested all three for the typology.
The nuts of Pinus edulis and Pinus monophylla were often ground into a flour, the primary
purpose of which was for food (Appendix D).
An epidermal phytolith indicative of Rosaceae Prunus was found on FS 2357. A seed of
Rosaceae Prunus had been found in the fill of Wolf Village (Dahle 2011). There are two Prunus
97
species in the valley (Welsh et al. 1987:537-539), and only one has documented ethnographic
uses: Prunus virginiana. I was able to test this plant for the typology. Prunus virginiana was
used both medicinally and for food. The bark would be ground medicinally to treat headaches
and head colds, and the berries would be mashed for food (Appendix D).
Poaceae phytoliths and starches were found on all six ground stones. Specifics include
Elymus, Stipea, Chloridoid, and Zea mays. Elymus starch has been found in teeth tartar from
Seamon’s Mound, Stipa caryopses have been found in the fill of Wolf Village and Woodard
Mound, and Zea mays kernels, pollen, and starches have been found throughout the valley in
fills, on ground stones, and even on teeth tartar (Appendix A). Eragrosis, Panicum, Phragmites,
and Sporobolus have also been identified at sites throughout the valley. Eighty-six grass species
are found in the valley (Welsh et al. 1987:684-788). Only sixteen species have documented
ethnographic uses (Appendix C). I tested seven of these species for phytoliths. Ethnographically,
three species were ground into flour for use in bread or mush: Panicum crus-galli, Sporobolus
airoides, Sporobolus cryptandrus Phytoliths indicative of Elymus were found on the ground
stones, yet there are no records of historic groups grinding Elymus for flour. Same with Stipea
and most Chloridoids. All Poaceae species were used for food. The sugary exudate from
Phragmites communis was also used medicinally. The Fremont likely used grasses for food as
well.
Typhaceae Typha starches were found on FS 219 and 2357. Typha pollen has been
recovered from ground stone, a ceramic bowl, and fill from sites such as Kay’s Cabin, Smoking
Pipe, and Woodard Mound (Appendix A). Two Typha species are found in the valley, and both
have documented ethnographic uses. I tested Typha latifolia for the typology. None of the
ethnographic sources record if Typha was ever ground or mashed, only that it was often prepared
98
roasted (Appendix E), which may have been a food source prepared by the Fremont using
grinding stones.
Conclusion
The creation of a regional phytolith typology and the use of ethnographic sources to
provide interpretations for plant use can improve our understanding of plant use by prehistoric
peoples. Specifically, the creation of a Utah Valley regional phytolith typology for the Utah
Valley Fremont that is based on plants with documented ethnographic medicinal and dietary
properties has added to what is known about Fremont plant use in the valley. Through this
ground stone analysis, additional evidences for the Fremont use of squash, chokecherry, and
pinyon pine have been found. The use of cacti has also been found.
Furthermore, this typology is valuable because it is narrow in scope, and the forms
observed in the typology have a higher chance of appearing in prehistoric contexts than forms
that have no known medicinal or dietary properties. There were few phytolith forms that I
observed on the ground stone that were not among the forms in the typology. The unrecognized
forms were often rare in count, and when shared with other archeobotanists, were also not
recognized. Regional phytolith typologies, such as the one created for the Utah Valley Fremont,
can increase understanding of prehistoric plant use.
If this work were to continue, Dr. Loreen Allphin in the Plant and Wildlife Sciences
Department and Scott Jensen with the Utah Valley BLM Shrub Science Lab would both be able
to assist in the gathering of plants to expand the phytolith typology. I do not believe that the
methods need to be changed.
99
Suggested future directions would be to test more native plants for phytoliths, to increase
what is known about plant phytolith production, to expand the typology, and to further explore
diagnostic forms. These plant types include those used by historic native groups listed in
Appendix C. Specific genera and species that come to mind are plants from the Elymus and
Prunus genera, as well as plants from the Apiaceae and Liliaceae family. Starches from this
family were found on several ground stones, yet I tested no plants from this family for the
phytolith typology. There may be phytoliths from species in this family that could be found
archaeologically.
Other future directions include testing more ground stone artifacts to see if the patterns
observed continue: that plants which were not ground would not be seen on ground stone
artifacts, in other words, that patterns in the presence and absence of particular plants continue,
especially on artifacts from sites throughout Utah Valley. It will also be important to test other
archaeological remains, such as soil samples and ceramic sherds, to see if the phytoliths observed
are different than those observed on the ground stone artifacts.
100
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121
Appendix A
This data table is an amalgamation of the archaeobotanical reports of twelve Utah Valley
Fremont archaeological sites. I list the identified plant as noted in the reports, and in a few cases
rectified discrepancies in naming. I note the nature of the remain, where the remain was found at
the site, the site the remain is from, and the botanical report where I found this information. For
consistency, all botanical remains akin to seeds, such as sunflower achenes and grass caryopsis,
are referred to as seeds. See Table 2.1.
Family/subfamily, genus,
Common Name
Adoxaceae Sambucus,
Elderberry
Amaranthaceae Amaranthus,
Pigweed
Amaranthaceae Atriplex,
Saltbush, orache
Botanical
Remain
Seed
Provenience
Site
Source
Fill
Richens 1983
Seed
Seed
Fruit and
seed
Seed
Coprolite
Fill
Fill
Amaranthaceae Cheno-ams,
Species and genus
unidentifiable
Pollen
Ground stone
artifact (GS)
Fill
Fill
GS, and Fill
Woodard
Mound
Spotten Cave
Wolf Village
Hinckley
Mounds
Woodard
Mound
Wolf Village
Wolf Village
Smoking Pipe
Hinckley
Mounds
Woodard
Mound
Kay’s Cabin
Dahle 2011
Scott 1984
Peterson 2016;
Puseman 2016
Richens 1983
Seed
Pollen
Pollen,
seed
Seed
Fill
Fill
Seed
Fill
Amaranthaceae
Chenopodium,
Goosefoot
Seed
Seed
Fill
Fill
Seed
Fill
Wolf Village
Hinckley
Mounds
Kay’s Cabin
Amaranthaceae Suaeda,
Seepweed
Apiaceae,
Parsley/carrot
Seed
Fill
Wolf Village
Starch
Pollen
Teeth
GS
Asteraceae
Pollen
Fill
Seamons Mound
Hinckley
Mounds
Smoking Pipe
122
Pearce 2012
Dahle 2011
Puseman 2016
Richens 1983
Cummings 2011
Puseman and
Cummings 2001
Dahle 2011
Puseman 2016
Puseman and
Cummings 2001
Dahle 2011
Yost 2009:6
Peterson 2016
Scott 1984
Family/subfamily, genus,
Common Name
Botanical
Remain
Pollen
Provenience
Site
Source
Fill and GS
Peterson 2016
Asteraceae Ambrosia,
ragweed
Asteraceae Artemisia,
Sagebrush
Pollen
Fill and GS
Pollen
Pollen
Pollen
GS
Fill
Fill and GS
Asteraceae, Artemisia
tridentata, sagebrush
Asteraceae, Chenopodium
berlandieri, pigweed
Asteraceae Cirsium,
Thistle
Asteraceae/ Compositae
Helianthus,
Sunflower
Fragments
fill
Hinckley
Mounds
Hinckley
Mounds
Wolf Village
Smoking Pipe
Hinckley
Mounds
Smoking Pipe
Seed
Fill
Puseman 2016
Seed
Fill
Hinckley
Mounds
Wolf Village
Seed
Fill
Wolf Village
Dahle 2011
Seed
Seed
Coprolite
Fill
Spotten Cave
Kay’s Cabin
Asteraceae, High-spine
Aster, rabbitbrush,
snakeweed, sunflower
Pollen
Pollen
Pollen
GS
Fill
Metate
Asteraceae Iva axillaris,
Poverty Weed
Asteraceae Liguliflorae,
dandelion
Asteraceae Low-spine
Ragweed, cocklebur
Seed
Coprolite
Wolf Village
Smoking Pipe
Woodard
Mound
Spotten Cave
Pearce 2012
Puseman and
Cummings 2001
Cummings 2011
Scott 1984
Richens 1983
Pollen
Pollen
Pollen
Pollen
Pollen
GS
Fill
GS
Fill
Metate
Asteraceae Taraxacum,
dandelion
Betulaceae, Birch
Betulaceae Alnus,
Alder
Boraginaceae Amsinckia,
Fiddleneck
Boraginaceae Cryptantha,
popcorn flowers
Brassicaceae, mustard
Pollen
Fill and GS
Pollen
Pollen
Pollen
Seed
Fill
GS
Fill
Fill
Seed
Fill
Pollen
Pollen
Pollen
GS
Fill
GS
Brassicaceae Brassica,
Mustard
Seed
Fill
123
Peterson 2016
Cummings 2011
Scott 1984
Peterson 2016
Billat 1985
Dahle 2011
Pearce 2012
Wolf Village
Smoking Pipe
Wolf Village
Smoking Pipe
Woodard
Mound
Hinckley
Mounds
Smoking Pipe
Wolf Village
Smoking Pipe
Wolf Village
Cummings 2011
Scott 1984
Cummings 2011
Scott 1984
Richens 1983
Woodard
Mound
Wolf Village
Smoking Pipe
Hinckley
Mounds
Wolf Village
Richens 1983
Peterson 2016
Scott 1984
Cummings 2011
Scott 1984
Dahle 2011
Cummings 2011
Scott 1984
Peterson 2016
Dahle 2011
Family/subfamily, genus,
Common Name
Brassicaceae Lepidium,
Pepperweed
Caryophyllaceae, carnation
family
Caryophyllaceae Silene,
Campion, catchfly
Cleomaceae Cleome,
Beeweed
Cucurbitaceae
Cucurbita, Squash
Cupressaceae
Juniperus, Juniper
Cupressaceae
Juniperus monosperma
One-seeded Juniper
Cyperaceae, sedge
Cyperaceae Scirpus,
Bulrush, tule
Ephedraceae
Ephedra nevadensis,
Mormon Tea
Euphorbiaceae
Euphorbia prostrata,
Creaping spurge
Fabaceae Leguminosae
Phaseolus,
bean
Fabaceae Leguminosae
Botanical
Remain
Seed
Provenience
Site
Source
Fill
Wolf Village
Dahle 2011
Seed
Fill
Richens 1983
Seed
Fill
Woodard
Mound
Wolf Village
Seed
Pollen
Uncharred
Rind
Pollen
Pollen
Pollen
Fill
Fill
Zone III
Wolf Village
Smoking Pipe
Spotten Cave
Dahle 2011
Scott 1984
Mock 1971
GS
Fill
Metate
Cummings 2011
Scott 1984
Richens 1983
Seed and
pollen
Seed
Seed
Fill and GS
Seed
Fill
Wolf Village
Smoking Pipe
Woodard
Mound
Hinckley
Mounds
Wolf Village
Woodard
Mound
Kay’s Cabin
Pollen
Fragments
GS
Fill
Wolf Village
Smoking Pipe
Pollen
Metate
Richens 1983
Pollen
Fill and GS
Seed
Seed
Coprolite
Fill
Seed
Seed
Fill
Fill
Pollen
GS
Woodard
Mound
Hinckley
Mounds
Spotten Cave
Hinckley
Mounds
Wolf Village
Woodard
Mound
Wolf Village
Seed
Fill
Likely introduced
Wolf Village
Dahle 2011
Welsh et al. 1987:303
Bean
Fill
Smoking Pipe
Charred
Pollen
Seed
Fill
GS
Fill
Wolf Village
Wolf Village
Woodard
Mound
Billat 19851; Forsyth
1984
Dahle 2011
Cummings 2011
Richens 1983
Fill
Fill
124
Dahle 2011
Peterson 2016;
Puseman 2016
Dahle 2011
Richens 1983
Puseman and
Cummings 2001
Cummings 2011
Billat 1985
Peterson 2016
Pearce 2012
Puseman 2016
Dahle 2011
Richens 1983: 111
Cummings 2011
Family/subfamily, genus,
Common Name
Fagaceae Quercus,
Oak
Malvaceae Sphaeralcea,
Globemallow
Nyctaginaceae Boerhaavia,
Spiderling
Papaveraceae Argemone,
Prickly poppy
Pinaceae Abies,
fir
Pinaceae Pinus,
pine
Plantaginaceae Plantago,
plantain
Poaceae,
Grass
Poaceae Eragrostis,
loveweed
Poaceae Panicum,
Panic grass
Poaceae Sporobolus,
dropseed
Poaceae Hordeum/Elymus,
wildrye
Poaceae Oryzopsis
hymenoides, Indian rice grass
Poaceae Phragmites,
reed
Poaceae Stipa hymenoides
Indian rice grass
Poaceae Zea mays,
corn
Botanical
Remain
Uncharred
bean
Pollen
Provenience
Site
Source
Coprolite
Spotten Cave
Pearce 2012
Fill
Smoking Pipe
Scott 1984
Seed
Fill
Wolf Village
Dahle 2011
Pollen
Fill
Smoking Pipe
Scott 1984
Seed
Fill
Wolf Village
Dahle 2011
Pollen
Pollen
Pollen
Pollen
Pollen
GS
Fill
GS
Fill
Fill and GS
Cummings 2011
Scott 1984
Cummings 2011
Scott 1984
Peterson 2016
Pollen
Metate
Pollen
Seed
Phytoliths
Pollen
Pollen
Seed and
pollen
Starch
Uncharred
seed
Seed
GS
Fill
Teeth
GS
Fill
Fill and GS
Wolf Village
Smoking Pipe
Wolf Village
Smoking Pipe
Hinckley
Mounds
Woodard
Mound
Wolf Village
Wolf Village
Seamons Mound
Wolf Village
Smoking Pipe
Hinckley
Mounds
Wolf Village
Spotten Cave
Puseman 2016
Seed
Fill
Hinckley
Mounds
Wolf Village
Seed
Fill
Wolf Village
Dahle 2011
Starch
Teeth
Seamons Mound
Yost 2009:6
Seed
Fill
Richens 1983
Seed
Fill
Seed
Fill
Woodard
Mound
Woodard
Mound
Wolf Village
Kernels,
cobs, stalk
Pollen
Kernels
Fill
Smoking Pipe
Fill
Fill
Smoking Pipe
Hinckley
Mounds
Billat 1985; Forsyth
1984
Scott 1984
Puseman 2016
GS
Coprolite
Fill
125
Richens 1983
Cummings 2011
Dahle 2011
Yost 2009
Cummings 2011
Scott 1984
Peterson 2016;
Puseman 2016
Cummings 2011
Pearce 2012
Dahle 2011
Richens 1983
Dahle 2011
Family/subfamily, genus,
Common Name
Botanical
Remain
Kernels
Kernels
Provenience
Site
Source
Fill
Fill
Dahle 2011
Richens 1983
Kernels,
cupule
Pollen
Pollen
Fill
Wolf Village
Woodard
Mound
Kay’s Cabin
Kernels
Fill
Starch,
phytoliths
Cob
Pollen
Pollen
Seed
Seed
Teeth
Seed
Fill
Uncharred
seed
Pollen
Coprolite
Wolf Village
Woodard
Mound
American Fork
Cave
Seamon’s
Mound
West Canyon
Wolf Village
Smoking Pipe
Wolf Village
Woodard
Mound
Hinckley
Mounds
Spotten Cave
GS
Wolf Village
Cummings 2011
Seed
Fill
Wolf Village
Dahle 2011
Seed
Seed
Fill
Fill
Dahle 2011
Puseman 2016
Portulacacea Portulaca,
Purslane
Ranunculus, Buttercup
Seed
Fill
Wolf Village
Hinckley
Mounds
Wolf Village
Pollen
GS
Peterson 2016
Rosaceae, Rose
Pollen
Pollen
Pollen
GS
Fill
GS
Rosaceae Amelanchier,
Service berry
Rosaceae Prunus,
Chokecherry, cherry, plum
Rosaceae Prunus Virginiana,
chokecherry
Rosaceae Rosa, rose
Seed
Fill
Hinckley
Mounds
Wolf Village
Smoking Pipe
Hinckley
Mounds
Wolf Village
Seed
Fill
Kay’s Cabin
Puseman and
Cummings 2001
Seed
Fill
Wolf Village
Dahle 2011
Seed
Fill
Kay’s Cabin
Puseman and
Cummings 2001
Polygonaceae Eriogonum,
Wild buckwheat
Polygonaceae Polygonum,
Knotweed
Polygonaceae Polygonum
bistortoides,
American bistort, Pursh
Polygonaceae Polygonum
lapathifolium, willowweed
Polygonaceae Rumex,
Dock
GS
Metate
Fill
GS
Fill
Fill
Fill
126
Puseman and
Cummings 2001
Cummings 2011
Richens 1983
Hansen 1941
Yost 2009
Wheeler 1968
Cummings 2011
Scott 1984
Dahle 2011
Richens 1983
Puseman 2016
Pearce 2012
Dahle 2011
Cummings 2011
Scott 1984
Peterson 2016
Dahle 2011
Family/subfamily, genus,
Common Name
Rosaceae Rubus,
Wild raspberry
Salicaeae Salix,
Willow
Botanical
Remain
Seed
Provenience
Site
Source
Fill
Wolf Village
Dahle 2011
Pollen
Metate
Richens 1983
Pollen
Pollen
Fill
Fill
Sapindaceae Acer,
Maple
Pollen
Pollen
GS
GS
Pollen
Metate
Sarcobataceae, formerly
Chenopodiaceae Sarcobatus,
greasewood
Pollen
Pollen
Pollen
GS
Fill
Fill and GS
Solanaceae, Potato
Solanaceae Physalis,
Ground cherry
Pollen
Seed
Uncharred
seed
Starch
Fill
Fill
Coprolite
Woodard
Mound
Smoking Pipe
Hinckley
Mounds
Wolf Village
Hinckley
Mounds
Woodard
Mound
Wolf Village
Smoking Pipe
Hinckley
Mounds
Smoking Pipe
Wolf Village
Spotten Cave
Teeth
Seamons Mound
Yost 2009
Pollen
Kay’s Cabin
Pollen,
seed
Seed
Pollen
Pollen
Ceramic
bowl
Fill and
GS
Fill
GS
GS
Puseman and
Cummings 2001
Peterson 2016;
Puseman 2016
Dahle 2011
Cummings 2011
Peterson 2016
Pollen
Pollen
Fill
Metate
Solanaceae Solanum jamesiitype, Wild potato
Typhaceae Typha,
Cattail
Typhaceae Typha latifolia,
Cattail
127
Hinckley
Mounds
Wolf Village
Wolf Village
Hinckley
Mounds
Smoking Pipe
Woodard
Mound
Scott 1984
Peterson 2016
Cummings 2011
Peterson 2016
Richens 1983
Cummings 2011
Scott 1984
Peterson 2016
Scott 1984
Dahle 2011
Pearce 2012
Scott 1984
Richens 1983
Appendix B
I researched the plants listed in the first column of the table in Appendix A to see if any
of the genera and families identified at Utah Valley Fremont archaeological sites had species that
were native to Utah County. In this table, I list all the species that are native to Utah County, and
what a cursory review of ethnographic reports say on whether or not that plant was used by
historic indigenous groups. Additionally, I include a source of where to find that plant in Welsh
et al. (1987, 2008).
Plants demarcated with a ^ are plants that were not identified at archaeological sites in
Utah Valley, but do appear significantly in ethnographies. Plants demarcated with a * represent
archaeobotanical remains that were identified at the species level at archaeological sites in Utah
Valley. No name synonyms are recorded in this appendix.
Scientific Name, Common Name
Adoxaceae Sambucus,
elderberry 1987:100
Alismataceae Sagittaria^,
arrowweed 1987:651
Amaranthaceae Suaeda, seepweed
1987:130
Amaranthaceae Amaranthus,
pigweed
1987:44-45
Amaranthaceae Atriplex,
saltbush, orache
1987:118-122
Amaranthaceae Cheno-ams,
species and genus unidentifiable
1987:44-46, 116-130
Amaranthaceae Chenopodium,
goosefoot,
1987:124-130
Used
Sambucus cerulea
Sambucus racemosa
Sagittaria latifolia
Not used
Suaeda calceoliformis
Suaeda torreyana
Amaranthus albus
Amaranthus
hypochondriacus
Amaranthus retroflexus
Atriplex canescens
Atriplex confertifolia
Atriplex powellii
Atriplex truncata
Allenrolfea occidentalis,
Ceratoides lanata
Salicornia europaea
Suaeda occidentalis
Chenopodium capitum var.
parvicapitum
Chenopodium fremontii
var. fremontii
128
Sagittaria cuneata
Amaranthus blitoides
Atriplex hortensis
Atriplex patula var. patula
Atriplex rosea
Grayia Spinosa
Kochia Americana
Monolepis nuttaliiana
Salicornia utahensis
Salsola iberica
Sarcobatus vermiculatus
Chenopodium ambrosioides
Chenopodium atrovirens
Chenopodium dessicatum
Chenopodium glaucum
Scientific Name, Common Name
Amaranthus Chenopodium
berlandieri* pigweed 2008:131
Anacardiaceae Rhus ^,
skunk bush 1987:46-47
Apiaceae,
parsley/carrot
1987:613-637
Asteraceae,
aster, daisy, sunflower
1987:131-240
Used
Chenopodium rubrum
Chenopodium berlandieri
Not used
Chenopodium hybridum
Aromatic var. trilobata
Angelica pinnata,
Carum/Perideridia
gairdneri
Cymopterus globosus
Cymopterus longpipe
Cymopterus purparascens,
Ferula multifida
Heracleum
maximum/lanatum,
Ligusticum filicinum
Orogenia linearifolia
Osmorhiza occidentalis
Achillea millefolium,
Agoseris aurantiaca var.
aurantiaca
Aster leucanthemifolius
(Machaerantherea
canescnes var.
leucanthemfolia)
Balsamorhiza hookeri var.
hispidula
Balsamorhiza sagittata
Brickellia grandiflora
Brickellia oblongifolia
Chaenactic alpine
Chaenactic douglasii
Chrysothamnus nauseosus
Chrysothamnus
viscidiflorus var.
viscidiflorus
Crepis acuminata
Crepis runcinata var.
glauca & var. hispidulosa
& var. Runcinata
Erigeron caespitosus
Erigeron speciosus var.
macranthus
Grindelia squarrosa var.
Squarossa
Gutierrezia sarothrae
Lactuca ludoviciana
129
Angelica roseana
Angelica wheekeri
Cicuta maculata
Conium maculatum – poisonous
Cymopterus hendersonii
Ligusticum porter
Lomatium ambiguum
Lomaticum grayi var. grayi
Lomatium juniperum
Lomatium kingii var. kingii
Lomatium triternatum var.
platycarpum
Osmorhiza chilensis
Sium suave
Zizia aptera
Agoseris glauca var. dasycephala,
& var. laciniata
Antennaria alpina
Antennaria dimorpha
Antennaria microphylla
Antennaria neglecta
Antennaria parviflora
Anthemis cotula
Arnica cordifolia
Arnica diversifolia
Arnica latifolia
Arnica longifolia
Arnica mollis
Arnica rydbergii
Aster brachyactis
Aster chilensis
Aster eatonii
Aster engelmannii
Aster falcatus
Aster foliaceus var. canbyi & var.
parryi
Aster frondosus
Aster glaucodes var. glaucodes
Aster hesperius
Aster kingii
Aster pauciflorus
Balsamorrhiza macrophylla
Bidens comosa
Bidens frondosa
Scientific Name, Common Name
Used
Lygodesmia grandiflora
var. dianthopsis
“Senecio”
Tetradymia canescens
Wyethia amplexicaulis
130
Not used
Brickellia californica
Brickellia microphylla var. watsonii
Chrysothamnus depressus
Chrysothamnus greenei
Chrysothamnus parryi var.
attenuatus
Chrysothamnus vaseyi
Conyza canadensis
Crepis atrabarba
Crepis intermedia
Crepis modocensis
Crepis nana
Crepis occidentalis var. costata
Erigeron arenoides
Erigeron argentatus
Erigeron composites
Erigeron coulteri
Erigeron divergens
Erigeron eatonii
Erigeron engelmannii
Erigeron garrettii
Erigeron glabellus
Erigeron goodrichii
Erigeron kachinensis
Erigeron lonchophyllus
Erigeron pumilus
Erigeron tener
Erigeron ursinus
Eupatorium maculatum
Gnaphalium chilense
Gnaphalium palustre
Gutierrezia microcephala
Haplopappus acaulis var. acaulis,
var. glabratus
Haplopappus lanceolatus
Haplopappus macronema
Haplopappus racemosus
Helenium autumnale
Helenium hoopesii
Heterotheca villosa var. foliosa,
var. hispida
Hieracium albiflorum
Hieracium cynoglossoides
Hymenopappus filifolius var.
nudipes
Hymenoxys grandiflora
Iva xanthifolia
Lactuca tatarica
Layia glandulosa
Leucelene ericoides
Scientific Name, Common Name
Used
Asteraceae Artemisia,
sagebrush
1987:145-150
Artemisia biennis
Artemisia carruthii
Artemisia dracunculus
Artemisia ludoviciana var.
incompta & var. latilopa
Artemisia michauxiana
Artemisia spinescens
Artemisia tridentate var.
pauciflora & var.
tridentata
Cirsium eatonii var eatonii
Cirsium undulatum var.
undulatum
Asteraceae Cirsium,
thistle 1987:171-175
Asteraceae Helianthus,
sunflower 1987:202-203
Helianthus annus
Helianthus uniflora,
131
Not used
Machaeranthera grindelioides var.
grindeliodes
Machaeranthera tanacetifolia
Madia glomerata
Microseris nutans
Perityle stansburyi
Rudbeckia occidentalis
Senecio amplectens
Senecio canus
Senecio crassulus
Senecio crocatus
Senecio dimorphophyllus var.
dimorphophyullus
Senecio eremophilus
Senecio fremontii var fremontii
Senecio hydrophilus
Senecio integerrimus
Senecio mulilopatus
Senecio serra var serra
Senecio streptanthifolius
Senecio triangularis
Senecio werneriifolius
Solidago multiradiata
Solidago nana
Solidago occidentalis
Solidago parryi
Solidago sparsiflora
Sphaeromeria diversifolia)
Stephanomeria exigua
Tetradymia spinosa
Townsendia florifer
Viguiera ciliata
Viguiera multiflora var. multiflora
Artemisia cana,
Artemisia frigid
Cirsium neomexicanum var.
utahense
Cirsium scariosum var. scariosum
Cirsium vulgare
Helianthus nuttallii
Scientific Name, Common Name
Asteraceae Iva axillaris*,
poverty weed 1987:209
Asteraceae Liguliflorae,
Cichorioideae
Asteraceae Ambrosia,
ragweed 2008:148-150
Asteraceae Taraxacum,
dandelion 2008:270
Betulaceae,
birch 2008:49-52
Betulaceae Alnus,
alder 1987:57
Boraginaceae Amsinckia,
fiddleneck 1987:60-61,84
Boraginaceae Cryptantha,
popcorn flowers
1987:64-75
Brassicaceae,
mustard
2008:289-348
Used
Iva axillaris
Not used
Subfamily in Asteraceae.
See general Astereaceae
listing.
Ambrosia psilostachya
Not found in Welsh et al. 1987 or
2008
Amsinckia tessellata
Descurainia pinnata var.
filipes,
Descurainia richardsonii
var. sonnei
Descurainia richardsonii
var. sonnei
Draba nemorsa
132
Ambrosia acanthicarpa
Ambrosia artemisiifolia
Taraxacum laevigatum
Betula glandulosa
Betula occidentalis
Alnus incana
Alnus serrulata
Amsinckia menziesii
Cryptantha affinis
Cryptantha flavoculata
Cryptantha gracilis
Cryptantha humilis
Cryptantha mensana
Cryptantha torreyana
Cryptantha watsoni
Arabis drummondii
Arabis glabra
Arabis hirsuta
Arabis holboellii var. pinetorum,
var. secunda
Arabis lyallii
Arabis microphylla
Arabis perannans
Arabis selbyi
Arabis sparsiflora
Barbarea orthoceras
Cardamine breweri
Cardamine cordifolia
Chlorocrambe hastatus
Descurainia californica
Draba aurea
Draba brachystylis
Draba cunefolia
Draba denisfolia
Draba lanceolata
Draba lonchocarpa
Draba oligosperma var.
oligosperma
Drapa reptans
Draba stenoloba
Draba verna
Erysium asperum
Scientific Name, Common Name
Brassicaceae Brassica,
mustard 1987:254-255,259
Brassicaceae Lepidium,
pepperweed
1987:257, 271-274
Used
Not used
Hutchinsia procumbens
Lesquerella garretti
Lesquerella hemiphysaria var.
hemiphysaria
Lesquerella utahensis
Rorippa curvipes var. alpina, var.
curvipes, var. integra
Rorippa islandica var. glabra, var.
hispida
Rorippa sphaerocarpa
Smelowskia calycina
Strephtanthus cordatus
Thelypodiopsis sagittata var.
sagittata
Thelypodiopsis vermicularis
Thelypodium integrifolium var.
integrifolium
Thelypodium laxiflorum
Thlaspi montanum
According to Welsh et al., all species are introduced.
Lepidium lasiocarpum var.
lasiocarpum
Cactaceae Opuntia^,
prickly pear 1987:88-91
Caryophyllaceae,
carnation
1987:101-114
Opuntia polycantha var.
polycantha
Arenaria congesta var.
congesta
Caryophyllaceae Silene,
campion, catchfly
1987:108-112
Cleomaceae Cleome,
beeweed
1987:97-98,284
Cucurbitaceae Cucurbita,
squash 1987:290-291
Silene acaulis
Silene douglasii
Silene menziesii
Cleome serrulata var.
serrulata
Cleome pinnata
Cucurbita maxima
Cucurbita moschata
133
Lepidium campestre
Lepidium densiflorum var.
densiflorum, var. pubicarpum, var.
ramosum
Lepidium integrifolium
Lepidium monatum var. jonesii, var.
montanum
Lepidium virginicum
Opuntia erinaceae var. utahensis
Opuntia fragilis
Arenaria fendleri var. glabrescens
Arenaria hookeri
Arenaria macradenia
Arenaria nuttallii
Arenaria rubella
Cerastium arvense
Cerastium beeringianum
Stellaria jamesiana
Stellaria longifolia
Stellaria obtuse
Cleome docecandra
Scientific Name, Common Name
Cupressaceae Juniperus,
juniper 1987:26-27
Cupressaceae Juniperus
monosperma*,
one-seeded juniper 1987:26
Cyperaceae,
sedge
1987:653-684
Cyperaceae Scirpus,
bulrush, tule
1987:680-684
Used
Juniperus communis
Juniperus osteosperma
Juniperus scopulorum
Scirpus acutus
Scirpus maritimus
Scirpus validus
134
Not used
Juniperus horizontalis
Juniperus monosperma
Carex aquatilis
Carex atherodes
Carex athrostachya
Carex atrata var. chalciolepis
Carex aurea
Carex backii
Carex capillaries
Carex disperma
Carex douglasii
Carex egglestonii
Carex elynoides
Carex geyeri
Carex haydeniana
Carex hoodii
Carex lanuginosa
Carex lenticularis
Carex microptera
Carex nebrascensis
Carex nova
Carex obtusata
Carex occidentalis
Carex parryana
Carex petasata
Carex phaeocephala
Carex praegracilis
Carex raynoldsii
Carex rossii
Carex rostrata
Carex stenophylla
Carex straminiformis
Carex vallicola
Cyperus aristatus
Cyperus erythrorhizos
Eleocharis acicularis
Eleocharis parishii
Eleocharis pauciflora
Scirpus americanus
Scirpus microcarpus
Scirpus ovatus
Scirpus palustris
Scirpus parvulus
Scirpus rostellatus
Scirpus spadiceus
Scientific Name, Common Name
Used
Elaeagnaceae^ Shepherdia^,
buffaloberry 1987:295
Ephedraceae Ephedra
nevadensis*, Mormon tea
1987:28-29
Ericaceae Arctostaphylos,
manzanita 1987:297
Euphorbiaceae Euphorbia
prostrata*,
creeping spurge
2008:358-364
Shepherdia canadensis
Shepherdia argentea
Ephedra nevadensis – not
found in Utah County
Ephedra virdis var. virdis
Arctostaphylos patula
Fabaceae,
bean,
1987:336-411
Glycyrrhiza lepidota
Hedysarum var. boreale
Astragalus agrophyllus
var. agrophyllus, var.
martini
Euphorbia prostrata (pp.
360-361) was introduced
from the eastern U.S.
135
Not used
Scirpus pallidus
Scirpus pungens
Euphorbia brachycera
Euphorbia dentata
Euphorbia glyptosperma
Euphorbia maculata
Euphorbia occellata
Euphorbia serpyllifolia
Astragalus agrestis
Astragalus beckwithii var.
beckwithii
Astragalus bisculatus var.
haydenianus
Astragalus calycosus var. calycosus
Astragalus canadensis var.
canadensis
Astragalus cibarius
Astragalus drummondii
Astragalus eurekensis
Astragalus kentrophyta var.
implexus
Astragalus lutosus
Astragalus miser var. oblongifolius
(poisonous)
Astragalus oophorus var.
caulescens
Astragalus scopulorum
Astragalus tenellus
Gleditsia triacanthos
Lathyrus brachycalyx var.
brachycalyx
Lathyrus lanszwertii var.
laetivirens, var. lanszwertii
Lathyrus pauciflorus
Lotus utahensis
Lupinus var. argenteus, var.
rubricaulis
Lupinus brevicaulis
Lupinus caudatus var. utahensis
Lupinus Lepidus var. utahensis
Lupinus polyphyllus var.
prunophilus
Scientific Name, Common Name
Used
Not used
Lupinus sericeus var. sericeus
Oxytropis viscida
Thermopsis montana
Trifolium gymnocarpon
Trifolium kingie
Trifolium longpipes var. reflexum
Vicia americana var. americana
Fabaceae Phaseolus,
bean 1987:400
Fagaceae Quercus,
oak 1987:304-306
Liliaceae^,
lily,
1987:800-811
Phaseolus vulgaris
Malvaceae Sphaeralcea,
globemallow
1987:420-424
Nyctaginaceae Boerhaavia,
spiderling 2008:516
Papaveraceae Argemone,
prickly poppy 1987:451
Pinaceae Abies,
fir 1987:29-31
Pinaceae Pinus,
pine 1987:31-33
Pinaceae Pinus edulis*,
pinyon pine 1987:32
Plantaginaceae Plantago,
plantain
1987:454-455
Poaceae Eragrostis,
lovegrass
2008:852-854
Poaceae, Graminae,
grass
1987:684-788
Sphaeralcea grossulariifolia var.
grossulariifolia
Sphaeralcea rivularis
No species found in Utah County.
Quercus gambelii
Quercus pacuiloba
Allium acuminatum
Allium biceptrum
Calochortus nuttallii
Fritillaria atropurpurea
Fritillaria pudica
Smilacina racemosa
Smilacina stellate
Veratrum californicum
Zigadenus paniculatus
Zigadenus venenosus
Sphaeralcea coccinea
Sphaeralcea munroana
Androstephium breviflorm
Disporum trachycarpum
Erythronium grandiflorum
Triteleia grandiflora
Zigadenus elegans
Argemone munita
Abies concolor
Abies lasiocarpa
Pinus flexilis
Pinus monophylla
Pinus edulis
Bromus carinatus
Cinna latifolia
Deschampia cespitosa
Festuca octoflora
Festuca ovina
Poa fendleriana
Puccinellia nuttalliana
Trisetum spicatum
136
Abies engelmanii (Picea
engelmanii; Picea glauca)
Pinus longaeva
Pinus ponderosa
Plantago elongata
Plantago eripoda
Plantago tweedyi
Eragrostis hypnoides
Eragrostis Mexicana
Eragrostis pectinaceae
Agrostis exarata
Agrostis humilis
Alopecurus aequalis
Aristida purpurea
Bouteloua gracilis
Bromus anomalus
Bromus catharticus
Bromus ciliates
Calamagrostis canadensis
Scientific Name, Common Name
Used
Poaceae Hordeum/Elymus,
wildrye, wheatgrass,
1987:723-728, 740-742
Elymus cinerus
Elymus glaucus
Hordeum jubatum
Poaceae Oryzopsis,
Indian rice grass
137
Not used
Calamagrostis scorpulorum
Calamagrostis stricta
Cenchrus longispinus
Danthonia californica
Danthonia intermedia
Deschampsia elongate
Distichlis spicata
Festuca rubra
Festuca sororia
Festuca thurberi
Glyceria borealis
Glyceria grandis
Glyceria striata
Koeleria macrantha
Leersia oryzoides
Leptochloa fascicularis
Leucopoa kingie
Melica bulbosa
Melica spectablis
Muhlenbergia andina
Muhlenbergia filiformis
Muhlenbergia mexicana
Muhlenbergia racemosa
Muhlenbergia richardsonis
Paspalum distichum
Phalaris arundinaceae
Phleum alpinum
Poa alpine
Poa curta
Poa glauca var. glauca
Poa leptocoma
Poa nervosa
Poa palustris
Poa reflexa
Poa secunda
Polypogon interruptus
Spartina gracilis
Sphenopholis obtusata
Trisetum wolfii
Elymus lanceolatus
Elymus salinus
Elymus scribneri
Elymus smithii
Elymus spicatus
Elymus traachycaulus
Elymus tritcoides
Elymus virginicus
Hordeum brachyantherum
Oryzopsis bloomerii
Stipa comata
Scientific Name, Common Name
1987:754,782-786
Used
Poaceae Panicum,
panic grass
1987:755-757,776,720, 719,701
Panicum crus-galli
Poaceae Phragmites,
reed 1987:759
Poaceae Sporobolus,
dropseed
1987:779-781,752,749
Poaceae Stipa hymenoides*,
Indian rice grass 1987:783
Poaceae Zea mays*, Corn,
Not found in Welsh et al.
Polygonaceae Eriogonum,
wild buckwheat
1987:474-487
Phragmites australis
(Phragmites communis)
Sporobolus airoides var.
airoides
Sporobolus cryptandrus
Stipa hymenoides
(Oryzopsis hymenoides)
Zea mays
Eriogonum microthecum
Eriogonum ovalifolium,
Eriogonum umbellatum
var. umbellatum
Polygonaceae Polygonum,
knotweed
1987:487-490
Polygonaceae Polygonum
bistortoides*,
American bistort 1987:489
Polygonaceae Polygonum
lapathifolium*,
willowweed 1987:490
Polygonaceae Rumex,
dock 1987:491-493,487
Portulacacea Portulaca,
purslane 1987:496
Ranunculus,
buttercup
2008:610-628
Not used
Stipa nelsonii
Stipa pinetorum
Panicum acuminatum
Panicum capillare
Panicum syzigachne
Panicum virgatum
Sporobolus asper
Sporobolus giganteaus
Eriogonum brevicaule var.
brevicaule var.laxifolium, var.
nanum
Eriogonum heracleoides
Eriogonum hookeri
Eriogonum jamesii
Eriogonum racemosum var.
racemosum
Polygonum amphibium
Polygonum aviculare
Polygonum douglasii var. douglasii
Polygonum persicaria
Polygonum ramosissimum
Polygonum sawatchense
(Polygonum douglasii var.
johnstonii)
Polygonum bistortoides
Adventive from Europe
Rumex salicifolus
Rumex venosus
Aquilegia coerulea
Ranunculus aqualitis var.
diffusus
Thalictrum fendleri
138
Portulaca oleracea
Aconitum columbianum
Actaea rubra
Anemone multifida
Aquilegia barnebyi
Aquilegia flavescens
Caltha leptosepala
Clematis columbiana
Clematis hirsutissima
Scientific Name, Common Name
Used
Rosaceae,
rose
1987:519-543
Cercocarpus ledifolius
Crataegus douglassii var.
rivularis
Fragaria vesca
Geum macrophyllum
Holodiscus dumosus
Petrophytum caespitosum
Potentilla glandulosa var.
intermedia
Purshia Mexicana
Purshia tridentata
Rosaceae Ameliancher,
service berry 1987:521
Rosaceae Prunus,
chokecherry, cherry, plum
1987:537-539
Rosaceae Prunus Virginiana*,
chokecherry 1987:539
Rosaceae Rosa,
rose 1987:540-541
Rosaceae Rubus,
wild raspberry 1987:541-542,528
Salicaeae Salix,
willow
Ameliancher alnifolia
Ameliancher utahensis
Not used
Clematis occidentalis
Delphinium barbeyi
Delphinium nuttallianum
Delphinium occidentale
Delphinium scaposum var.
andersonii
Myosurus apetalus
Ranunculus adoneus
Ranunculus flammula
Ranunculus jovis
Ranunculus macounii
Ranunculus orthorhynchus
Ranunculus sceleratus
Thalictrum occidentale
Thalictrum sparsiflorum
Cercocarpus intricatus
Cercocarpus montanus
Crataegus succulent
Fragaria virginiana
Geum aleppicum
Geum rossii
Ivesia gordonii
Ivesia utahensis
Physocarpus alternans
Physocarpus malvaceus
Physocarpus monogynus
Potentilla anserine
Potentialla arguta
Potentialla fruticosa
Potentialla gracilis var.
brunnescens, var.elmeri, var.
pulcherrima
Potentialla norvegiiva
Potentialla paradoxa
Sibbaldia procumbens
Sorbus scopulina
Prunus emarginata
Prunus virginiana
Rosa woodsii
Rubus leucodermis
Rubus parviflorus
Salix amygdaloides
Salix exigua
139
Rosa eglanteria
Rosa nutkana
Rubus idaeus
Salix artica
Salix bebbiana
Scientific Name, Common Name
1987:551-556
Used
Sapindaceae Acer,
maple 1987:41-42
Sarcobataceae Sarcobatus,
greasewood 1987:129
Solanaceae,
night shade, potato
2008:726-733
Acer negundo
Solanaceae Physalis,
ground cherry 1987:607-608
Solanaceae Solanum jamesii-type*,
wild potato 1987:609-610
Typhaceae Typha,
cattail 1987:823
Typhaceae Typha latifolia*, cattail
1987:823
Physalis longifolia
Sarcobatus vermiculatus
Nicotiana attenuata
Solanum tuberosum
Solanum jamesii-type (not
found in Utah valley)
Typha angustifolia var.
domingensis
Typha latifolia
140
Not used
Salix boothii
Salix brachycarpa
Salix drummondiana
Salix geyeriana
Salix lasiandra
Salix lasiolepis
Salix lutea
Salix reticulata
Salix scouleriana
Acer glabrum
Acer grandidentatum
Petunia hybrid
Solanum rostratum
Solanum sarrachoides
Solanum triflorum
Appendix C
This table is of the plants with documented ethnographic use by historic indigenous
groups who occupied Utah Valley and the surrounding regions. Specifically, this table is of
plants that I did not sample for the phytolith typology. See Appendix D for ethnographic uses of
the plants I sampled for the typology. Select species name synonyms are noted in parenthesis.
Family, Species
Alismataceae
Sagittaria latifolia^
Amaranthaceae
Allenrolfea occidentalis
Amaranthaceae
Amaranthus albus
Amaranthaceae
Amaranthus
hypochondriacus
Amaranthaceae
Amaranthus retroflexus
Common Name,
Source
arrowweed
1987:651
iodine bush,
pickelweed
1987:117
pale amaranth,
1987:44
grain amaranth,
princes feather
1987:45
redroot pigweed
1987:45
Amaranthaceae
Atriplex canescens
four-wing
saltbush
1987:118
Amaranthaceae
Atriplex confertifolia
shadscale
1987:118
Amaranthaceae
Atriplex powellii
powell orach
1987:121
Amaranthaceae
Ceratoides lanata
(Erotia lanata)
Amaranthaceae
Chenopodium capitum
var. parvicapitum
Amaranthaceae
Chenopodium fremontii
var. fremontii
winterfat,
white sage
1987:123
strawberry
spinach
1987:124
Fremont
goosefoot
1987:125
Documented Ethnographic Use
Goshute used the fruit for food (Fowler 1986:69).
Tubers were eaten boiled or roasted (Yanovsky
1936:7).
Seeds used for food by the Goshute and Southern
Paiute (Fowler 1986:72). Seeds were ground and made
into bread or mush (Yanovsky 1936:21).
Seeds used for food by the Utah Southern Paiute
(Fowler 1986:69; Rainey and Adams 2004).
Seeds used for food by the Western Shoshone, and
Southern Paiute (Fowler 1986:70; Rainey and Adams
2004).
Seeds collected in autumn were eaten raw or ground
into flour for cakes, or boiled. Young shoots and stems
were eaten raw or boiled by the Ute (Callaway et al.
1986:338). Seeds used for food and above ground plant
eaten as greens by Utah Southern Paiute (Fowler
1986:70; Rainey and Adams 2004).
Seeds used for food by the Goshute and Southern
Paiute (Chamberlin 1964:363; Fowler 1986:72). Fresh
roots and salt used as a medicinal physic by the
Shoshone (Train et al. 1941:50).
Seeds used for food by the Goshute and Southern
Paiute (Chamberlin 1964:340, 363; Fowler 1986:72;
Rainey and Adams 2004).
Seeds used for food, often ground into a meal for bread
or mush, by the Southern Paiute (Fowler 1986:72;
Rainey and Adams 2004).
Used medicinally for intermittent fevers by the
Goshute (Chamberlin 1964:352, 369).
Seeds gathered in large supplies and used for food by
the Goshute (Chamberlin 1964:366).
Leaves used as greens (Yanovsky 1936:22).
Seeds and leaves used for food by the Southern Paiute
(Fowler 1986:73).
141
Family, Species
Amaranthaceae
Chenopodium rubrum
Amaranthaceae
Salicornia europaea
Common Name,
Source
red goosefoot
1987:126
annual samphire
1987:128
Amaranthaceae
Suaeda calceoliformis
broom seepweed
1987:130
Amaranthaceae
Suaeda torreyana
torrey seepweed
1987:130
Apiaceae
Angelica pinnata
small-leaved
angelica
1987:617
false yarrow
1987:636
Apiaceae
Carum gairdneri
(Perideridia gairdneri)
Apiaceae
Cicuta maculata
Apiaceae
Conium maculatum
Apiaceae
Cymopterus globosus
(C. montanus)
water hemlock
1987:618
poison hemlock
1987:619
golfball
springparsley
1987:622
Apiaceae
Cymopterus longpipe
long-stalk
springparsley
1987:623
widewing
springparsley
1987:624
giant lomatium
1987:629
Apiaceae
Cymopterus
purparascens
Apiaceae
Ferula multifida
(Leptotaenia multifida;
Lomatium dissectum)
Documented Ethnographic Use
Gathered and eaten by the Goshute like Chenopodium
capitum (Chamberlin 1964:366).
Seeds used for food by the Goshute who cooked meal
from seeds tasted like “sweet bread” (Fowler 1986:73;
Chamberlin 1964:380).
Seeds used for food by the Goshute and Paiute, the
latter would clean and grind the seeds into flour for
biscuits (Chamberlin 1964:383; Palmer 1878:653).
Seeds used for food by the Utah Southern Paiute
(Fowler 1986:73). The Shoshone and Paiute made a tea
out of leaves for bladder and kidney troubles. The fresh
plants were crushed and rubbed on itchy sores like
chicken pox (Train et al. 1941:95).
Roots used as medicine by the Goshute (Chamberlin
1964:361).
The sweet, pleasant, nutritious, and starchy roots were
eaten cooked in pits overnight or boiled, and were also
cached for winter, Goshute (Chamberlin 1964: 339,
365). Roots were also eaten by the western, northern,
and eastern Shoshone (Fowler 1986:71).
No data yet – poisonous.
No data yet – poisonous.
Seeds and underground parts eaten, but not the leaves,
by the Goshute (Chamberlin 1964:367).
Boiled root water used as insecticide by the Paiute to
kill mites on chickens (Train et al. 1941: 42).
Leaves of the early spring plant were consumed, often
boiled by the Goshute and Ute (Chamberlain 1909:33;
1964:338, 367).
Seeds used for food by the Goshute (Fowler 1986:70).
The Ute would apply the pulped root to wounds and
bruises, and would create smoke of the root for
distemper in horses (Chamberlin 1909:34). The
Goshute used this plant medicinally and for food.
Medicinally, the mashed roots were used for wounds
and bruises, and for distemper in horses. The seeds
were eaten, but the mature plant was deemed inedible
because it was “too strong to taste” (Chamberlin
1964:338, 348, 369). Some considered this plant
poisonous, and others ate the young sprouts (Yanovsky
1936:48).
142
Family, Species
Common Name,
Source
Apiaceae
Heracleum maximum
(H. lanatum)
cow parsnip
1987:626
Apiaceae
Ligusticum filicinum
fernleaf
ligusticum
1987:627
Indian potato
1987:634
western
sweetcicely
1987:635
Apiaceae
Orogenia linearifolia
Apiaceae
Osmorhiza occidentalis
Documented Ethnographic Use
The root was seen as a panacea among the Paiute and
Shoshone, who used it for coughs, colds, hay fevers,
bronchitis, influenza, pneumonia, and tuberculosis. The
root was peeled, sliced, and dried for winter months.
They would mix the roots with Osmorhiza occidentalis
to make a decoction drunk as a tea for colds, sore
throat, and influenza. When boiled with the terminal
twigs of Juniperus utahensis, it became a tea for
influenza. Raw root was chewed for sore throats. When
combined with Leptotaenia roots and Achillea
lanulosa, it became a tea for gonorrhea. When boiled
alone, or with Osmorhiza occidentalis, or Rumex
venosus, it was a tea for veneral diseases. The roots
were used as an antiseptic, an external wash for
smallpox, as a healing agent for rashes, cuts, or sores.
The pulped raw root pulped was applied directly to
cuts, umbilical cords of newborns, or fresh slices
placed on sores. A poultice of raw or boiled roots was
applied to swellings sprains, or rheumatism. An oily
sap from fresh roots was used for trachoma or
gonorrheal infections of the eye. Dried root,
sometimes mixed with Pinus monophylla root, was
smoked for congestion and asthma (Train et al.
1941:65-67).
This plant was used by the Shoshone and Paiute. These
groups used the raw root for toothache and cavity pain,
or they would mash the root, soak it in water, and then
gargle the water for a sore throat while applying the
poultice around the throat. Mashed roots were also
used for rheumatism, and root slave for healing
wounds. A root tea was taken for diarrhea or for longer
for tuberculosis. A root decoction when mixed with
burnt whiskey was taken for coughs and chest colds.
The inhaling of smoke from the root coupled with
Pinus monophylla pitch was for head colds (Train et al.
1941:57).
Root occasionally used as a cough remedy by the
Paiute (Train et al. 1941:67).
Bears sometimes eat the bulbs of this plant, according
to the Goshute (Chamberlin 1964:375)
Used much like Leptotaenia multifida. The Paiute and
Shoshone primarily used the roots of this plant. A root
decoction was made for fevers, to regulate menstrual
disorder, for colds, pulmonary disorders, pneumonia,
influenza, and when consumed for a long period of
time, for venereal disease. A decoction was sometimes
prepared by soaking the roots in cold water.
143
Family, Species
Asteraceae
Agoseris aurantiaca var.
aurantiaca (Troximon
aurantiacum)
Asteraceae
Ambrosia psilostachya
Asteraceae
Artemisia carruthii
Asteraceae
Artemisia michauxiana
(Artemisia discolor)
Asteraceae
Artemisia spinescens
Asteraceae
Aster leucanthemifolius
(Machaerantherea
canescens var.
leucanthemfolia)
Common Name,
Source
orange agoseris
1987:137
Documented Ethnographic Use
Hot teas of the root were for fever, diarrhea, a palliative
for stomachaches, gas pains, indigestion, and as a
physic, with strength depending on brew concentration.
A root tonic was for boils, and when sugar was added
for colds and sore throats. This hot tea was also used
for whooping cough. The treatments were enhanced by
adding Artemisia gnaphalodes and Leptotaenia
multifida. A hot wash applied externally for lice on
humans, or chicken lice. An external antiseptic for
measles, venereal sores, skin rashes, and eyewash. Raw
roots were applied for sores, cuts, bruises, swellings,
snake bites. Raw roots were chewed for sore throat,
applied to toothaches, smoked for colds, or inserted
into nostrils for headaches (Train et al. 1941:73-74).
Leaves used for food by the Goshute, Ute, and
Northern Ute (Chamberlin 1964:338, 383; 1909:36;
Fowler 1986:71).
western rageweed The Goshute would make remedies for sore eyes by
1987:139
steeping leaves in hot water and bandaging them over
the eye (Chamberlin 1964:361).
carruth
Seeds used for food by the Utah Southern Paiute
wormwood
(Fowler 1986:71).
1987:147
michaux
Seeds used for food by the Goshute (Fowler 1986:71;
wormwood
Chamberlain 1964:362).
1987:148
budsage
This plant was used medicinally by the Paiute and
1987:150
Shoshone. They would make a green leaf or young
branch poultice for swellings. Green leaves were mixed
and mashed with chewing tobacco for sores or bruises,
and bedridden individuals were rubbed daily.
Additionally, this poultice was used for nosebleeds. A
whole plant poultice, fresh or boiled, was used for
rashes and itches. Mashed leaves were used to draw out
boils. A boiled branch tea was for hemorrhages,
especially from tuberculosis. A fresh flower and leaf
cold tea was for the bladder, and a similar hot tea for
stomach troubles, cramps, or indigestion. A root hot tea
for chest congestion, coughs, or colds. A whole plant
tea for influenza or as a wash. Boiled stems and leaves
were used as a wash for rheumatism, rashes, or skin
irritations (Train et al. 1941:28).
hoary tansy aster The Shoshone would boil the whole plant and take it
1987:213
twice a week as a blood tonic. They brewed the tops
into physics (Train et al. 1941:31).
144
Family, Species
Asteraceae
Balsamorhiza hookeri
var. hispidula
Asteraceae
Brickellia grandiflora
Asteraceae
Brickellia oblongifolia
Asteraceae
Chaenactic douglasii
Asteraceae
Chrysothamnus
viscidiflorus var.
viscidiflorus
Common Name,
Source
hooker
balsamroot
1987:158
tasselflower
1987:160
Mohave
brickellbush
1987:161
Douglas dustymaiden
1987:163
Documented Ethnographic Use
Seeds used for food by the Goshute and Eastern
Shoshone (Chamberlin 1964:363; Fowler 1986:71).
The Goshute would mix these seeds with meal made of
seeds from other plants made cakes better because they
acted as a baking powder. However, use with care as
overuse can be poisonous. The roots were also used as
a medicine (Chamberlin 1964:364).
Stems and leaves boiled and taken as a stomach
medicine by the Shoshone (Train et al. 1941:34).
The Goshute used this plant as follows: “Minced or
mashed and rubbed on limbs, etc., for soreness or
aching” (Chamberlin 1964:365). The Paiute and
Shoshone used crushed the leaves into a poultice
against swelling. They would boil leaves or the whole
plant into a drink for coughs or colds, or for indigestion
or sour stomach. Leaves were also used for snake bites
(Train et al. 1941:36-37).
viscid rabbitbrush The Shoshone and Paiute boiled young shoots, or
1987:169
crushed and soaked leaves, for a drink for colds. The
boiled plant with roots of Leptotaenia multifida used
for influenza. A stem and leaf poultice made for
rheumatism, and mashed leaves for tooth cavities and
toothaches (Train et al. 1941:38). Chewing gum made
from roots (Yanovsky 1936:60).
Asteraceae
Chrysothamnus
viscidiflorus var.
stenophyllus
(Bigelovia douglasii)
Asteraceae
Cirsium eatonii var
eatonii (Cnicus eatonii)
Asteraceae
Cirsium undulatum var.
undulatum
Asteraceae
Crepis acuminata
viscid rabbitbrush Chewing gum made from roots by the Goshute
1987:169
(Chamberlin 1964:344, 364).
Asteraceae
Crepis runcinata var.
glauca & var.
meadow
hawksbeard
1987:177-78
Eaton thistle
1987:172
gray thistle
1987:174
mountain hawksbeard
1987:176
Stems were used for food by the Goshute (Fowler 71).
They also used this plant on wounds, cuts, or sores
(Chamberlin 1964:349, 366).
Stems used for food by the Goshute and Eastern
Shoshone (Fowler 71986:1).
The Shoshone believed that the fine dust from the
ground root when sprinkled into eye would remove
foreign objects. The crushed the whole into a poultice
that would be applied on breasts induce milk flow after
childbirth, or to relieve sore breasts (Train et al.
1941:41).
Leaves used for food by the Goshute (Fowler 1986:71).
145
Family, Species
hispidulosa & var.
runcinata
Asteraceae
Erigeron caespitosus
Common Name,
Source
Documented Ethnographic Use
tufted flea bane
1987:184
Asteraceae
Erigeron speciosus var.
macranthus
(Erigeron grandifloras)
Asteraceae
Grindelia squarrosa var.
squarrosa
Asteraceae
Helianthella uniflora
Oregon daisy
1987:190
A cool solution from the boiled roots was applied as an
eyewash among the Paiute. They also made a boiled
root tea to stop diarrhea (Train et al. 1941:46).
Roots used in arrow poison and used as a fleabane
among the Goshute (Chamberlin 1964:368).
curly gumweed
1987:197
Roots made into cough medicine by the Utes
(Chamberlin 1909:34, 1964:371).
onehead
sunflower
1987:203
Asteraceae
Helianthus annus
common
sunflower
1987:203
Asteraceae
Iva axillaris*
poverty weed,
devil’s weed*
1987:209
Asteraceae
Lactuca ludoviciana
(Sonchus ludoviciana)
Asteraceae
Lygodesmia grandiflora
(Erthremia grandiflora)
var. dianthopsis
Asteraceae
Senecio amplectens
S. canus
S. crassulus
S. crocatus
S. dimorphophyllus
S. eremophilus
prairie lettuce
1987:210
A root poultice, sometimes heated, was applied on
swellings and sprains, and when not heated was used
for rheumatism of shoulder or knee. A mashed root
cold water infusion was used as a wash or cold
compress for headaches by the Paiute and Shoshone
(Train et al. 1941:56).
Seeds were highly prized food source that was often
made into oil made (Chamberlin 1964:371). Seeds
were eaten parched, raw, or into cakes by the Eastern
Shoshone, Utah Southern Paiute (Fowler 1986:71;
Rainey and Adams 2004). The Paiute made a root
decoction for rheumatism (Train et al. 1941:56).
Occasionally used as medicine by the Ute (Chamberlin
1909:35). For the Shoshone and Paiute, this was a
favorite stomachache or cramp remedy, especially for
young children. They would boil or steep whole plant,
the roots, or the leafy stems in a tea. The same
decoction was used for diarrhea and children’s colds.
Roots were eaten raw, roasted, or boiled for
indigestion. Mashed leaves were applied externally for
sores, and boiled as a wash for sores, rashes, and itches
(Train et al. 1941:61).
Leaves used for food by the Goshute (Chamberlin
1964:373) and Western Shoshone (Fowler 1986:71).
showy rushpink
1987:212
A horse medicine for the Goshute (Chamberlin
1964:374).
groundsel
1987:222-227
No species was specified, only that Senecio was used
by the Goshute as chewing gum (Chamberlin
1964:344, 381), and by the Ute as medicine
(Chamberlin 1909:36).
146
Family, Species
S. fremontii
S. hydrophilus
S. integerrimus
S. mulilopatus
S. serra var serra
S. streptanthifolius
S. triangularis
S. werneriifolius
Asteraceae
Solidago nana
Asteraceae
Tetradymia canescens
Asteraceae
Wyethia amplexicaulis
Common Name,
Source
Documented Ethnographic Use
dwarf goldenrod
1987:228
common
horsebrush
1987:233
mule ears
1987:238
Seeds eaten (Yanovsky 1936:63).
Boraginaceae
Amsinckia tessellata
Brassicaceae
Descurainia pinnata var.
filipes
bristly fiddleneck
1987:61
tansy mustard
1987:260
Brassicaceae
Descurainia
richardsonii var. sonnei
gray tansymustard
1987:261
Brassicaceae
Draba nemorosa
Brassicaceae
Lepidium lasiocarpum
var. lasiocarpum
Caryophyllaceae
Arenaria congesta var.
congesta
mustard
1987:266
hairy pod
pepperwort
1987:272
ballhead
sandwort
1987:102
Caryophyllaceae
Silene acaulis
moss campion
1987:110
When soaked or boiled dried, the plant used as a
physic. A boiled solution was also taken for venereal
diseases by the Shoshone (Train et al. 1941:96).
Seeds gathered for food by the Goshute, who also
applied the roots externally as a remedy for bruises and
swollen limbs. They also made a root tea and found the
leaves edible (Chamberlin 1964:341, 349, 384). The
Shoshone and Paiute would grind the roots, soak them
in water, and drink it as an emetic. A pulped root
poultice was used for swellings. This plant was also
used like Purshia tridentata as a wash for measles
(Train et al. 1941:99).
Seeds used for food by the Goshute (Chamberlin
1964:361).
Seeds used for food by the Utah Southern Paiute
(Fowler 1986:72). Greens were boiled and eaten, seeds
ground into a meal for mush or made into soup, by the
Southern Paiute (Rainey and Adams 2004). Used as
medicine by the Ute (Chamberlin 1909:36).
Seeds used as an in-season food or stored for winter, or
made into a mush, by the Goshute (Chamberlin
1964:340, 382). Seeds used for food by the Utah
Southern Paiute Fowler 1986:72).
Seeds used for food by the Northern Ute (Fowler
1986:72).
Seeds used for food by the Utah Southern Paiute
(Fowler 1986:72).
Used as bowel medicine for the Goshute (Chamberlin
1964:362) A steeped leaf poultice used for swellings,
and when coupled with blossoms for sun exposure and
gonorrheal ulcers by the Shoshone (Murphey 1990:42,
47). Infusion of flowers and seeds for a blood purifier
by the Shoshone (Nickerson 1966:47).
Used for colic in children by Goshute (Chamberlin
1964:381).
147
Family, Species
Caryophyllaceae
Silene douglasii
(multicaulis)
Caryophyllaceae
Silene menziesii
Cleomaceae
Cleome pinnata
(Stanleya pinnata)
Common Name,
Source
Douglas’s
catchfly
1987:110
Menzie’s catchfly
1987:111
prince’s plume
1987:284
Cleomaceae
Cleome serrulata var.
serrulata
Cyperaceae
Scirpus acutus
rocky mountain
beeplant
1987:97
hardstem bulrush
1987:682
Cyperaceae
Scirpus maritimus
alkali bulrush
saltmarsh bulrush
1987:683
Cyperaceae
Scirpus validus
Euphorbiaceae
Euphorbia prostrata
Fabaceae
Astragalus convallarius
var convallarius (A.
junceus)
Fabaceae
Glycyrrhiza lepidota
Fagaceae
Quercus gambelii
(undulata)
great Bulrush
1987:684
prostrate spurge
1987:303
lesser rushy milk
vetch
1987:359
Liliaceae^
Allium acuminatum
hooker’s onion
1987:802
American licorice
1987:384
gambel oak^
1987:305
Documented Ethnographic Use
Used as an emetic for stomach pain. The Goshute
would mash it and put into warm water as a drink. Also
used as a horse medicine (Chamberlin 1964:381).
Leaves smoked as a tobacco once dried and powdered
by the Goshute (Chamberlin 1964:345, 381).
Leaves, stems, and seeds used for food by the Western
Shoshone and Utah Southern Paiute (Fowler 1986:72).
The leaves and young stems were washed and boiled
before eating. Seeds were ground into mush (Yanovsky
1936:28). The Shoshone and Paiute used the root used
medicinally. A tonic tea was given for debilities after
an illness. The pulped root was used in the mouth for
cavities or toothaches, applied hot for earaches, and
rheumatic pains, and when mashed was applied
externally for throat pain and congestion (Train et al.
1941:94-95).
Leaves used for food by the Goshute and Eastern
Shoshone (Fowler 1986:72)
Seeds and roots used for food by the Goshute and Utah
Southern Paiute (Fowler 1986:73). Seed used for food
by the Southern Paiute (Rainey and Adams 2004).
Rootstocks eaten raw or used to make bread. Seeds and
young shoots were also eaten (Yanovsky 1936:10).
Seeds used for food by the Utah Southern Paiute
(Fowler 1986:73). Root and stems when tender (the
lower portions) used for food by the Goshute (Rainey
and Adams 2004).
Lower, tender portions eaten by the Ute (Chamberlin
1909:36).
Introduced from the eastern U.S.
A horse medicine for the Goshute (Chamberlin
1964:363).
Paiute noted tonic effects when eaten (Palmer
1878:653).
Fruit used for food by the Northern Ute (Fowler
1986:74). Acorns used for food in season and were not
preserved for winter by the Goshute (Chamberlin
1964:343, 379). Acorns eaten, raw, roasted, parched, or
ground and boiled for mush, by the Ute and Shoshone
(Stewart 1942:250).
Leaves used for food by the Goshute, Western
Shoshone, Eastern Shoshone, and Northern Ute
(Fowler 1986:75). Bulbs were eaten in spring and early
148
Family, Species
Common Name,
Source
Liliaceae^
Allium bisceptrum
twincrest onion
1987:802
Liliaceae^
Calochortus nuttallii
sego lily
1987:805
Liliaceae^
Fritillaria atropurpurea
spotted fritillaria
1987:807
Liliaceae^
Fritillaria pudica
yellow fritillaria
1987:807
Liliaceae^
Smilacina racemosa (S.
amplexicaulis)
Liliaceae^
Smilacina stellata
false spikenard
1987:809
Liliaceae^
Veratrum californicum
skunk cabbage,
California false
hellebore
1987:810
false Solomon’s
seal 1987:809
Documented Ethnographic Use
summer by the Goshute (Chamberlin 1964:360). Bulbs
and leaves eaten by the Ute (Chamberlin 1909:32).
Bulb of wild onions eaten in season but not saved for
winter by the Goshute (Chamberlain 1964:339, 360).
Bulbs and leaves eaten for food by Northern Ute and
Ute (Fowler 1986:75; Chamberlin 1909:32).
Bulbs gathered for food, eaten in season, eaten raw, or
dried for winter. When dried they were cooked with
meat in the form of stews by the Goshute (Chamberlain
1964:339, 364).
Bulbs eaten for food by the Western Shoshone, Eastern
Shoshone, Utah Southern Paiute, and Ute (Fowler
1986:75; Chamberlin 1909:33).
Bulbs eaten for food by the Utah Southern Paiute
(Fowler 1986:75). Ute used the bulbs and roots used as
a medicinal decoction but used sparingly because large
quantities were poisonous (Chamberlin 1909:34).
Less important than the sego lily, but the bulb was
edible (Chamberlin 1964 339, 370). Bulb used for food
by the Eastern Shoshone, Northern Ute, and Ute
(Fowler 1986:75; Chamberlin 1909:34).
Goshute considered this a “bear food plant”
(Chamberlin 1964:382).
Goshute would pound the roots and rub on limbs for
rheumatism. Bears ate the berries (Chamberlin
1964:382). The Shoshone and Goshute had several uses
for this plant. A fresh root poultice used for boils,
sprains, swellings. Pulverized root powder for bleeding
wounds. Liquid from soaked mashed roots as wash for
eye inflammations. The same liquid used was as an
antiseptic for blood poisoning. The pulped root eased
earaches. A boiled root tea used for menstrual
disorders, venereal diseases, and stomach trouble, and
in high concentrations used as a tonic. Exudate from
plant eaten as candy or used as cough syrup (Train et
al. 1941:92-93).
The Shoshone and Paiute used this plant as a
contraceptive. The boiled root, drunk three times a day,
for three weeks, insured sterility. Was taken by both
sexes. A root decoction also taken for venereal
diseases. The raw root was chewed for sore throats,
inflamed tonsils, and heavy colds. The mashed root
was applied externally for swellings, sore throat,
enlarged neck glands, rheumatism, boils, sores, cuts,
sore nipples, infections, and blood poisoning. The
pulped substance served as a liniment for snake bites.
149
Family, Species
Common Name,
Source
Liliaceae^
Zigadenus paniculatus
death camas
1987:811
Liliaceae^
Zigadenus venenosus
elegant death
camas
1987:811
Malvaceae
Sphaeralcea coccinea
(Malvastrum coccineum)
globemallow
1987:422
Pinaceae
Abies lasiocarpa
Poaceae
Bromus carinatus
Poaceae
Cinna latifolia
(Agrostis latifolia)
Poaceae
Festuca octoflora
Poaceae
Hordeum jubatum
Poaceae
Panicum crus-galli
(Echinochloa crus-gali)
Poaceae
Phragmites australis
(Phragmites communis)
subalpine fir
1987:29
mountain brome
1987:707
drooping
woodreed
1987:715
six-week fescue
1987:735
foxtail barley
1987:741
barnyard grass
cockspur grass
1987:720
common reed
1987:759
Poaceae
Puccinellia nuttalliana
Poaceae
Sporobolus cryptandrus
alkali grass
1987:771
prairie grass
sand dropseed
1987:780
Poaceae
Trisetum spicatum
bristle grass
1987:787
Documented Ethnographic Use
Th dried root powder was sprinkled on sores (Train et
al. 1941:98-99).
A crushed bulb poultice or wet dressing was applied by
the Shoshone and Paiute for rheumatism, sprains,
lameness, neuralgia, toothache, or swelling. Bulbs were
sometimes roasted, crushed, and applied as a hot
poultice. Boiled bulbs were used as an emetic tea. The
plant, though, is poisonous (Train et al. 1941:99-100).
Used much like Zigadenus paniculatus by the Paiute. A
crushed bulb poultice or wet dressing was used for
burns, rattlesnake bites, rheumatic pains, and swellings.
Plant is poisonous (Train et al. 1941:100).
The Goshute would pound the plant in water into a
gummy paste, then apply the paste over the rough inner
surfaces of earthen dishes. Also, sometimes was used
on wicker vessels after they had been pitched with pine
gum (Chamberlin 1964:374).
Infusion of needles or of resin blisters taken for colds
by the Shoshone (Murphey 1990:37).
Seeds used for pinole (Yanovsky 1936:7).
Seeds used for food (Yanovsky 1936:7).
Seeds used for food (Yanovsky 1936:8).
Seeds used for food by the Goshute (Fowler 1986:76).
Bread or mush made from flour of ground seeds
(Yanovsky 1936:7).
Honey dew formed on leaves by aphides used as a
sugar by the Goshute (Chamberlin 1964:376).
Sweet gum from stems used for food (Yanovsky
1936:8). Paiute considered this plant as a source of
sugar that they collected in fall. The sugary exudate
was eaten as a candy, but also used to loosen phlegm of
pneumonia patients or to sooth lung pains (Train et al.
1941:77)
Seeds used for food by the Goshute (Yanovsky 1936:9
– noted in Chamberlin 1964:370 as Glyceria distans).
Seeds parched, ground, mixed with water or milk, or
made into a mush or biscuit (Yanovsky 1936:9).
Seeds used as food by the Utah Southern Paiute
(Fowler 1986:77).
Seeds used for food by the Goshute (Yanovsky 1936:9
–noted in Chamberlin 1964:383 as Trisetum
subspicatum).
150
Family, Species
Polygonaceae
Eriogonum microthecum
Common Name,
Source
wild buckwheat
1987:482
Polygonaceae
Rumex salicifolus
(R. mexicanus)
Mexican dock
1987:492
Polygonaceae
Rumex venosus
veined dock
1987:493
Ranunculus
Aquilegia coerulea
Colorado
columbine
2008:613
Ranunculus
Ranunculus aquatilis
water crowfoot,
thread waterbuttercup
2008:623
Fendler’s
meadowrue
2008:627
black hawthorn
1987:524
Ranunculus
Thalictrum fendleri
Rosaceae
Crataegus douglasii var.
rivularis
Rosaceae
Geum macrophyllum
Rosaceae
Petrophytum
caespitosum (Spiraea
caespitosum)
Rosaceae
Potentilla glandulosa
var. intermedia
Rosaceae
Rosa nutkana
Documented Ethnographic Use
The Shoshone and Paiute would make a tea of boiled
roots, and sometimes plant tops, for tuberculosis. The
dried roots and tops were boiled and drunk for
tubercular cough. The liquid keeps and was stored in
quantity. Tea of boiled stems and leaves for bladder
trouble. Boiled whole plant for a wash or hot compress
for lameness or rheumatism (Train et al. 1941:47-48).
A remedy referred to as “blood medicine.” A root
decoction was used as an injection in the rectum for
severe constipation by the Goshute (Chamberlin
1964:380).
Root considered in standard treatments for burns,
wounds, sores, and swellings by the Shoshone and
Paiute. The roots were also dried, pulverized, and
applied as a powder, or mashed and applied as a wet
dressing or poultice. Root sometimes boiled and
applied as an antiseptic wash for drying sores, such as
from syphilis. A boiled root tea was used for venereal
diseases, as a blood purifier, for rheumatism,
pneumonia, influenza, coughs, colds, kidney disorders,
inflamed gall bladder, stomach aches and troubles, and
diarrhea (Train et al. 1941:88).
The Goshute said this was a medicinal plant for the
heart. Seeds were chewed as medicine. A tea was made
from the roots for abdominal pains and when one was
“sick all over” (Chamberlin 1964:362).
The Goshute would sometimes eat the entire plant after
boiling the plant to remove any acridity (Chamberlin
1964:379).
Shoshone believed that a weak tea of the roots, taken
over a long time, would cure gonorrhea. (Train et al.
1941:07).
Fruits eaten (Yanovsky 1936:31).
big leaf avens
1987:526
rock spiraea
tufted rockmat
1987:529
“Decoction from roots used as medicine” by the
Goshute (Chamberlin 1964:370).
Roots boiled and reduced to a pulpy mass were applied
to burns. The leaves were used as bowel medicine by
the Goshute (Chamberlin 1964:349-350, 382-383).
cinquefoil
1987:533
Roots used medicinally in a poultice applied to
swellings. Also used internally, by the Goshute
(Chamberlin 1964 378).
According to the Goshute, this plant is poisonous
(Chamberlin 1964:379).
nootka rose
1987:540
151
Family, Species
Rosaceae Rubus
leucodermis
Rosaceae Rubus
parviflorus
(R. nutkanus)
Salicaeae Salix
amygdaloides
Salicaeae Salix exigua
(Salix interior Rowlee)
Sapindaceae Acer
negundo, (Negundo
aceroides)
Solanaceae Physalis
longifolia
Solanaceae Solanum
jamesii-type*
Solanaceae Solanum
tuberosum
Typhaceae Typha
angustifolia var.
domingensis
Common Name,
Source
blackcap
raspberry
1987:542
Documented Ethnographic Use
Berries were eaten in season by the Goshute, but not
preserved (Chamberlin 1964:344, 380).
Fruits used as food by the Western Shoshone (Fowler
1986:78). Stems pounded into a powder and used as a
dressing for cuts and wounds by the Shoshone (Train et
al. 1941:87).
Berries used for food by the Goshute (Chamberlin
1964:380).
salmonberry;
thimbleberry
1987:542
peach-leaf willow Wood used for baskets and fishing weirs by the
1987:551
Goshute (Chamberlin 1964:380), and Ute (Chamberlin
1909:36).
coyote willow;
It is unclear if Indian groups distinguished between the
sandbar willow
different species. The Paiute and Shoshone used this
1987:553
plant for various, non-uniform remedies. A sitz bath of
twigs or a boiled root and bark tea was for venereal
disease. Gonorrhea was treated with a potion made of
ashes from burnt stems mixed with water. The
powdered, dried root was used to dry sores. A root
decoction was used as a blood purifier or for stomach
aches. The roots were burned to charcoal, powdered,
and made into a pill for dysentery, intestinal influenza,
failure to urinate, diarrhea. Boiled bark was used as a
spring tonic. The young twigs steeped in water with
salt used as a laxative, and the boiled woody stems for
a physic. Powdered, dried bark was used as a healing
agent for navels of young babies. A mashed root
poultice for toothaches. A solution of boiled leaves and
young twigs was rubbed onto scalp to deter dandruff.
(Train et al. 1941:89-90).
The Ute used this tree in basketry (Chamberlin
1909:360).
ash-leaf maple,
Sugar made from tree sap (Yanovsky 1936:41).
box elder
1987:42
ground cherry
Fruit consumed raw, boiled, made into a sauce, or
1987:608
cooked (Yanovsky 1936: 56); or made into preserves
(Krochmal et al. 1954: 14). A common food source in
the Great Basin (Cummings 2004: 208-216).
wild potato
No data yet (not found in Utah valley).
1987:609
wild potato
Spring beauty-bulbs were eaten by the Goshute. This
1987:610
plant was also cultivated by Goshute (Chamberlin
1964:382).
lesser bulrush;
Used like Typha latifolia (Yanovsky 1936:6).
narrowleaf cattail
1987:823
152
Appendix D
These are not all the plants that were identified in the archaeological record or in the
ethnobotanical research, but those that I was able to collect. I also included frequent associations,
meaning plants or plant communities a particular species is frequently found among or with. For
plant descriptions, see Welsh et al. 2008, plant identification manuals, such as Woody Plants of
Utah (Van Buren et al. 2011) or Wasatch Wildflowers: A Field Guide (Hegji 2010), and
plants.usda.gov.
Two notes on the ethnographic sources: Ebeling (1986:73) often referenced the Diggers,
which was a name given to bands of Ute, Paiute, Shoshone, and Goshute. Plant use in Yanovsky
(1936) was only included if the plant was used by indigenous groups in Utah.
ABIES CONCOLOR
Family: Pinaceae
Genus: Abies
Species: concolor
Common Name: white fir (Welsh et al. 2008:18).
Alternate Names: Piceae concolor, Pinus concolor (Welsh et al. 2008:18).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES28 Western hardwoods FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES37 Mountain meadow
(Zouhara 2001a).
Production Time:
Seeds are shed from late September to early October. Good crops occur every two to five
years. Seeds germinate in spring (USDA NRCS Plant Materials Program 2002). About
185 to 295 seeds are produced per cone (Zouhara 2001a).
Frequent Associations:
Can be found in stands with other white firs, or alone amid other trees such as oak and
maple (Welsh et al. 2008:18). Grows where precipitation exceeds 20 inches annually and
can grow on soils from almost any parent material (USDA NRCS Plant Materials
Program 2002).
Archaeological Artifacts:
Abies pollen was found on groundstone from Wolf Village (Dahle 2011) and in fill from
Smoking Pipe (Scott 1984).
153
Ethnographic Use:
This plant was mainly used by the Shoshone and Paiute for several uses. The bark was
boiled and drunk in a tea, or the soft resin of the bark was eaten, as a cure for
tuberculosis. The pitch was warmed and made into a poultice for sores and boils. Fresh
pitch was applied to cuts. A tea of needles and sometimes resin was used for pulmonary
troubles (Train et al. 1941:19).
Collected:
I collected needles of this plant from Red Butte Gardens during the summer of 2015. The
needles required three digestions. This sample was sonicated.
Microfossiles Production:
Evett et al. (2006:356) found no diagnostic morphotypes in Abies concolor, with leaf
silica percentage measuring 0.1%. I also found no diagnostic morphotypes in the needles
I digested. There was some residual organic matter, but it is doubtful such residue would
survive in soils later to be found in archaeological contexts.
Elongate polyhedrons have been found in the needles of Abies balsamea (Bozarth
1993:100), and polyhedrons, silicified transfusion cells, and tracheids been found in the
wood and needles of Abies alba (Carnelli et al. 2003). Abies grandis has been found to
produce common yet poorly silicified blocky mesophyll and epidermal cells in the
needles, and sporadic tracheids in the twigs (McCune 2014:105). No phytoliths have been
found in Abies amabilis, and tracheids, blocky types, and epidermal polygonals have
been found in Abies lasiocarpa (Blinnikov 2005:78).
ACHILLEA MILLEFOLIUM
Family: Asteraceae
Genus: Achillea
Species: millefolium (Welsh et al. 2008:145).
Common Name: common yarrow, filfoil yarrow
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
FRES41 Wet grasslands
FRES44 Alpine
(Aleksoff 1999).
Production Time:
Yarrow has a long flowering season in Utah. At higher elevations the flowers bloom from
June to July, and seeds are ripe and disseminated from September to October, at which
point the plant has dried out. In lower elevations, flowers bloom as early as April until
July, seeds can ripen as early as August and will disseminate in September. By October
the plant has dried (Aleksoff 1999). The yarrow I grew in a pot stayed in bloom until
October.
Archaeological Artifacts:
154
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016), on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Goshute would make a tea of yarrow for biliousness and headaches. Externally they
applied it for rheumatism or bruises (Chamberlin 1964:350, 360). The Ute also externally
applied yarrow on bruises and used yarrow as a tea for sickness (Chamberlin 1909:32).
The Paiutes would make yarrow into a tea for stomach problems (Palmer 1878:651). The
Paiute and Shoshone would make a poultice of the boiled plant that they applied to pains,
sores, or swellings, and a poultice of fresh roots used to deaden pain, or as an anesthetic
to painful wounds. A decoction of flowers was brewed for stomachaches, itching, or
indigestion, or as a liniment for muscle pains. A decoction of leaves was taken for colic
or dyspepsia. A decoction of roots was used as a preliminary soak to extract splinters. A
decoction of the plant was taken for diarrhea, or upset stomach, or for colds; and a
decoction of leaves, or smelling of the crushed green plant for headaches or toothaches
(Train et al. 1941:20). The Shoshone also made a poultice of the whole plant for abscess
or boils (Murphey 1990:43).
Collected:
Samples from this plant came from Spanish Fork Canyon and from a specimen I grew
myself. I collected and digested the flower tops and leaves. This sample was sonicated.
Microfossil Production:
Yarrow is considered a common producer of phytoliths (Morris et al. 2009:342). The
leaves produce the majority of phytoliths while the stems are not highly silicified. In the
leaves are found hairs, such as acicular psilate unsegmented hairs, as well as polyhedral
epidermal sheets and anticlinal epidermal sheets, some of which have striations, and
lastly tracheids (Morris 2008: Plate Ia and Plate Ib; Morris et al. 2009:342). Blocky forms
have also been found in Yarrow (Blinnikov et al. 2013:108). The following table notes
my findings, with PI referring to the production index (see chapter 4).
Species
Achillea
millefolium
Plant Tissue
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Inflorescence
Inflorescence
Phytolith
Sinuate epidermal with striate texture, E1a
Lancelote hair with psilate texture, unsegmented, H1b
Irregular sub-spheroid form with ruminate texture, S1b
Tracheid, V1
Elongate with pilate margins, Achillea type, L1a
Papillate with liguate margins, tuberculated, P1a
Irregular sub-spheroid form with ruminate texture, S1b
Tracheid, V1
AMELANCHIER ALNIFOLIA
Family: Rosaceae
Genus: Amelanchier
Species: alnifolia
Common Name: juneberry, dwarf shadbush, saskatoon (Welsh et al. 2008:632).
Forest-Range Environmental Study Ecosystems:
FRES10 White-red-jack pine
FRES11 Spruce-fir
155
PI
C
C
C
C
C
U
U
U
Figures
4.4.F
4.3.C
4.9.A
4.7.A
4.10.F
4.8.J
4.9.A
4.7.A
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES24 Hemlock-Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
(Fryer 1997).
Production Time:
Fragrant white flowers bloom from April to May and are followed in late summer by
purple fruits that resemble blueberries (Andersen and Holmgren 1996; Selland 2003).
Frequent Associations:
Commonly found in riparian areas (Fryer 1997). Flowers and fruits “are borne in terminal
clusters” (Fryer 1997).
Archaeological Artifacts:
Service berry seeds were identified in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
For the Goshute, the berries were an important food source that they collected and ate in
season, boiled, or raw. They also preserved the berries, as mashed and/or dried, for later
(Chamberlin 1964:343, 361; Rainey and Adams 2004). The Northern Ute and Utah
Southern Paiute ate the fruit (Fowler 1986:78; Rainey and Adams 2004). The berries
were important for the Ute as well; they were used in season or dried for winter
(Chamberlin 1909:32). A tea would also sometimes be made from dried leaves
(Yanovsky 1936:30).
Collected:
Plant material was collected from the Sego Lily Gardens and Central Utah Gardens. I
digested the berries of Juneberry after removing the seeds. This sample was not
sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Amelanchier
alnifolia does produce abundant starch grains that range in size from 2.8-8.4 um (mean
3.5 um), with an open central hilum, a small red dot, and a visible cross-polarization
(Zarrillo and Kooyman 2006:488). Other Amelanchier species, such as A. arborea, do
not produce phytoliths (Tedford 2009:191). The following table notes my findings, with
PI referring to the production index (see chapter 4).
Species
Plant Tissue
Amelanchier Berry
alnifolia
Berry
Berry
Phytolith
Styloid cluster, O2a
Crystal sand, O3
Lancelote hair with psilate texture, segmented, H1c
AMELANCHIER UTAHENSIS
Family: Rosaceae
Genus: Amelanchier
Species: utahensis
156
PI
U
U
U
Figure
4.1.R
4.2.A
4.3.D
Common Name: Utah serviceberry (Welsh et al. 2008:632).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
(Zlatnik 1999a).
Production Time:
Leafs and blooms earlier than other species (Zlatnik 1999a). Is very similar to the
Saskatoon serviceberry.
Frequent Associations:
“Utah service berry hybridizes with Saskatoon serviceberry” and is often found
associated with Gambel oak or mountain mahogany (Zlatnik 1999a).
Archaeological Artifacts:
Service berry seeds were identified in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
The Utah Southern Paiute ate the fruit for food (Fowler 1986:78). The Shoshone and
Paiute would boil the green, inner bark and drink it for snow blindness (Train et al.
1941:21).
Collected:
Berries and wood were collected from Central Utah Gardens. Both required two
digestions. These samples were sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Other
Amelanchier species, such as A. arborea, do not produce phytoliths (Tedford 2009:191).
The following table notes my findings, with PI referring to the production index (see
chapter 4).
Species
Amelanchier
utahensis
Plant
Tissue
Berry
Berry
Berry
Berry
Wood
Wood
Phytolith
PI Figures
Small raphide-types connected, different orientations, O1c
Styloid, single, O2
Single, rectangular, prismatic O5
Tracheid, V1
Tracheid, V1
Indeterminate vascular tissue V2
U
U
U
U
U
C
ARCTOSTAPHYLOS PATULA
Family: Ericaceae
Genus: Arctostaphylos
Species: patula
Common Name: greenleaf manzanita (Welsh et al. 2008:356).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES27 Redwood
FRES28 Western hardwoods
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
(Hauser 2007).
157
4.1.G
4.1.L
4.2.K
4.7.B
4.7.B
4.8.A
Production Time:
Large seed crops are produced almost yearly. Flowers bloom from April to June (Hauser
2007).
Archaeological Artifacts
Seeds were found in the fill from South Temple (Cummings et al. 2004).
Ethnographic Use:
Leaves were boiled and drunk for venereal disease by the Shoshone (Train et al.
1941:24).
Collected:
Leaves were collected from Grow Wild Nursery, Salt Lake City. This sample was
sonicated.
Microfossil Production:
Evett et al. (2006:356) found no diagnostic morphotypes in Arctostaphylos patula, with
leaf silica percentage measuring less than 0.1%. Very few phytoliths have been found in
A. pungens leaves, such as ovoid forms with radiation striations, polygonal epidermals,
blocky crystalline, and tracheid shapes (McNamee 2013:36, 48, 63). A. uva-ursi also
produces phytoliths, including polyhedrals, jigsaw epidermal cells, and trichomes in the
leaves (Carnelli et al 2003). I found no phytoliths in the leaves of this sample.
ARTEMISIA BIENNIS
Family: Asteraceae, Compositae
Genus: Artemisia
Species: biennis
Common Name: biennial wormwood (Welsh et al. 2008:158).
Forest-Range Environmental Study Ecosystems:
Grows in most plant communities, especially those that have been disturbed or are of a
wetland-riparian nature (CalFlora 2005).
Production Time:
Blooms in the fall (CalFlora 2005).
Archaeological Artifacts:
Artemisia pollen was found on groundstone from Wolf Village (Cummings 2011), on
groundstone and in fill from the Hinckley Mounds (Peterson 2016), and in fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
The Goshute extensively gathered the seeds of biennis and used them for food
(Chamberlin 1964:362).
Collected:
Seeds were collected from the Brigham Young University Monte L. Bean Herbarium.
This sample was not sonicated.
Microfossil Production:
I found no scholarly sources on microfossil production in this species. Other Artemisia
species do produce phytoliths. The following table notes my findings, with PI referring to
the production index (see chapter 4).
158
Species
Plant Tissue
Artemisia Leafy seeds
biennis* Leafy seeds
Leafy seeds
Leafy seeds
Leafy seeds
Phytolith
PI Figures
Sinuate epidermal with striate texture, E1a C 4.4.G
Astrosclerid, C1
U 4.3.A
Ligulate epidermal, psilate texture, E3 U none
Spheroids with ruminate texture, S3b
U 4.9.bb
Tracheid, V1
U none
ARTEMISIA DRACUNCULUS
Family: Asteraceae, Compositae
Genus: Artemisia
Species: dracunculus
Common Name: tarragon (Welsh et al. 2008:159).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES29 Sagebrush
FRES30 Desert shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
(Groen 2005).
Production Time:
Infrequent seed production. Flowers from July to October (Groen 2005).
Frequent Associations:
Often associated with quaking aspen, firs and spruces, rabbitbrush, sagebrush, and
wildrye (Groen 2005).
Archaeological Artifacts:
Artemisia pollen was found on groundstone from Wolf Village (Cummings 2011), on
groundstone and in fill from the Hinckley Mounds (Peterson 2016), and in fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
The oily and nutritious seeds of Artemisia dracunculoides, another Tarragon, were a
favorite dish for the Goshute and were extensively gathered (Chamberlin 1964:363).
The Southern Paiute would make a tea for colds, headaches, worms, and as a stimulant
for women in childbirth. They used the leaves to stop nosebleeds, and the seeds were
ground into mush (Rainey and Adams 2004). The Shoshone would make a hot decoction
of branches as a physic for colds. Steam from boiling the plant was used for eye trouble.
A decoction of the whole plant was a wash for nettle stings and for venereal diseases. A
poultice of pulped, green plant was applied externally for sore throat or neck glands
(Train et al. 1941:25-26). They also steeped the seeds added to dishes to provide flavor
(Murphey 1990:29).
Collected:
Leaves and plant tops were collected from the Red Butte Gardens, Salt Lake City. This
sample was sonicated.
Microfossil Production:
Abundant blocky-form phytoliths in the leaves, minimal amounts in the branches, and
none present in the seeds (McNamee 2013:34, 45). The following table notes my
findings, with PI referring to the production index (see chapter 4).
159
Species
Plant
Tissue
Artemisia
Leaves
dracunculus Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
New growth
Phytolith
PI Figures
Small raphide-types connected, different orientations, O1c
Crystal sand, O3
Sinuate epidermal with heavy to light striate texture E1b
Tracheid V1
Ligulate epidermal with psilate exture, E3
Crenate epidermal, E6
Acicular hair, with striate texture, unsegmented, H2a
Irregular sub-spheroid form with ruminate texture, S1b
Blocky with psilate to facetate texture, S2
Stomates, V3
Sinuate epidermal with heavy to light striate texture E1b
U
R
C
C
U
U
U
U
U
U
U
4.1.H
none
4.4.J
4.7.C
none
4.5.M
4.3.I
4.9.B
4.9.Q
4.6.C
4.4.J
ARTEMISIA LUDOVICIANA
Family: Asteraceae, Compositae
Genus: Artemisia
Species: ludoviciana
Common Name: Louisiana wormwood (Welsh et al. 2008:160).
Variations: Five variations are present in Utah, with variations incompta, latiloba, and
ludoviciana present in Utah Valley.
Forest-Range Environmental Study Ecosystems:
FRES14 Oak-pine
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES44 Alpine
(Anderson 2005)
Production Time:
Small green flowers bloom from late July to early September (Selland 2003). Seeds
mature and disseminate from October to December (Anderson 2005).
Frequent Associations:
Includes other sagebrush, grasses, and junipers (Anderson 2005).
Archaeological Artifacts:
Artemisia pollen was found on groundstone from Wolf Village (Cummings 2011), on
groundstone and in fill from the Hinckley Mounds (Peterson 2016), and in fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
The Utah Southern Paiute used the seeds for food (Fowler 1986:71). The Paiute saw
medicinal virtue in this plant and made it into a tea or stuffed it into their nostrils (Palmer
1878:652). The Southern Paiute would make a tea for colds, worms, and headaches, and
as a stimulant for women in childbirth. The leaves were used to stop nosebleeds and
seeds were ground into mush (Rainey and Adams 2004). The Paiute and the Shoshone
160
would make a decoction of leaves for headaches, colds, and coughs, and a decoction of
branches for colds, coughs, influenza. A decoction of tops, sometimes with roots, for
colds, diarrhea, coughs, and severe infection. A decoction of the whole plant, or
sometimes shoots, as a physic for heavy colds, coughs, fevers, and stomach aches, or as a
wash for rashes, itching, or skin eruptions. The steeped leaves would be made into a
poultice used to compress fevers, especially for babies. An infusion of leaves was used to
regulate menstrual disorders or as an eyewash. (Train et al. 1941:26-27)
Collected:
Leaves and flower tops were collected from the Sego Lily Gardens, Sandy City. This
sample was sonicated.
Microfossil Production:
Has been found to produce anticlinal epidermal sheets, sometimes with striations, as well
as tracheids (Morris 2008:131). Polyhedrals and jigsaw-puzzle pieces have also been
found, as well as branched tracheary elements (Bozarth 1992:194). The following table
notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Artemisia
Leaves
ludoviciana Leaves
Leaves
Leaves
Leaves
Inflorescence
Phytolith
Crystal sand, O3
Sinuate epidermal with psilate texture, E1
Blocky with psilate to granulate texture, S2a
Tracheid, V1
Parenchyma, Y1
Tracheid, V1
PI
U/R
C
C
C
U
C
Figures
none
4.4.A
4.9.S
4.7.D
4.6.E
4.7.D
ARTEMISIA TRIDENTATA
Family: Asteraceae, Compositae
Genus: Artemisia
Species: tridentata
Common Name: big dagebrush, common sagebrush (Welsh et al. 2008:162).
Variations: Four variations are found in Utah, with variations tridentata and vaseyana both
present in Utah Valley.
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands
(Tirmenstein 1999b).
Production Time:
Small yellow blossoms bloom in late summer to fall (Anderson and Holmgren 1996;
Selland 2003). Plants need to be at least two years old to produce seed, and seeds are
produced from October to December (Tirmenstein 1999b).
Frequent Associations:
Often hybridizes with its subspecies. Often grows alongside cheatgrass and other grasses,
broom snakeweed, and rabbitbrush (Tirmenstein 1999b).
Archaeological Artifacts:
161
Various fragments of tridentata were found in the fill of Smoking Pipe (Billat 1985:91).
Additionally, Artemisia pollen was found on groundstone from Wolf Village (Cummings
2011), on groundstone and in fill from the Hinckley Mounds (Peterson 2016), and in fill
from Smoking Pipe (Scott 1984).
Ethnographic Use:
The Goshute would make a tea of leaves that they used as medicine in feverish
conditions. They also used the leaves to cover food preserved in caches (Chamberlin
1964:351, 363). The Ute also used the leaves medicinally (Chamberlin 1909:32). Great
Basin peoples in general used the seeds (Sutton 1989:245-246). Specifically, the Utah
Southern Paiute and Western Shoshone (Fowler 1986:71). The Paiutes would make a tea
for colds, headaches, and worms, or as a stimulant (Palmer 1878:651; Rainey and Adams
2004). The Shoshone would chew the leaves for indigestion (Murphey 1990:45). They
would also make a decoction of branches for stomach cramps, red ant bites, and take with
salt for coughs, or used as a wash for lumbago or muscular cramps. A hot poultice of
branches was made for aches and pains, and rheumatism. A headache wash was made of
a decoction of leaves, and a poultice of crushed leaves for headaches. Steeped leaves
were made into a poultice for inflamed eyes or sores, cuts, and wounds (powdered
branches were also used). A decoction of leaves was taken to cause sweating, to break a
fever, or as an antiseptic wash for cuts and as an antiseptic bath for newborns, wounds,
sores, or taken with salt for pneumonia coughs. A decoction of leaves, or leaves chewed
for a poison antidote or for colds. Raw leaves were chewed for indigestion. A poultice of
mashed leaves for toothaches. A decoction of plant tops for colds, and an over dosage
serves as an emetic. A decoction of plant taken as a tonic after childbirth (Train et al.
1941:44-47).
Collected:
Leaves, flower tops, and twigs were collected from Rock Canyon Park, Provo. This
sample was partially sonicated.
Microfossil Production:
The leaves have been found to produce a few blocky shapes, and the stems produce
several small blocky forms, usually less than 20mu (Morris 2008:145). Leaf phytoliths
were minimal, and stem phytoliths were common (Morris 2008:168, 178). Produces
tracheids, as well as smooth (psilate) or cavate blocky cubic or globose shapes and
smooth epidermal polygonal irregular shapes (Blinnikov 2005:78-79). The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Artemisia Leaves
tridentata Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Inflorescence
Phytolith
Raphide bundle of same orientation, O1a
Raphide bundle of different orientation, O1b
Small raphide-types connected, different orientations, O1c
Crystal sanad, O3
Prismatic, single, rectangle, O5
Prismatic, hexagon, single, O5b
Sinuate epidermal with psilate texture, E1
Irregular sub-spheroid form, ruminate to facetate texture, S1b
Blocky with facetate texture, S2b
Parenchyma Y1
Irregular sub-spheroid form, ruminate to facetate texture, S1b
162
PI
U
U
U
U
U
U
U
U
U
R
C
Figures
4.1.C
4.1.E
4.1.I
4.2.B
4.2.L
4.2.R
4.4.B
4.9.E
none
4.6.F
4.9.E
Species
Plant Tissue
Inflorescence
Twigs
Phytolith
Tracheid V1
Indeterminate vascular tissue V2
PI Figures
U 4.7.E
C none
ATRIPLEX TRUNCATA
Family: Amaranthaceae, Chenopodiaceae
Genus: Atriplex
Species: truncata
Common Name: wedge orach (Welsh et al. 2008:128).
Forest-Range Environmental Study Ecosystems:
Often in saline salt-grass-greasewood-rabbitbrush communities, and in palustrine or
lacustrine habitats (eFlroas 2017). Creosote Bush Scrub, Pinyon-Juniper Woodland,
Yellow Pine Forest, Red Fir Forest, Lodgepole Forest, Subalpine Forest, or wetlandriparian zones (CalFlora 2015).
Production Time:
Blooms in the summer and fall (CalFlora 2015).
Archaeological Artifacts:
The fruit and seeds of Atriplex were found in the fill of Hinckley Mound (Puseman 2016)
and Woodard Mound (Richens 1983:111).
Ethnographic Use:
The Goshute would collect and eat the seeds (Chamberlin 1964:340, 363)
Collected:
A flower head with seeds was collected from Thanksgiving Point Gardens, Lehi. This
sample was not sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. No phytoliths
have been found in Atriplex canescens (McNamee 2013:36). The following table notes
my findings, with PI referring to the production index (see chapter 4).
Species Plant Tissue
Atriplex Inflorescence
truncata Inflorescence
Inflorescence
Phytolith
Blocky with psilate to facetate texture, S2
Blocky with facetate texture, S2b
Tracheid, V1
BALSAMORHIZA SAGITTATA
Family: Asteraceae, Compositae
Genus: Balsamorhiza
Species: sagittata
Common Name: arrowleaf balsamroot (Welsh et al. 2008:172).
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
163
PI
U
U
C
Figures
none
4.9.U
none
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES44 Alpine
(McWilliams 2002).
Production Time:
Shoots sprout in April or May from a large taproot, and in summer the plant goes
dormant. Balsamroots need to be at least five years old before they produce flowers
(Selland 2003).
Frequent Associations:
Often grows alongside Balsamorhiza macrophylla (Anderson and Holmgren 1996), and
singleleaf pinyon and curl-leaf mountain mahogany (McWilliams 2002).
Archaeological Artifacts:
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016), on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Goshute would gather the leaves and petioles in the early season and eat them as a
green or boiled in water. They gathered and ate the seeds late in year. The root was used
as remedy for fresh wounds, and was also chewed or pounded into a paste (Chamberlain
1964:338, 348, 363). The Eastern Shoshone, Northern Ute, Utah Southern Paiute would
use the seeds, roots, young leaves as food (Fowler 1986:71). The Shoshone would make a
root decoction as an eyewash, and a mashed root poultice for insect bites, and syphilitic
sores (Train et al. 1941:50-51). The Ute would eat the young shoots and leaves, and
sometimes roots (Chamberlin 1909:32-33).
Collected:
Leaves, tops, and flowers were collected from Red Butte Gardens, Salt Lake City. This
sample was sonicated.
Microfossil Production:
Segmented hairs commonly found in the leaf and flowering head. These hairs are thinly
silicified and tend to break and disfigure. They also produce tracheids (Morris 2008:132).
Siliceous hairs have been found in the roots (Witty 1962:25). The following table notes
my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Basalmhoriza Leafy tops
sagittata
Leafy tops
Phytolith
Lancelote hair with psilate texture, segmented, H1c
Irregular sub-spheroid form, ruminate to facetate
texture, S1b
Leafy tops
Blocky with facetate texture, S2b
Leafy tops
Parenchyma, Y1
Inflorescence Irregular sub-spheroid form with granulate texture, S1c
Inflorescence Tracheid, V1
CERCOCARPUS LEDIFOLIUS
Family: Rosaceae
Genus: Cercocarpus
Species: ledifolius
Common Name: curl-leaf mountain mahogany (Welsh et al. 2008:633).
164
PI Figures
C 4.3.E
U none
U
U
U
U
none
4.6.G
4.9.G
4.7.F
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES26 Lodgepole pine
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
(Gucker 2006).
Production Time:
Pale yellow or cream blossoms bloom from May to June, and seeds ripen in the fall
(Anderson and Holmgren 1996; Selland 2003). Trees are usually mature when at least ten
years old (Gucker 2006).
Frequent Associations:
Often hybridizes with other mountain mahogany species (Gucker 2006).
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in fill from Smoking Pipe (Scott 1984).
Ethnographic Use:
The Goshute would use the wood for bows, and the charcoal of the wood mixed with
water was used for burns (Chamberlin 1964:350, 365). The Shoshone would use this
plant for pulmonary disorders and tuberculosis. The bark would be dried and then boiled
for decoctions, and sometimes boiled with Purshia tridentata or Populus trichocarp. The
dried bark or steeped leaves would be made into a decoction taken for coughs and colds.
The dried bark was made into a paste and applied to sores, cuts, burns, and wounds, and
leaves and bark made into a poultice for swellings. The leaves and bark in decoction for
heart disorders. A bark decoction also used for stomachache, venereal disease, diarrhea,
stomach ulcers, pneumonia, diphtheria, and eye disease (Train et al. 1941:35-36).
Collected:
Leaves and wood were collected from Central Utah Gardens, Orem. Both required two
digestions. This sample was sonicated.
Microfossil Production:
Found to produce abundant blocky, prismatic and elongate rectangular styloid forms
(Morris 2008:144). Phytoliths were observed in minimal amounts (Morris 2008:168,
178). McNamee (2013:38) found no phytoliths in Cercocarpus spp. The following table
notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Cercocarpus Leaves
ledifolius
Wood
Leaves
Leaves
Leaves
Phytolith
Inderterminate vascular tissue V2
Raphide bundle of different orientation, O1b
Small raphide-types connected together and of different
orientations, O1c
Prismatic, rectangle, single, O5
CHRYSOTHAMNUS NAUSEOSUS
Family: Asteraceae, Compositae
Genus: Chrysothamnus
Species: nauseosus
Synonym: Ericameria nauseosa
165
PI
NP
U
U
U
Figures
U
4.2.M
4.8.B
none
4.1.F
Common Name: rubber rabbitbrush (Welsh et al. 2008:184).
Variations: Fifteen variations are found in Utah.
Forest-Range Environmental Study Ecosystems:
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands
(Tirmenstein 1999e).
Production Time:
Clusters of yellow blossoms bloom in late summer and can continue into the fall; mature
seeds follow a few days later (Anderson and Holmgren 1996; Selland 2003)
Archaeological Artifacts:
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016), on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Shoshone and the Paiute would steep the leaves into a tea that was drunk for stomach
disorders and colds. The dried leaves and flowers would be steeped as a general tonic. A
tea from the roots and tops for diarrhea. Boiled stems and leaves for a cough medicine
(Train et al. 1941:37).
Collected:
Leaves and flower tops were collected from Red Butte Gardens, Salt Lake City. This
sample was sonicated.
Microfossil Production:
Poorly silicifies and only produces a limited amount of segmented hairs (Blinnikov
2005:78, 82). C. viscidiflorus was found to produce minimal amounts of phytoliths
(Morris 2008:168). The following table notes my findings, with PI referring to the
production index (see chapter 4).
Species
Plant Tissue
Chrysothamnus Leaves
nauseous
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Inflorescence
Inflorescence
Phytolith
Irregular sub-spheroid form, granulate texture, S1c
Blocky with psilate to granulate texture, S2a
Blocky with facetate texture, S2b
Tracheid, V1
Irregular sub-spheroid form, granulate texture, S1c
Blocky with psilate to granulate texture, S2a
Blocky with facetate texture, S2b
Tracheid, V1
PI
C
C
C
C
C
C
C
C
CRATAEGUS DOUGLASII
Family: Rosaceae
Genus: Crataegus
Species: douglasii
Common Name: river hawthorn (Welsh et al. 2008:637).
Variations: Two variations exist, with variation rivularis being found in Utah Valley.
166
Figures
4.9.H
4.9.T
4.9.V
4.7.G
4.9.H
4.9.T
4.9.V
4.7.G
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral - mountain shrub FRES35 Pinyon – juniper
FRES36 Mountain grasslands
(Habeck 1991).
Production Time:
White fragrant blossoms bloom in April and May, fruits follow later in the summer
(Anderson and Holmgren 1996).
Frequent Associations:
Cottonwoods, aspens, pines, Wood’s rose, and chokecherry (Habeck 1991).
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The fruits were eaten (Yanovsky 1936:31).
Collected:
Berries were collected from Sego Lily Gardens, Sandy City. They required three
digestions. This sample was not sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. On PhytCore
(2017) there are hair bases, parenchyma, epidermal ground-mass polyhedrals, and blocky
shapes recorded for Crataegus aronia. The following table notes my findings, with PI
referring to the production index (see chapter 4).
Species
Plant Tissue Phytolith
PI Figures
Craetaegus douglasii* Berry
Blocky with psilate to facetate texture, S2 U 4.9.R
Berry
Indeterminate vasculare tissue, V2
U none
CUCURBITA
Family: Cucurbitaceae
Genus: Cucurbita
Species: maxima and moschata
Common Name: fall squash, winter squash, pumpkin, zucchini, and cushaw, butternut, winter
crookneck (Welsh et al. 2008:349).
Forest-Range Environmental Study Ecosystems:
This cultivated species prefers well-drained soils and full sun.
Production Time:
Since squashes are highly susceptible to frost, plant when soil temperatures are above
60oF. The fruits are usually ready for harvesting about seven to eight weeks after
planting.
Archaeological Artifacts:
Uncharred squash rinds were found in the fill of the Fremont Zone of Spotten Cave
(Pearce 2012).
Ethnographic Use:
The Plateau Shoshoneans would consume this plant (Lowie 1924:200).
167
Collected:
No samples were collected for this study given the prolific publication and analysis that
currently exists on Cucurbita.
Microfossil Production:
All parts of squash plants produce phytoliths, with the most useful being found in the
rinds (Piperno 2006:65). Squash rinds produce deeply scalloped spheroidal phytoliths
that is diagnostic of Cucurbita (Piperno et al. 2000).
DESCHAMPIA CESPITOSA
Family: Poaceae, Graminae
Genus: Deschampia
Species: cespitosa
Common Name: tufted hairgrass (Welsh et al. 2008:841).
Forest-Range Environmental Study Ecosystems:
FRES10 White-red-jack pine
FRES11 Spruce-fir
FRES13 Loblolly-shortleaf pine
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES24 Hemlock-Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES27 Redwood
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES41 Wet grasslands
FRES42 Annual grasslands
FRES44 Alpine
(Walsh 1995).
Production Time:
Growth begins in early spring and grass remains green through summer, and seeds tend
to mature towards the end of summer (Walsh 1995).
Archaeological Artifacts:
Poaceae phytoliths were found in teeth tartar from the Provo Mounds (Yost 2009:6), as
pollen on groundstone from Wolf Village (Cummings 2011), and Hinckley Mounds
(Peterson 2016), and as starch on groundstone from Wolf Village (Cummings 2011).
Additionally, Poaceae seeds and caryopsis were found in coprolites from Spotten Cave
(Pearce 2012), and in the fill of Hinckley Mounds (Puseman 2016).
Ethnographic Use:
The Goshute would eat the seeds (Chamberlin 1964:367).
Collected:
Florets with seeds were collected from Central Utah Gardens, Orem. This sample was not
sonicated.
Microfossil Production:
168
In the basal leaf sheath and in the culm leaf blade are papillae and elongates (Blackman
1971:771). No scutiform opals were found (Blackman 1971:778). The following table
notes my findings, with PI referring to the production index (see chapter 4).
Species
Cell Type
Deschampia
Long
cespitosa
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
PI Figures
Sinuate epidermal with psilate texture, E1 U
none
Entire epidermal with psilate texture, E4
C
4.5.A
Blocky epidermal, lateral striations, E7
U
none
Elongate with pilate margins, L1
C 4.10.A
Elongate with entire margins, L2
C 4.10.H
Elongate, aculeate margins, L5
C 4.11.E
Elongate, aculeate margins, curled, L5a
C 4.11.L
Tracheid, V1
U
4.7.H
Round/oblong rondels, G1a
C 4.12.A
Square/rectangular rondels, G1b
C 4.12.E
Keeled rondels, G1c
C 4.12.H
Pyramidal rondel, G1d
C 4.12.M
Pyramidal rondel, aculeated, G1d1
C 4.12.P
Lancelote style hair trichomes, G6a
U 4.12.Y
Hair base trichome, G6b
U
none
General, polylobe/bilobe rondel, G7b
U 4.12.jj
Parenchyma, Y1
U
4.6.H
ELYMUS CINEREUS
Family: Poaceae, Graminae
Genus: Elymus
Species: cinereus
Common Name: Great Basin wildrye (Welsh et al. 2008:846).
Synonym: Leymus cinereus
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands
(Anderson 2002a).
Production Time:
Starts to grow in early spring, flowers in the summer, seeds ripen as early as July and as
late as September (Anderson 2002a).
Archaeological Artifacts:
Elymus starch grains were found in teeth tartar from Provo Mounds (Yost 2009:6).
Ethnographic Use:
The Eastern Shoshone, Goshute, and Utah Southern Paiute ate the seeds (Fowler
1986:76).
Collected:
169
Florets with seeds were gathered from Red Butte Gardens, Salt Lake City. This sample
was sonicated.
Microfossil Production:
According to Kerns (2001:286), Elymus species are in the subfamily Pooideae and
produce crenates, rondels, and pyramids. There is reason to believe that the morphotypes
in the genus Elymus do not group together (Morris 2008:140). Elymus spp. have been
found to produce distinct articulated dendritic phytoliths. Noted to have a high research
potential (Serpa 2008:104, 108) Specifically, E. cinereus has been found to be an
abundant producer of phytoliths, dominated by long, deeply indented and long indented
cells, also oblong knobby hair bases and keeled rondels (Morris et al. 2009:341, 346).
Blinnikov (2005:84) found “smooth rods,” or rectangular long cells, in cinereus.
Blackman (1971:772) found intercostal, costal, and abaxial Hats in the basal leaf sheath
and blade, and culm leaf sheath and blade.
Thin, elongate phytoliths have been found in the roots of Elymus canadensis (Sangster
and Hodson 1992:242). Square plates, along with several other forms such tracheids, long
cells, trichomes, and rondels, have been found in E. elymoides (McNamee 2013:90, 9899). The following table notes my findings, with PI referring to the production index (see
chapter 4).
Species Cell Type
Elymus
Long
Cinereus Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
Ligulate to collumnate epidermal with psilate texture, E3a
Entire epidermal with psilate texture, E4
Elongate, dendritic margins, L3
Elongate, crenate margins, L4
Elongate, aculeate margins, L5
Elongate, aculeate margins, granulate texture, L5c
Tracheid, V1
Square/rectangular rondels, G1b
Keeled rondels, G1c
Pyramidal rondel, G1d
Lancelote style hair trichomes, G6a
Hair base trichome, G6b
Acicular hair with psilate texture, unsegmented, H2b
Acicular hair with ovoid base with tubucurlate processes, H2b
Papillae with ligulate margins, P1
Papillae with pitted edges, P1b
Ellipsoids with tuberculate processes, S5
ELYMUS GLAUCUS
Family: Poaceae, Graminae
Genus: Elymus
Species: glaucus
Common Name: blue wildrye (Welsh et al. 2008:847).
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
170
PI
U
U
A
U
C
C
U
C
C
C
U
U
U
U
C
C
U
Figures
4.4.O
4.5.b
4.10.S
4.11.A
4.11.F
4.11.O
4.7.I
4.12.F
4.12.I
4.12.Q
4.12.Z
None
4.3.J
4.3.L
4.8.H
4.8.K
4.9.dd
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
FRES42 Annual grasslands
(Johnson 1999).
Production Time:
Flowers from June to August (Johnson 1999).
Frequent Associations:
Often hybridizes with other Elymus species, and often found alongside alders, maples,
sage grass, strawberry, yarrow, asters, and other grasses (Johnson 1999).
Archaeological Artifacts:
Elymus starch grains were found in teeth tartar from Provo Mounds (Yost 2009:6).
Ethnographic Use:
The seeds were eaten (Yanovsky 1936:8).
Collected:
Florets with seeds were gathered from Red Butte Gardens, Salt Lake City. This sample
was sonicated.
Microfossil Production:
According to Kerns (2001:286), Elymus species are in the subfamily Pooideae and
produce crenates, rondels, and pyramids. McCune (2014:100) found rondels (10um),
papillae, and dendriform elongate shapes in abundance, and echinate elongate and
polygonal epidermal/hairbases shapes common, and tuberculate/echinate elongate and
scrobiculate elongate (>50um) shapes sporadic in the inflorescences. In the vegetation,
rondels (10um) and globular ovate shapes were abundant, with echinate elongate and
crenate elongate shapes common, and scrobiculate elongate (>50um) shapes rare
(McCune 2014:100). The following table notes my findings, with PI referring to the
production index (see chapter 4).
Species Cell type
Elymus Long
glaucus Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
Ligulate to collumnate epidermal with psilate texture, E3a
Entire epidermal with psilate texture, E4
Blocky epidermal, lateral striations, E7
Elongate, dendritic margins, L3
Elongate, crenate margins, L4
Elongate, aculeate margins, L5
Keeled rondels, G1c
Pyramidal rondel, G1d
Pyramidal rondel, aculeated, G1d1
Reniform shape, G1f
Lancelote style hair trichomes, G6a
Acicular hair with ovoid base with tubucurlate processes, H2b
Papillae with ligulate margins, P1
Papillae with pitted edges, P1b
Ellipsoids with tuberculate processes, S5
171
PI
U
U
U
A
C
C
C
C
C
C
C
C
C
C
C
Figure
4.4.P
4.5.C
none
4.10.T
4.11.B
4.11.G
4.12.J
4.12.N
4.12.R
4.12.V
4.12.aa
4.3.M
none
4.8.L
4.9.ee
EPHEDRA NEVADENSIS
Family: Ephedraceae
Genus: Ephedra
Species: nevadensis
Common Name: Nevada ephedra (Welsh et al. 2008:17).
Forest-Range Environmental Study Ecosystems:
FRES29 Sagebrush
FRES30 Desert shrub
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES40 Desert grasslands
(Anderson 2004a).
Production Time:
Produces leaves in spring and again in September after rains; forms buds in spring, and
cones open from March to May, with seeds maturing in early summer (Anderson 2004a).
Archaeological Artifacts:
Ephedra nevadensis pollen was found on groundstone from Wolf Village (Cummings
2011). However, nevadensis is not commonly found in Utah Valley, but E. viridis is.
Both species were included.
Ethnographic Use:
The Shoshone and Paiute also would brew a tea twigs and branches for venereal diseases.
Gilia congesta was sometimes boiled with the twigs. A tea was also taken as urination
stimulant, and a poultice from powdered twigs and branches used for sores (Rainey and
Adams 2004; Train et al. 1941:45).
Collected:
Green stem tops and woody twigs were gathered from Red Butte Gardens, Salt Lake
City. This sample was sonicated and required two and three digestions, respectively.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Ephedra nevadensis Green stems
Green stems
Wooden twigs
Phytolith
Blocky with facetate texture, S2b
Tracheid, V1
-
PI Figures
U
4.9.W
C
4.7.J
NP
EPHEDRA VIRIDIS
Family: Ephedraceae
Genus: Ephedra
Species: viridis
Common Name: green tea, Mormon tea, Brigham’s tea (Welsh et al. 2008:17).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES33 Southwestern shrub steppe
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES40 Desert grasslands
(Anderson 2001).
172
Production Time
Vegetative growth occurs during the cool season, although seeds start to develop in
spring and mature in late summer (Anderson 2001).
Frequent Associations:
Creosote bush, shadscale, fourwing saltbush, sagebrush, dropseed, serviceberry, and
junipers (Anderson 2001).
Archaeological Artifacts:
Ephedra nevadensis pollen was found on groundstone from Wolf Village (Cummings
2011). However, nevadensis is not commonly found in Utah Valley, but E. viridis is.
Ethnographic Use:
The Shoshone and Paiute would make a tea of small stems for syphilis that was also a
physic if boiled in salted water. A tea of roots was also a physic. A tea of twigs or stems
was made for venereal disesases, bladder disorders, kidney regulation, and colds, and as
a tonic or blood purifier. The tea once boiled down into a thick consistency was used for
colds as well. The tea was believed to aid blood circulation, and so was given to the
elderly. A tea was also made for stomach disorders and ulcers, and rheumatism. A tea of
dried twigs and the bark of Purshia tridentata for gonorrhea. A warm water tea of
scraped bark from Cercocarpus ledifolius and Ephedra root for diarrhea. The powder
from dried and pulverized stem applied to sores, or mixed with Pinus monophylla pitch to
make a salve. The moistened powder was applied to burns (Rainey and Adams 2004;
Train et al. 1941:45-46)
Collected:
Green stem tops and wooden twigs were gathered from Central Utah Gardens, Orem.
This sample was partially sonicated and required one and two digestions, respectively.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species Plant Tissue
Ephedra Green stems
viridis
Wooden twigs
Wooden twigs
Leaves
Leaves
Phytolith
Tracheid, V1
Irregular sub-spheroid form with ruminate texture, S1b
Blocky with facetate texture, S2b
Single raphide, O1
Small raphide-types connected together and of different
orientations, O1c
PI
C
U
U
U
U
Figures
4.7.K
none
none
4.1.A
4.1.J
ERIOGONUM OVALIFOLIUM
Family: Polygonaceae
Genus: Eriogonum
Species: ovalifolium
Common Name: cushion buckwheat (Welsh et al. 2008:586).
Forest-Range Environmental Study Ecosystems:
Sagebrush Scrub, Northern Juniper Woodland, Pinyon-Juniper Woodland, Red Fir
Forest, Lodgepole Forest, Subalpine Forest, and Alpine Fell-fields (CalFlora 2008).
Production Time:
Blooms in the summer (CalFlora 2008).
173
Archaeological Artifacts:
Eriogonum pollen was found on groundstone from Wolf Village (Cummings 2011), and
in fill from Smoking Pipe (Scott 1984). Seeds were also found in the fill of Wolf Village
(Dahle 2011).
Ethnographic Use:
The Goshute used this plant as an eye medicine, for stomach aches, and for venereal
diseases and other afflictions of sexual organs in the form of a wash or poultice
(Chamberlin 1964:351, 369). The Paiute and Shoshone would make a tea for colds made
from boiled roots (Train et al. 1941:48). The Ute also used this plant medicinally
(Chamberlin 1909:34).
Collected:
Roots and leaves were collected from Central Utah Gardens, Orem. These samples were
not sonicated.
Microfossil Production:
These phytoliths were minimally observed by Morris (2008:168). Minimal amounts of
phytoliths were also found in E. heracleoides, E. microthecum, E. racemosum and E.
wrightii (McNamee 2013:40; Morris 2008:168). These took the form of globular rugulate
echinates and cystoliths with attached hairs, with the cystoliths considered as a diagnostic
form for E. racemosum (McNamee 2013:30, 33, 40). Other forms in these species include
polygonal epidermals, blocky, and blocky angled, globulars, tracheids, segmented hairs,
and hair bases. The following table notes my findings, with PI referring to the production
index (see chapter 4).
Species
Plant Tissue Phytolith
PI Figures
Eriogonum Roots
U 4.8.C
Vascular tissue indeterminate,V2
ovalifolium Leaves
Irregular sub-spheroid form with granulate texture, S1c U 4.9.I
ERIOGONUM UMBELLATUM
Family: Polygonaceae
Genus: Eriogonum
Species: umbellatum
Common Name: sulfur buckwheat (Welsh et al. 2008:590).
Forest-Range Environmental Study Ecosystems:
Sagebrush Scrub, Northern Juniper Woodland, Yellow Pine Forest, Subalpine Forest,
Alpine Fell-fields, Pinyon-Juniper Woodland, Lodgepole Forest, Red Fir Forest, and
Foothill Woodland (CalFlora 2013).
Production Time:
Red-orange or cream hinted yellow flowers bloom in June and July (Selland 2003).
Archaeological Artifacts:
Eriogonum pollen was found on groundstone from Wolf Village (Cummings 2011), and
in fill from Smoking Pipe (Scott 1984). Seeds were also found in the fill of Wolf Village
(Dahle 2011).
Ethnographic Use:
174
The Shoshone and Paiute would mash the leaves which, sometimes combined with boiled
roots, was used for poultices for lameness or rheumatism. A decoction of roots was taken
hot for colds or for stomach aches (Train et al. 1941:48).
Collected:
Leaves were collected from Red Butte Gardens, Salt Lake City. This sample was not
sonicated, but was rinsed in distilled water.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Minimal
amounts of phytoliths have been found in Eriogonum ovalifolium, E. racemosum and E.
wrightii (McNamee 2013:40; Morris 2008:168). The following table notes my findings,
with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Eriogonum Leaves
umbellatum Leaves
Leaves
Leaves
Phytolith
Irregular sub-spheroid form with ruminate texture, S1b
Sinuate epidermal with psilate texture, E1
Favose epidermal, E5
Tracheid, V1
PI
C
U
U
U
Figures
4.9.C
4.4.C
4.5.L
4.7.L
FESTUCA OVINA
Family: Poaceae, Graminae
Genus: Festuca
Species: ovina
Common Name: sheep fescue (Welsh et al. 2008:857).
Forest-Range Environmental Study Ecosystems:
Grows between 1,000 and 13,000 feet above sea level in almost any habitat.
Production Time:
Blooms in late spring, matures in mid-summer (Ogle et al. 2010).
Frequent Associations:
Can hybridize with Idaho fescue and western fescue (Ogle et al. 2010). Often found
alongside other grasses such as bromes and wheatgrasses, as well as ponderosa pine and
sagebrush (Ogle et al. 2010).
Archaeological Artifacts:
Poaceae phytoliths were found in teeth tartar from the Provo Mounds (Yost 2009:6), as
pollen on groundstone from Wolf Village (Cummings 2011), and Hinckley Mounds
(Peterson 2016), and as starch on groundstone from Wolf Village (Cummings 2011).
Additionally, Poaceae seeds and caryopsis were found in coprolites from Spotten Cave
(Pearce 2012), and in the fill of Hinckley Mounds (Puseman 2016).
Ethnographic Use:
The Goshute ate the seeds (Chamberlin 1964:369).
Collected:
Florets with seeds were collected from Grow Wild Nursery, Salt Lake City. This sample
was not sonicated.
Microfossil Production:
According to Kerns (2001:286), Festuca species are in the subfamily Pooideae and
produce rondels. They should also produce pyramids. In the inflorescence and vegetation
of F. occidentalis, F. roemeri, F. rubra, and F. subulata, rondels and papillae were
175
abundant, crenate longates were abundant in most of the samples, and various elongates
were found in some of the samples ranging from rare to abundant, hairbases and
scorbiculate elongates were the most uncommon (McCune and Pellatt 2013:63). In F.
arizonica, long cells, trichomes, plates, and rondels have been found (McNamee
2013:99). In F. idahoensis, long wavy plates made up half the morphotypes, the rest
being long cells and keeled rondels (Morris 2008:285). The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species Cell type
Festuca Long
ovina* Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
Elongate with pilate margins, L1
Elongate with pilate to clavate margins, L1b
Elongate with entire margins, L2
Elongate with entire margins, granulate texture, L2a
Elongate, aculeate margins, L5
Elongate, aculeate margins, curled, L5a
Elongate, aculeate margins, granulate texture, L5c
Tracheid, V1
Round/oblong rondels, G1a
Keeled rondels, G1c
Pyramidal rondel, G1d
Pyramidal rondel, aculeated, G1d1
Lancelote style hair trichomes, G6a
Hair base trichome, G6b
Acicular hair, needle/rod like, H2d
Parenchyma, Y1
PI
C
U
C
C
A
A
C
U
C
U
U
U
C
U
U
U
FRAGARIA VESCA
Family: Rosaceae
Genus: Fragaria
Species: vesca
Common Name: starvling strawberry (Welsh et al. 2008:638).
Forest-Range Environmental Study Ecosystems:
FRES10 White-red-jack pine
FRES11 Spruce-fir
FRES13 Loblolly-shortleaf pine
FRES14 Oak-pine
FRES15 Oak-hickory
FRES16 Oak-gum-cypress
FRES17 Elm-ash-cottonwood
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES24 Hemlock-Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES27 Redwood
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES41 Wet grasslands
FRES44 Alpine
(Munger 2006).
176
Figure
4.10.B
4.10.G
4.10.I
4.10.M
4.11.H
4.11.M
4.11.P
none
4.12.B
4.12.K
4.12.O
4.12.S
4.12.bb
none
4.3.N
4.6.I
Production Time:
White flowers bloom in May and red fruit ripens in early summer (Anderson and
Holmgren 1996).
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The Goshute at the fruit when it was in season (Chamberlin 1964:344, 370)
Collected:
Berries were collected from Red Butte Gardens, Salt Lake City. They were sonicated.
Microfossil Production:
Evett et al. (2006:356) found no diagnostic morphotypes in Fragaria vesca, with leaf
silica percentage measuring 0.2%. The following table notes my findings, with PI
referring to the production index (see chapter 4).
Species
Plant Tissue
Fragaria vesca Berry
Berry
Berry
Berry
Phytolith
Styloid, single, O2
Styloid cluster, O2a
Prismatic, rectangle, single O5
-
PI
U
U
U
NP
Figures
4.1.M
4.1.S
4.2.N
GUTIERREZIA SAROTHRAE
Family: Asteraceae, Compositae
Genus: Gutierrezia
Species: sarothrae
Common Name: broom snakeweed, common matchweed (Welsh et al. 2008:225).
Forest-Range Environmental Study Ecosystems:
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES28 Western hardwoods
FRES33 Southwestern shrubsteppe
(Tirmenstein 1999c).
Production Time:
Yellow flowers bloom from May to October (Anderson and Holmgren 1996).
Archaeological Artifacts:
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016) on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Shoshone and Paiute would boil the leaves, wrap them in a cloth, and apply them as
a poultice for sprains and rheumatism. The application, though, could burn skin. A
decoction of the plant was taken to cure a cold. An antiseptic measles wash was made
177
when the plant was boiled with the needles of Pinus monophylla. The boiled leaves
applied in a wet cloth to top of head will stop a nosebleed (Train et al. 1941:55).
Collected:
The tops with leaves were gathered from Red Butte Gardens. This sample was sonicated.
Microfossil Production:
McLaren and Coder (2003:28) report the absence of phytoliths in the flora and seeds, but
the presence of phytoliths in the vegetative parts, such as the leaves. In particular, they
found phytoliths that can be described as “spherical, rugalose to grainy texture” and
“Subrectangular rounded ends; grainy textures lightly silicified; distinguished from grass
phytoliths by absence of visible cell nucleus” (McLaren and Coder 2003:23).
These phytoliths were minimally observed by Morris (2008:167). Yet McNamee
(2013:35) found no phytoliths when the whole plant was sampled. The following table
notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Gutierezia Leafy tops
sarothrae Leafy tops
Leafy tops
Leafy tops
Leafy tops
Leafy tops
Phytolith
Sinuate epidermal with striate texture E1a
Elongate with entire margins, granulate texture, L2a
Irregular sub-spheroid form with granulate texture, S1c
Tracheid, V1
Parenchyma, Y1
Astrosclerid, C1
PI
U
U
U
U
U
R
Figures
4.4.H
4.10.N
4.9.J
4.7.M
4.6.J
4.3.B
HEDYSARUM BOREALE
Family: Fabaceae, Leguminosae
Genus: Hedysarum
Species: boreale
Common Name: northern sweetvetch (Welsh et al.2008:457).
Forest-Range Environmental Study Ecosystems:
Found in “mountain brush, ponderosa pine, pinyon-juniper, and big sage brush vegetative
zones” (Rosales 2017).
Production Time:
In the first season of growth, no seeds are produced. Maximum yields can be acquired on
the third year, and then again every other year (Rosales 2017).
Archaeological Artifacts:
Fabaceae pollen was found on groundstone from Wolf Village (Cummings 2011), as a
seed in the fill of Woodard Mound (Richens 1983:111), and in a coprolite from Spotten
Cave (Pearce 2012.).
Ethnographic Use:
The root was used as a medicine by the Ute (Chamberlin 1909:35).
Collected:
Roots were collected from Red Butte Gardens, Salt Lake City. They were rinsed in
distilled water and needed two digestions.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I observed
entire epidermal with striate texture (E4) produced rarely in the roots. See figure 4.5.I.
178
HOLODISCUS DUMOSA
Family: Rosaceae
Genus: Holodiscus
Species: dumosus
Common Name: mountain spray, rockspirea (Welsh et al. 2008:640).
Synonym: Spiraeae dumosa, Holodiscus discolor
Forest-Range Environmental Study Ecosystems:
Cold- to warm-temperate coniferous, hardwood, and shrubland communities, particularly
Rocky Mountain Douglas-fir communities, and in curl-leaf mountain mahogany
communities (Fryer 2010).
Production Time:
Tiny, cream-white flowers bloom from June to August (Anderson and Holmgren 1996).
Fruits ripen late in summer and are dispersed by November (Fryer 2010).
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The Paiute and Shoshone would boil the leaf or the stem in a decoction for venereal
disease. This treatment was needed for a long period of time. A decoction of leaves and
stems for stomachaches, and a boiled root decoction for stomach disorders and diarrhea.
A leaf decoction was made as an emetic, and a tea of boiled stems for colds. An external
antiseptic wash was made from boiled leaves, flowers, and upper stems (Train et al.
1941:59).
Collected:
Leaves and flowers were collected from Sego Lily Gardens, Sandy City, and Red Butte
Gardens, Salt Lake City. They were sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Holodiscus Leaves
dumosa
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Leaves
Inflorescence
Inflorescence
Phytolith
Styloid, single, O2
Styloid cluster, O2a
Prismatic, rectangular, single, O5
Prismatic, hexagon, single, O5b
Sinuate epidermal with striate texture, E1a
Entire epidermal with psilate texture, E4
Crenate epidermal, E6
Acicular hair with psilate texture, unsegmented, H2b
Tracheid, V1
Parenchyma, Y1
Acicular hair with psilate texture, unsegmented, H2b
Vascular tissue indeterminate, V2
179
PI
U
U
U
U
U
U
U
U
U
U
C
U
Figures
4.1.N
4.1.T
4.2.O
4.2.S
4.4.I
4.5.D
4.5.N
4.3.K
4.7.N
4.6.K
4.3.K
4.8.D
JUNIPERUS COMMUNIS
Family: Cupressaceae
Genus: Juniperus
Species: communis
Common Name: common juniper (Welsh et al. 2008:15).
Forest-Range Environmental Study Ecosystems:
FRES11 Spruce-fir
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES44 Alpine
(Tirmenstein 1999d).
Production Time:
Junipers begin new growth in the spring, and flowers develop later from April to June.
These flowers, or cones, will remain on the plant for two years, often dropping during
August (Tirmenstein 1999d).
Frequent Associations:
Common snowberry, gooseberry currant, Oregon-grape, hairy telegraphplant, timber
milkvetch, silvery lupine, Thurber fescue, elk sedge, and bottlebrush squirreltail
(Tirmenstein 1999d).
Archaeological Artifacts:
Juniper pollen was found on groundstone from Wolf Village (Cummings 2011) and
Woodard Mound (Richens 1983:116), on groundstone and in fill from Hinckley Mounds
(Peterson 2016). Juniper seeds were found in the fill of Kay’s Cabin (Puseman and
Cummings 2001), Woodard Mound (Richens 1983:111), Wolf Village (Dahle 2011),
andHinckley Mounds (Puseman 2016).
Ethnographic Use:
The Shoshone and Paiute would boil the younger growth at branch ends, which would
produce a reddish liquid that was drunk as a blood tonic. A cold tea from boiled twigs
was drunk for venereal diseases. The seeds were eaten as a blood tonic, and for lumbago
(Train et al. 1941:61).
Collected:
Twigs and young shoots were gathered from Red Butte Gardens, Salt Lake City. This
sample was sonicated, and the twigs required two digestions.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I observed no
phytoliths in the twigs, but in the young shoots I rarely observed entire epidermal with
psilate texture (E4), and tracheids (V1). See Figure 4.5.E for E4
JUNIPERUS OSTEOSPERMA
Family: Cupressaceae
Genus: Juniperus
180
Species: osteosperma
Common Name: Utah juniper, Utah cedar (Welsh et al. 2008:15).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES40 Desert grasslands
(Zlatnik 1999b).
Production Time:
Seeds germinate and cones open in spring (Zlatnik 1999b).
Frequent Associations:
Can hybridize with other Juniper species (Zlatnik 1999b).
Archaeological Artifacts:
Juniper pollen was found on groundstone from Wolf Village (Cummings 2011) and
Woodard Mound (Richens 1983:116), on groundstone and in fill from Hinckley Mounds
(Peterson 2016). Juniper seeds were found in the fill of Kay’s Cabin (Puseman and
Cummings 2001), Woodard Mound (Richens 1983:111), and Wolf Village (Dahle 2011),
Hinckley Mounds (Puseman 2016).
Ethnographic Use:
The Southern Paiute ate the seeds and berries (Fowler 1986:73; Rainey and Adams
2004). The Goshute used the bark to line pits that stored dried fruit. The leaves were used
in a tea for coughs and colds, and the berries eaten in fall and winter after boiling
(Chamberlin 1964:372). The Ute used this plant for kindling and slow matches
(Chamberlin 1909:35)
Collected:
Berries and leaves were gathered from Red Butte Gardens, Salt Lake City. This sample
was sonicated and the berries needed two digestions.
Microfossil Production:
Morris (2008:168) and McNamee (2013:43) found J. osteosperma to be a non-producer
of phytoliths. J. deppeana also produced no phytoliths in the bark, berries, or needles
(McNamee 2013:43). The following table notes my findings, with PI referring to the
production index (see chapter 4).
Species
Plant Tissue
Juniperus
Berry
osteosperma Berries
Leaves
Phytolith
Crystal sand, O3
Tracheid, V1
Spheroids with granulate texture, S3a
JUNIPERUS SCOPULORUM
Family: Cupressaceae
Genus: Juniperus
Species: scopulorum
Common Name: rocky mountain juniper (Welsh et al. 2008:16).
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
181
PI
U/R
U
U
Figures
none
none
4.9.Z
FRES25 Larch
FRES26 Lodgepole pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
(Scher 2002).
Production Time:
Flowers develop in late summer and bloom in April, with the berries ripening by Autumn
and maturing in November or December before falling the following spring (Scher 2002).
Frequent Associations:
Can hybridize with other Juniper species (Scher 2002).
Archaeological Artifacts:
Juniper pollen was found on groundstone from Wolf Village (Cummings 2011) and
Woodard Mound (Richens 1983:116), on groundstone and in fill from Hinckley Mounds
(Peterson 2016). Juniper seeds were found in the fill of Kay’s Cabin (Puseman and
Cummings 2001), Woodard Mound (Richens 1983:111), Wolf Village (Dahle 2011), and
Hinckley Mounds (Puseman 2016).
Ethnographic Use:
The Shoshone and Paiute would make a tea of boiled terminal twigs used for venereal
disease (Train et al. 1941:62).
Collected:
Twigs were gathered from Sego Lily Gardens, Sandy City. This sample was sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I also
observed no forms in the twigs.
NICOTIANA ATTENUATA
Family: Solanaceae
Genus: Nicotiana
Species: attenuata
Common Name: coyote tobacco (Welsh et al. 2008:729).
Forest-Range Environmental Study Ecosystems:
Yellow Pine Forest, Red Fir Forest, Lodgepole Forest, Subalpine Forest, Foothill
Woodland, Chaparral, Valley Grassland, wetland-riparian (CalFlora 2009).
Production Time:
Blooms from May to October (CalFlora 2009).
Archaeological Artifacts:
Solanaceae pollen was found in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The Shoshone and Paiute employed this plant as a smoking tobacco. Often applied
externally, rarely taken internally. A tablespoonful of boiled leaves was taken internally
three times a day for worms. A weak solution of leaves, boiled or raw, for a physic or
emetic. A wet dressing of crushed leaves plus tubers of Cyperus esculentus was applied
for athlete’s foot. A decoction of boiled leaves for hives or other skin irritations. Smoked
182
leave when mixed with dried Salvia carnosa or Leptotaenia multifida for colds, asthma,
or tuberculosis. The crushed leaves were applied as is or as a poultice for swellings,
rheumatism, toothache, eczema and similar skin conditions. The chewed leaves were
applied to cuts, or bound on snakebites after poison has been sucked out. Pulverized
tobacco was applied to sores (Rainey and Adams 2004; Train et al. 1941:71-72). The
Goshute also use this plant as a source of tobacco. They used the leaves dried and alone
or with kinnikinick, Cornus stolonifera, bark (Chamberlin 1964:375). The Ute and
Shoshone also gathered tobacco from previously burned areas (Stewart 1942:251).
Collected:
A packet of seeds was received at a research conference. I attempted to grow some, but
did not produce a sufficient sample to test.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. McNamee
(2013:42) reports no phytoliths produced in Nicotiana obtusifolia. Bozarth (1997) reports
that in N. rustica leaves that large plates with irregular curvilinear edges can be found,
although they are rare.
OPUNTIA POLYCANTHA
Family: Cactaceae
Genus: Opuntia
Species: polycantha
Common Name: central prickly pear (Welsh et al. 2008:86).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands (Johnson 2000a).
Production Time:
Yellow, pink, or red flowers bloom in June and July (Selland 2003). Fruit ripens about
two or more months after flowering (Johnson 2000a).
Frequent Associations:
Pinyon, juniper, mountain-mahogany, oak, serviceberry, sage brushes, and several
grasses, among others (Johnson 2000a).
Archaeological Artifacts:
Evidences of this plant were not found in any Fremont Utah Valley sites. This plant was
included, however, because of its use by Native Historic groups and because it is found in
Utah Valley.
Ethnographic Use:
The Goshute and Utah Southern Paiute use the stems and buds for food once the spines
were removed (Chamberlin 1964:375; Fowler 1986:72).
Collected:
A pad and a bud were collected from Red Butte Gardens, Salt Lake City. This sample
was sonicated and required two digestions.
Microfossil Production:
183
I found no scholarly articles on the production of phytoliths in this species. However,
small and large druse phytoliths have been found in other Opuntia species in the piths,
cortexes, and epidermis/hypodermis (Jones and Bryant 1992). The following table notes
my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Opuntia
Bud
polycantha Bud
Pad
Spines/hairs
Pads
Pads
Buds
Spines/hairs
Phytolith
Tracheid, V1
Vascular tissue indeterminate, V2
Tracheid, V1
Irregular sub-spheroid form, ruminate to facetate texture,
S1b
Druse, O4
Druse-like, O4a
Druse-like, O4a
Druse-like, O4a
PI
C
U
U
U
Figures
4.7.O
none
4.7.O
4.9.F
C
A
C
C
4.2.H
4.2.H
4.2.H
4.2.H
PHASEOLUS VULGARIS
Family: Fabaceae, Leguminosae
Genus: Phaseolus
Species: vulgaris
Common Name: kidney bean, green bean, snap bean (Welsh et al. 2008:477).
Forest-Range Environmental Study Ecosystems:
A cultivated species that prefers well drained soil and full sun.
Production Time:
Often planted once soil temperatures are above 60oF, and matures 50-60 days after
planting.
Archaeological Artifacts:
Beans were found in the fill of Smoking Pipe (Billat 1985:91; Forsyth 1984:18), and in
the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
The Utah Southern Paiute ate the beans (Fowler 1986:73).
Collected:
No samples were collected for this study given the prolific publication and analysis that
currently exists on Phaseolus.
Microfossil Production:
Produces hook-shaped, silicified hairs (Bozarth 1990; Poveda-Diaz et al. 2016)
PINUS EDULIS
Family: Pinaceae
Genus: Pinus
Species: edulis
Common Name: pinyon, two-needle pinyon, Colorado pinyon (Welsh et al. 2008:21).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES29 Sagebrush
FRES30 Desert shrub
184
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES38 Plains grasslands
FRES40 Desert grasslands
(Anderson 2002).
Production Time:
Cones emerge in May or June and grow until the end of August when growth is paused
until the following May. Cones and seeds are fully mature by the September of their
second year of growth, after which they open (Anderson 2002).
Frequent Associations:
Often grows above or overlapping Juniperus osteosperma (Anderson 2002; Anderson
and Holmgren 1996).
Archaeological Artifacts:
Pine pollen was found on groundstone from Wolf Village (Cummings 2011), Woodard
Mound (Richens 1983:116), and Hinckley Mounds (Peterson 2016), and in the fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
When accessible, pinyon nuts were gathered, often in the fall, by most all groups,
including the Goshute, Shoshone, Ute, and Paiute (Chamberlin 1964:377; Fowler
1986:76). A few groups would travel to known groves to harvest them (Lowie 1924:201).
The nuts would be consumed raw or roasted, or ground into flour (Janetski 1991:39;
Stewart 1942:251; Yanovsky 1936:5). The inner bark was also sometimes used for food,
and the sap as a chewing gum, by the Southern Paiute (Rainey and Adams 2004). The
pitch was also sometimes made into medicine (Janetski 1991:39; Stewart 1942:251).
Collected:
Nuts, sap, and needles were collected at Red Butte Gardens, Salt Lake City. This sample
was not sonicated and required up to three digestions.
Microfossil Production:
McNamee (2013:25, 43) found no phytoliths in the needles, wood, bark, or cones, but did
find globular psilate central nuclei, half spherical psilate outline, and half spherical
granulate outline phytolith forms in the shells. Bozarth (1997) identified thick, crescent
shaped phytoliths in the pine nuts. Leaf cell types in pines are either weakly or heavily
silicified, and the phytoliths are often distinguished from other plants by the presence of
bordered pit impressions on tracheary elements (Bozarth 1993:96). I also observed the
half nuclei/crescent shapes in the nuts. The following table notes my findings, with PI
referring to the production index (see chapter 4).
Species
Plant Tissue
Pinus edulis Twigs
Nuts
Sap
Needles
Twigs
Phytolith
Prismatic, rectangular, clusters, O5a
Crescents, half-nucleir, S4
Vascular tissue indeterminate, V2
PINUS FLEXILIS
Family: Pinaceae
Genus: Pinus
Species: flexilis
185
PI
C
U
NP
NP
U
Figures
4.2.Q
4.9.cc
4.8.E
Common Name: limber pine (Welsh et al. 2008:21).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
(Johnson 2001).
Production Time:
Cones ripen from August to September and seeds are dispersed shortly thereafter
(Johnson 2001).
Frequent Associations:
Bristlecones, ponderosa, lodgepole, pinyons, and other pines, as well as firs, junipers,
aspens, maples, oaks, and sagebrush, among others (Johnson 2001).
Archaeological Artifacts:
Pine pollen was found on groundstone from Wolf Village (Cummings 2011), Woodard
Mound (Richens 1983:116), and Hinckley Mounds (Peterson 2016), and in the fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
The Western Shoshone ate the seeds (Fowler 1986:76).
Collected:
An entire cone with seeds came from Grow Wild Nursery, Salt Lake City. This sample
was sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Phytoliths
have been found in P. monophylla. Spiny body phytoliths have been reported in P.
ponderosa, but not in P. edulis (Kerns 2001:286, 292). McNamee (2013:28, 43) reports
blocky forms, elongates, globulars, and tracheary elements for P. ponderosa needles, and
no phytoliths from P. leiophylla stems or needles. In P. contorta, there were sporadic
tracheary elements, and abundant pilates and scorbiculate in the needles, and abundant
scorbiculate in the twigs (McCune and Pellatt 2013:64). The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Pinus flexulis Seeds
Nuts
Nuts
Nuts
Nuts
Nuts
Phytolith
Tracheid, V1
Styloid, single, O2
Styloid cluster, O2a
Druse-like, O4a?
Crystal sand, O3
Prismatic, rectangular, single, O5
PI
U
C
C
U/R
R
R
PINUS MONOPHYLLA
Family: Pinaceae
Genus: Pinus
Species: monophylla
Common Name: singleleaf pinyon (Welsh et al. 2008:21).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES28 Western hardwoods
186
Figures
none
4.1.O
4.1.U
4.2.I
4.2.C
4.2.P
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES40 Desert grasslands
(Zouhar 2001b).
Production Time:
Cones slowly develop in the first growing season, and grow rapidly be the second
growing season, with the seeds maturing in late summer (Zouhar 2001b).
Frequent Associations:
Can hybridize with other Pinus species. Often grows alongside junipers, sage brushes,
and other shrubs (Zouhar 2001b).
Archaeological Artifacts:
Pine pollen was found on groundstone from Wolf Village (Cummings 2011), Woodard
Mound (Richens 1983:116), and Hinckley Mounds (Peterson 2016), and in the fill from
Smoking Pipe (Scott 1984).
Ethnographic Use:
The Ute would turn pinyon nuts into a mush that was stored for winter (Sutton 1989:245).
The Western Shoshone and Utah Southern Paiute ate the nuts (Fowler 1986:76). The tree
resin was an important remedy for both groups. A tea of boiled resin was taken for colds,
venereal diseases, rheumatism, tuberculosis, influenza, chronic indigestion, bowel
trouble, fevers, nausea, diarrhea, for general debility, postpartum, and as a tonic. The
terminal twigs of Juniperus utahensis were sometimes added to the boiled resin tea to
improve flavor, and as a kidney medicine. Needles or young twigs could be used in place
of resin. The resin was also chewed for sore throats or swallowed as a pill to stop
diarrhea. A dressing of heated resin was used to draw out boils or embedded slivers, and
also used, when combined with crushed plants of Psathyrotes ramosissima or finely
chopped terminal twigs of Juniperus utahensis, for sores, cuts, swellings, and insect bites.
The pulverized resin was used as a drying agent for syphilitic sores. The heated resin
when smeared on a cloth was used to treat pneumonia, ruptures, sciatic pains, and
muscular soreness. This same hot resin poultice mixed with crushed Salvia carnos leaves
was used for chest congestion (Train et al. 1941:78-79). The Goshute would boil the gum
in water and drink it hot to fight intestinal parasites (Chamberlin 1964:350, 377). This
tree was also an important food source and a pine-nut harvest was a great event. Nuts
were consumed raw or roasted (Yanovsky 1936:5).
Collected:
Resin and needles were collected from Sego Lily Gardens, Sandy City. This sample was
sonicated and the needles required two digestions.
Microfossil Production:
Pinus monophylla was found to produce abundant blocky, prismatic and elongate
rectangular styloid forms (Plate III) (Morris 2008:144). Phytoliths were minimally
observed by Morris (2008:168). McNamee (2013:25, 43) found globular psilate central
nuclei, half spherical psilate outline, and half spherical granulate outline phytolith forms
in the shells. I did not test any shells. The following table notes my findings, with PI
referring to the production index (see chapter 4).
Species
Plant Tissue Phytolith
Pinus monophylla Resin
Tree
Needles
Elongate with aculeate margins, non grass, L5b
Needles
Tracheid, V1
187
PI Figures
NP
U
4.11.N
U
none
POA FENDLERIANA
Family: Poaceae, Graminae
Genus: Poa
Species: fendleriana
Common Name: muttongrass (Welsh et al. 2008:884).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES40 Desert grasslands
FRES44 Alpine
(Howard 1997).
Production Time:
Flowers during the spring and summer, with seeds ripening shortly thereafter (Howard
1997).
Frequent Associations:
Can hybridize with other Poa species (Howard 1997).
Archaeological Artifacts:
Poaceae phytoliths were found in teeth tartar from the Provo Mounds (Yost 2009:6), as
pollen on groundstone from Wolf Village (Cummings 2011), and Hinckley Mounds
(Peterson 2016), and as starch on groundstone from Wolf Village (Cummings 2011).
Additionally, Poaceae seeds and caryopsis were found in coprolites from Spotten Cave
(Pearce 2012), and in the fill of Hinckley Mounds (Puseman 2016).
Ethnographic Use:
Seeds were eaten (Yanovsky 1936:9).
Collected:
Florets with seeds were collected at Sego Lily Gardens, Sandy City. This sample was not
sonicated.
Microfossil Production:
According to Kerns (2001:286), Poa species are in the subfamily Pooideae and produce
rondels. They should also produce pyramids. Of a 100-short cell count of phytoliths from
leaf epidermis, 96 rondels, 3 pyramids, and one crenate were found (Kerns 2001:287).
In P. bulbosa and P. pratensis, rondels were abundant in the inflorescence and
vegetation, elongates were also frequently found, tuberculates were rare, papillae were
only found in the inflorescence of P. pratensis. Crentates and scorbiculate were rare
(McCune and Pellatt 2013:63). P. secunda produced abundant phytoliths that took the
form of long cells, plates, hair bases, papillae, and rondels (Morris 2008:171). The
following table notes my findings, with PI referring to the production index (see chapter
4).
Species
Cell Type
Poa
Long
fendleriana Long
Long
Long
Phytolith
Elongate with pilate margins, L1
Elongate with entire margins, L2
Elongate, crenate margins, L4
Elongate, aculeate margins, L5
188
PI
C
C
C
C
Figure
4.10.C
4.10.J
4.11.C
4.11.I
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Short
Elongate, aculeate margins, granulate texture, L5c
Tracheid, V1
Round/oblong rondels, G1a
Square/rectangular rondels, G1b
Keeled rondels, G1c
Trapeziform, G1e
Lancelote style hair trichomes, G6a
General, polylobe/bilobe rondel, G7b
Lancelote style hair, granulate texture, unsegmented, H1d
Papillae with ligulate margins, p1
Papillae with pitted edges, P1b
C
U
C
C
C
C
C
U
C
U
U
4.11.Q
none
4.11.C
4.11.G
4.11.L
4.11.T
4.11.cc
4.11.jj
4.3.H
4.8.I
none
PRUNUS VIRGINIANA
Family: Rosaceae
Genus: Prunus
Species: virginiana
Common Name: chokecherry (Welsh et al. 2008:654).
Forest-Range Environmental Study Ecosystems:
FRES10 White-red-jack pine
FRES11 Spruce-fir
FRES13 Loblolly-shortleaf pine
FRES14 Oak-pine
FRES15 Oak-hickory
FRES17 Elm-ash-cottonwood
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES39 Prairie
(Johnson 2000b).
Production Time:
White flowers bloom in late spring, and red fruits ripen into a deep purple in late summer
(Selland 2003; Johnson 2000b).
Frequent Associations:
Sagebrush, serviceberry, shadscale, rabbitbrushes, oaks, wood’s rose, mahogany, juniper,
and firs, among others (Johnson 2000b).
Archaeological Artifacts:
Chokecherry seeds were found in the fill of Kay’s Cabin (Puseman and Cummings 2001)
and in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
The Eastern Shoshone, Goshute, Utah Southern Paiute, and Western Shoshone ate the
fruit (Fowler 1986:78; Sutton 1989:249). The Ute gathered berries which were mashed,
formed into lumps, dried, and stored until spring (Lowie 1924:202). The Goshute would
use scrapings of wood in a decoction for bowel trouble (Chamberlin 1964:350).
The Paiute and Shoshone would make a tea from leaves or bark, or dried root for
tuberculosis, coughs and colds. A decoction of bark for indigestion and upset stomach.
Holding the head over boiling bark with steam rising into eyes was a cure for snow
189
blindness. The dried, pulverized bark was smoked for headaches and head colds, or used
as a drying powder for sores (Train et al. 1941:82-83).
Collected:
Berries, leaves, and roots were collected from Sego Lily Gardens, Sandy City, and Red
Butte Gardens, Salt Lake City. This sample was sonicated.
Microfossil Production:
Leaves have been found to produce silicified anticlinal epidermal cells, tracheids, and a
unique rhombohedral calcium oxalate crystal (Morris 2008:145, 168). The forms in the
leaves were common and the stem phytoliths were minimal. Calcium oxalate crystals
have also been found in P. laurocerasus in the form of blocky mesophyll/epidermal cells
in the leaves and stems, both of which were poorly silicified. Tracheids were also found
in the leaves, and rhombohedral CaOx crystals in the stems (McCune and Pellatt
2013:65). I also observed the rhombohedral CaOx crystals. The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Prunus
Berries
virginiana Leaves
Leaves
Leaves
Leaves
Roots
Roots
Leaves
Leaves
Phytolith
Vascular tissue indeterminate, V2
Ligulate epidermal with psilate texture, E3
Tracheid, V1
Vascular tissue indeterminate, V2
Parenchyma, Y1
Irregular sub-spheroid form with granulate texture, S1c
Vascular tissue indeterminate, V2
Single raphide, O1
Rhombohedral, O6
PI
C
C
C
C
C
C
C
R
C
Figures
none
4.4.N
4.7.P
none
4.6.L
4.9.K
none
none
4.2.V
PURSHIA MEXICANA
Family: Rosaceae
Genus: Purshia
Species: mexicana
Common Name: cliff-rose (Welsh et al. 2008:655).
Synonym: Cowania mexicana
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
(Howard 1995).
Production Time:
Yellow and white blossoms bloom from April to September (Anderson and Holmgren
1996). Seeds mature and disperse in summer up to October (Howard 1995).
Frequent Associations:
Can hybridize with other Purshia species (Howard 1995).
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
190
The Goshute used the leaves medicinally (Chamberlin 1964:367). The Paiute and
Shoshone would boil the leaves, powdered rock lichens, and another plant into a tea that
was taken twice daily as a smallpox cure. Another smallpox remedy was made by boiling
the plant tops with pitch of Pinus monophylla. An antiseptic wash was made by bowling
young tops, flowers, and leaves, or by boiling leaves with pine pitch. A tea was made
from leaves, young stems, and flowers for venereal diseases, as a physic, for colds, or for
back pains over the kidneys (Train et al. 1941:40).
Collected:
Flowers and leaves were collected from Sego Lily Gardens, Sandy City, and Central Utah
Gardens, Orem. This sample was partially sonicated.
Microfossil Production:
McNamee (2013:40) tested the whole plant and found it to be a non-producer of
phytoliths. Yet in P. subintegra, there were abundant phytoliths in the leaves and
inflorescence (none in the bark) that took the forms of tracheids, pitted vessels, and wavy
single celled hairs (McNamee 2013:41). The following table notes my findings, with PI
referring to the production index (see chapter 4).
Species
Plant Tissue
Purshia
Inflorescence
mexicana Leaves
Leaves
Phytolith
Irregular sub-spheroid form with granulate texture, S1c
Vascular tissue indeterminate, V2
PI Figures
NP
R
4.9.L
U
none
PURSHIA TRIDENTATA
Family: Rosaceae
Genus: Purshia
Species: tridentata
Common Name: bitterbrush (Welsh et al. 2008:655).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
(Zlatnik 1999c).
Production Time:
Yellow blossoms bloom in spring (Anderson and Holmgren 1996) to July, fruits ripen
from July to September (Zlatnik 1999c).
Frequent Associations:
Often grows alongside sagebrush (Anderson and Holmgren 1996). Can hybridize with
other Purshia species (Zlatnik 1999c), and grow with firs, pinyons, pines, sage brushes,
and spruces.
Archaeological Artifacts:
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
This plant was used extensively by the Paiute and Shoshone, in particular to treat
smallpox, chicken pox, and measles by an internal mendicant by boiling leaves, and
sometimes young branches and flowers. Sometimes it quickened the appearance of the
191
measles rash. A decoction was applied as a wash that also treated these three diseases. A
decoction of the plant as an external wash was used as an antiseptic for itches, rashes,
skin eruptions, scratches, or insect bites. The green leaves were mashed and applied as a
wet dressing for sores, or dried leaves dusted as a powder for sores. A tea of a boiled leaf
decoction, inner trunk bark, or roots for venereal disease and gonorrhea. This tea would
be prepared in quantity and stored. A tea of boiled leaves for colds, pneumonia, liver
trouble, or as a blood or general tonic. An inner bark tea for internal rupture. The inner
bark strips were used in a wash over swellings. An external wash made of boiled young
twigs for rashes. A tea of inner white or outer dried bark for tuberculosis. A tea of boiled
leaves, and/or twigs used as a physic or emetic. The strength of the physic or emetic was
determined by water to leaf ratio. A physic was also made by boiling unground seeds
(Train et al. 1941:86-87).
Collected:
Leaves were collected from Red Butte Gardens, Salt Lake City. This sample was
sonicated.
Microfossil Production:
Produces minimal amounts of phytoliths in the leaf and no phytoliths in the stem (Morris
2008:168). Blinnikov (2005:82) found no phytoliths in P. tridentata at all. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species Plant Tissue Phytolith
PI Figures
Purshia Leaves
Irregular sub-spheroid form with granulate texture, S1c U 4.9.M
tridenta Leaves
Blocky with facetate texture, S2b
U 4.9.X
RHUS AROMATICA
Family: Anacardiaceae
Genus: Rhus
Species: aromatica
Common Name: skunkbush (Welsh et al. 2008:36-37).
Variation: Two variations exist in Utah, with variation trilobata present in Utah Valley.
Forest-Range Environmental Study Ecosystems:
FRES14 Oak-pine
FRES13 Loblolly-shortleaf pine
FRES15 Oak-hickory
FRES18 Maple-beech-birch
FRES38 Plains grasslands
FRES39 Prairie
(Taylor 2004).
Production Time:
Yellow blossoms bloom before leaves emerge in March or April (Anderson and
Holmgren 1996). Fruits develop and mature about eight to nine weeks later (Taylor
2004).
Archaeological Artifacts:
Evidences of this plant were not found in any Fremont Utah Valley sites. This plant was
included, however, because of its use by Native Historic groups and because it is found in
Utah Valley.
Ethnographic Use:
192
The Shoshone, Goshute, Northern Ute, Ute, and Utah Southern Paiute ate the berries
(Chamberlin 1964:883; Chamberlin 1909:36; Fowler 1986:70). Both green and ripe
berries were eaten, and dried berries were also gathered. The berries were also stored for
winter. A drink was made with fresh berries (Ebeling 1986:126). The Paiute would make
an astringent for smallpox sores used the dried and powdered fruits (Train et al. 1941:86).
The root was used as a yellow dye. The leaf was sometimes mixed with tobacco (Rainey
and Adams 2004).
Collected:
Berries and leaves were collected from Central Utah Gardens, Orem, and Sego Lily
Gardens, Sandy City. This sample was not sonicated.
Microfossil Production:
Other Rhus species, such as choriophylla, have been found to be non-producers of
phytoliths (McNamee 2013:32). The following table notes my findings, with PI referring
to the production index (see chapter 4).
Species
Plant Tissue
Rhus aromatic Berries
Berries
Leaves
Leaves
Leaves
Leaves
Phytolith
Vascular tissue indeterminate, V2
Parenchyma, Y1
Entire epidermal with striate texture, E4a
Irregular sub-spheroid form with granulate texture, S1c
Tracheid, V1
Parenchyma, Y1
PI
U
U
U
U
U
U
Figures
4.8.F
4.6.M
4.5.J
4.9.N
4.7.Q
4.6.M
RIBES AUREUM
Family: Saxifragaceae
Genus: Ribes
Species: aureum
Common Name: golden currant, Lewis’ currant (Welsh et al. 2008:684).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir-spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands
(Marshall 1995).
Production Time:
Bright yellow flowers bloom in early spring, as early as March (Selland 2003). Seeds
mature in summer (Marshall 1995).
Archaeological Artifacts:
Evidences of this plant were not found in any Fremont Utah Valley sites. This plant was
included, however, because of its use by Native Historic groups and because it is found in
Utah Valley.
Ethnographic Use:
The Goshute ate the fruits, which they also dried in quantity and preserved for winter
(Chamberlin 1964:379). The Ute ate the berries (Chamberlin 1909:36). The Paiute and
193
Shoshone would use the inner bark, dried, pulverized, and applied as a powder for sores,
or made into a tea for leg swellings (Train et al. 1941:86).
Collected:
Berries were collected from Central Utah Gardens, Orem. This sample was sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I observed
irregular sub-spheroid form with ruminate texture (S1b) produced uncommonyl in the
berries. See Figure 4.9.D.
ROSA WOODSII
Family: Rosaceae
Genus: Rosa
Species: woodsii
Common Name: Wood’s rose (Welsh et al. 2008:657), mountain rose, wild rose
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES19 Aspen-birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES24 Hemlock-Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES27 Redwood
FRES28 Western hardwoods
FRES29 Sagebrush
FRES31 Shinnery
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
FRES44 Alpine
(Hauser 2006).
Production Time:
Pale to dark pink flowers bloom in late May (Selland 2003) up to July (Hauser 2006).
Rose hips that contain seeds turn bright red and mature in the fall (Anderson and
Holmgren 1996).
Archaeological Artifacts:
Rosa seeds were found in the fill of Kay’s Cabin (Puseman and Cummings 2001).
Rosaceae pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), and in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The Goshute, Eastern Shoshone, Utah Southern Paiute, Southern Ute, and Ute ate the
berries (Chamberlin 1909:36; Chamberlin 1964:379; Fowler 1986:78). The Paiute and
Shoshone used the ripe fruits as food. The pulpy seed was also used to soothe the lower
intestinal tract. A tea of steeped leaves was a popular beverage. A tea of boiled roots or
inner stem bark was used for colds. The plant was used as a tonic or cold remedy as a
physic. A root decoction was effective in stopping diarrhea and used for intestinal
influenza and bloody diarrhea, and for urination failure. The plant was used to dress
sores, cuts, wounds, burns, and swellings. Parts of the plant, roots, wood, or inner bark
stem were applied dry or moistened, or scraped into a fine powder and then inserted into
194
a wound and covered with a bandage. Claims were made that this improved healing
(Train et al. 1941:88-89).
Collected:
Berries and leaves were gathered from Central Utah Gardens, Orem, and Red Butte
Gardens, Salt Lake City. This sample was sonicated and the berries needed two
digestions.
Microfossil Production:
Produces phytoliths in the leaves, and production can be considered common; there is no
phytolith production in the stems or thorns (McNamee 2013:42; Morris 2008:144, 168).
The leaves were found to produce polygonal epidermal, polygonal thick epidermal, hair
base or hair base fragments, and stomates (McNamee 2013:24, 26, 41). Calcium oxalates
have been found in the leaves and stems of R. gymnocarpa, both spherical and blocky
forms. Puzzle piece epidermal forms were also found in the leaves (McCune and Pellatt
2013:65). The following table notes my findings, with PI referring to the production
index (see chapter 4).
Species
Rosa
woodsii
Plant
Tissue
Berries
Leaves
Leaves
Leaves
Leaves
Leaves
Phytolith
PI Figures
Spheroids with ruminate texture, S3b
Sinuate epidermal with psilate texture, E1
Polygonal epidermal with psilate texture, E2
Tracheid, V1
Parenchyma, Y1
Prismatic, hexagon, single, O5b
U
C
C
C
U
U
none
4.4.D
4.4.K
4.7.R
4.6.N
4.2.T
SAMBUCUS CAERULEA
Family: Caprifoliaceae, Adoxaceae
Genus: Sambucus
Species: caerulea
Common Name: blue elderberry (Welsh et al. 2008:98).
Alternate:
A profusion of synonyms and alternate spellings create a confusing taxonomy (Crane
1989).
Forest-Range Environmental Study Ecosystems:
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir - spruce
FRES24 Hemlock - Sitka spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES27 Redwood
FRES28 Western hardwoods
FRES34 Chaparral - mountain shrub
FRES35 Pinyon – juniper
(Crane 1989)
Production Time:
Small white flowers bloom in late spring and early summer (Selland 2003). Powdercovered dark-blue fruit develops in late summer (Anderson and Holmgren 1996).
Frequent Associations:
Serviceberries, chokecherries, roses, sage brushes, among others (Crane 1989).
Archaeological Artifacts:
195
Seeds were found in the fill of Woodard Mound (Richens 1983:111).
Ethnographic Use:
The Goshute ate the berries fresh, dried, or cooked (Chamberlin 1964:380; Yanovsky
1936:57).
Collected:
Berries were gathered from Central Utah Gardens, Orem. This sample was not sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I observed no
phytolith forms in the berries. I did observe CaOx crystal sand (O3) as uncommonly
produced in the berries. See Figure 4.2.D.
SAMBUCUS RACEMOSA
Family: Caprifoliaceae, Adoxaceae
Genus: Sambucus
Species: racemosa
Common Name: red elderberry (Welsh et al. 2008:99).
Forest-Range Environmental Study Ecosystems:
Quaking aspen/red elderberry communities, in riparian zones, in fir-spruce conifer
communities (Fryer 2008).
Production Time:
Flowers bloom in June and July, fruits and seeds mature in late July to mid-August
(Fryer 2008).
Archaeological Artifacts:
Seeds were found in the fill of Woodard Mound (Richens 1983:111).
Ethnographic Use:
The Eastern Shoshone, Utah Southern Paiute, Western Shoshone and Goshute ate the
berries in season (Chamberlin 1964:380; Fowler 1986:72).
Collected:
Berries were gathered from the BYU Herbarium. This sample was sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. No phytoliths
have been found in S. canadensis (Tedford 2009:189). I found no silica phytoliths in the
berries.
SARCOBATUS VERMICULATUS
Family: Amaranthaceae, Chenopodiaceae
Genus: Sarcobatus
Species: vermiculatus
Common Name: greasewood (Welsh et al. 2008:138).
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES29 Sagebrush
FRES30 Desert shrub
FRES33 Southwestern shrubsteppe
FRES35 Pinyon-juniper
FRES36 Mountain grasslands
FRES38 Plains grasslands
FRES40 Desert grasslands
196
(Anderson 2004b).
Production Time:
Flowers in May up to August, seeds mature from September to November (Anderson
2004b).
Frequent Associations:
Saltbushes, rabbitbrushes, sage brushes, wildryes, sacatons, wheatgrasses, among others
(Anderson 2004b).
Archaeological Artifacts:
Sarcobatus pollen was found on groundstone from Wolf Village (Cummings 2011) and
Hinckley Mounds (Peterson 2016), as well as in the fill of Smoking Pipe (Scott 1984) and
Hinckley Mounds (Peterson 2016).
Ethnographic Use:
The Utah Southern Paiute ate the seeds (Fowler 1986:73). The Paiute claimed this was a
remedy plant of past generations. They burnt the whole plant to charcoal, mixed the
charcoal with water, and drank it three times daily for diarrhea. A charcoal of branches
was made into a drink for diarrhea or rectal bleeding (Train et al. 1941:92).
Collected:
Leaves were gathered from Nine Mile Canyon. This plant was sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. I observed
tracheids (V1; Figure 4.7.S) and blocky forms with facetate texture (S2b) produced
uncommonly in the leaves.
SHEPHERDIA ARGENTEA
Family: Elaeagnaceae
Genus: Shepherdia
Species: argentea
Common Name: silver buffaloberry (Welsh et al. 2008:354).
Forest-Range Environmental Study Ecosystems:
FRES17 Elm-ash-cottonwood
FRES21 Ponderosa pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
(Esser 1995).
Production Time:
Flowers bloom before leaves emerge in early spring (Selland 2003). Fruits ripen later in
the summer, and seeds disperse in the fall.
Frequent Associations:
Often grows alongside serviceberry, sage brushes, rabbitbrush, greasewood, grasses,
among others (Esser 1995).
Archaeological Artifacts:
Evidences of this plant were not found in any Fremont Utah Valley sites. This plant was
included, however, because of its use by Native Historic groups and because it is found in
Utah Valley.
Ethnographic Use:
197
The Western Shoshone, Utah Southern Paiute, Northern Ute, Ute, and Goshute ate the
berries (Chamberlin 1909:36; Chamberlin 1964:381; Fowler 1986:73).
Collected:
Berries and leaves were gathered Sego Lily Gardens, Sandy City. This sample was
sonicated.
Microfossil Production:
Found to produce umbrella shaped, peltate trichomes “up to 0.5 mm in diameter, with a
radiation pattern of linear cells” (Warner 1989:235). Piperno (2006) does not note
anything on phytolith production of plants in the Elaeagnaceae family. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Shepherdia Berries
argentea
Berries
Leaves
Leaves
Leaves
Phytolith
Entire epidermal with psilate texture, E4
Umbrela peltate trichome, V4a
Polygonal epidermal with psilate texture, E2
Tracheid, V1
Umbrella peltate trichome, V4a
PI
R
C
U
U
U
Figures
4.5.F
4.6.P
4.4.L
4.7.T
4.6.P
SHEPHERDIA CANADENSIS
Family: Elaeagnaceae
Genus: Shepherdia
Species: canadensis
Common Name: soapberry, Canada buffalo berry (Welsh et al. 2008:355).
Forest-Range Environmental Study Ecosystems
FRES10 White - red - jack pine
FRES11 Spruce - fir
FRES15 Oak – hickory
FRES17 Elm - ash – cottonwood
FRES19 Aspen – birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir - spruce
FRES25 Larch
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper
FRES38 Plains grasslands
FRES44 Alpine
(Walkup 1991).
Production Time:
Yellow or brown flowers bloom in April to June; red, bitter berries ripen from July to
August (Walkup 1991).
Archaeological Artifacts:
Evidences of this plant were not found in any Fremont Utah Valley sites. This plant was
included, however, because of its use by Native Historic groups and because it is found in
Utah Valley.
Ethnographic Use:
The berries were eaten for food by the Eastern Shoshone (Fowler 1986:73).
Collected:
Berries and leaves were gathered from Thanksgiving Point, Lehi. This plant was partially
sonicated and required up to three digestions.
Microfossil Production:
198
Found to produce umbrella shaped, peltate trichomes “up to 0.5 mm in diameter, with a
radiation pattern of linear cells” (Warner 1989:235). Piperno (2006) does not note
anything on phytolith production of plants in the Elaeagnaceae family. The following
table notes my findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Shepherdia Berries
canadensis Leaves
Leaves
Phytolith
Blocky with facetate texture, S2b
Irregular sub-spheroid form with granulate texture, S1c
Tracheid, V1
PI
U
U
U
Figures
4.9.Y
4.9.O
none
SOLANUM JAMESII
Family: Solanaceae
Genus: Solanum
Species: jamesii
Common Name: James’ potato (Welsh et al. 2008:732).
Forest-Range Environmental Study Ecosystems:
Prefers slightly acidic soil, dislikes wet or heavy clay soils (Hanson 2007:70-71). Can be
found in pinyon-juniper and mountain-brush communities (Welsh et al 2008:732).
Production Time:
Plant in spring after the last frost, and then harvest in autumn after frost has killed the
foliage. Tubers are small and grow at the ends of the root tips. Advised not to eat raw
(Hanson 2007:70-71).
Archaeological Artifacts:
Solanum jamesii-type starch was found in the tooth tartar of an individual from the Provo
Mounds (Yost 2009:6).
Ethnographic Use:
While there is no documentation for the use of Solanum jamesii, there is documentation
for the consumption and cultivation of Solanum tuberosum by the Goshute (Chamberlin
1964:382).
Collected:
Tubers and leaves were gathered by Dr. Allison from his backyard. This sample was
sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Phytoliths
have been found in S. elaegnifolium, S. dulcamara, and S. sarrachoides (McNamee
2013:42; Morris 2008:170). Blocky tabular scrobiculate, blocky crystalline, elongate fusiform
thin psilate and tabular scrobiculate, and globular thick silica outlines, and tracheids in the leaves
for S. elaegnifolium (McNamee 2013:28-30, 42). The following table notes my findings,
with PI referring to the production index (see chapter 4).
Species Plant Tissue
Solanum Leaves
jamesii
Leaves
Leaves
Leaves
Leaves
Phytolith
Styloid, single, O2
Styloid cluster, O2a
Dumbbell bundle, O1d
Crystal sand, O3
Druse, O4
PI
C
C
C
U
R
199
Figures
4.1.P
4.1.V
4.1.K
4.2.F
4.2.J
Species
Plant Tissue
Tubers
Leaves
Leaves
Leaves
Leaves
Phytolith
Tracheid, V1
Irregular sub-spheroid form, ruminate to facetate texture, S1b
Spheroid with granulate texture, S3a
Tracheid, V1
Indeterminate vsculare issue, V2
PI
R
U
U
U
U
Figures
4.7.U
none
4.9.aa
4.7.U
4.8.G
SOLIDAGO CANADENSIS
Family: Asteraceae, Compositae
Genus: Solidago
Species: canadensis
Common Name: common goldenrod (Welsh et al. 2008:264).
Forest-Range Environmental Study Ecosystems:
Grows in most ecosystems, cover types, and alongside most plants (Coladonato 1993).
Production Time:
Pale golden-yellow flowers bloom from July to October (Anderson and Holmgren 1996).
Seeds are dispersed during fall and winter.
Archaeological Artifacts:
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016), on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Goshute would collect and eat the seeds (Chamberlin 1964:382).
Collected:
Florets were gathered from Red Butte Gardens, Salt Lake City. This sample was
sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Jigsaw-puzzle
pieces, branched tracheary elements, and silicified hairs have been noted in Solidago
rigida (Bozarth 1992). The following table notes my findings, with PI referring to the
production index (see chapter 4).
Species
Plant Tissue
Solidago
Inflorescence
canadensis Inflorescence
Inflorescence
Inflorescence
Phytolith
Lancelote hair with psilate texture, segmented, H1c
Irregular sub-spheroid form with granulate texture, S1c
Tracheid. V1
Indeterminate vascular tissue, V2
SPHAERALCEA MUNROANA
Family: Malvaceae
Genus: Sphaeralcea
Species: munroana
Common Name: Munro’s globemallow (Welsh et al. 2008:509-510).
Forest-Range Environmental Study Ecosystems:
200
PI
U
U
U
U
Figures
4.3.F
4.9.P
4.7.V
none
Often found alongside sagebrush in desert plains to low mountain slopes (Pavek et al.
2011).
Production Time:
Tangerine to brick-red colored flowers bloom from May to September (Anderson and
Holmgren 1996).
Archaeological Artifacts:
Sphaeralcea seeds were found in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
It is unclear how often the Shoshone distinguished between the different species of
Sphaeralcea. A drink of boiled roots or whole plant was taken for a long period of time
for venereal diseases and gonorrhea; this treatment also acted as a physic and emetic. A
tea of boiled roots was taken as a contraceptive, and a weak solution of this could be used
for upset stomach. A solution from boiled leaves was used as an eyewash or taken
internally as a hot tea for colds. The crushed raw root was applied to swellings. The entire
boiled plant was used as a dressing for cuts on horses. The plant once wilted in hot water
was bandaged to rheumatic sores or swellings (Train et al. 1941:93-94). The Goshute
pounded the plant in water into a gummy paste, which they then applied over rough inner
surfaces of earthen dishes. They also sometimes used this paste on wicker vessels after
they had been pitched with pine gum (Chamberlin 1964:374)
Collected:
Leaves were collected from Red Butte Gardens, Salt Lake City. This sample was
sonicated.
Microfossil Production:
I found no scholarly articles on the production of phytoliths in this species. Phytoliths not
present in Sphaeralcea ambigua (McNamee 2013:39). Minimal amounts of phytoliths
were found in S. grossulariifolia (Morris 2013:168). The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Sphaeralcea Leaves
munroano
Leaves
Leaves
Leaves
Leaves
Phytolith
Polygonal epidermal with granulate texture, E2a
Entire epidermal with striate texture, E4a
Tracheid, V1
Stomata, V3
Parenchyma, Y1
SPOROBOLUS AIROIDES
Family: Poaceae, Graminae
Genus: Sporobolus
Species: airoides
Common Name: alkali sacaton (Welsh et al. 2008:897).
Forest-Range Environmental Study Ecosystems:
FRES29 Sagebrush
FRES30 Desert shrub
FRES32 Texas savanna
FRES33 Southwestern shrubsteppe
FRES35 Pinyon-juniper
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
(Johnson 2000c).
201
PI
U
U
U
U
U
Figures
4.4.M
4.5.K
none
4.6.D
4.6.O
Production Time:
Blooms in the summer, seeds produced thereafter and into October (Johnson 2000c).
Frequent Associations:
Can hybridize with other Sporobolus species. Often grows alongside other grasses,
shrubs, sages, and Juniper (Johnson 2000c).
Archaeological Artifacts:
Sporobolus seeds were found in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
The seeds were parched, ground, eaten dry or made into a mush (Yanovsky 1936:9)
Collected:
Florets with seeds were gathered from Red Butte Gardens, Salt Lake City. This sample
was not sonicated.
Microfossil Production:
Forms in airoides includes saddles, long indented cells, and pyramidal rondels, as well as
other shapes such as bilobates (Morris 2008:143). In the blade and stem of S. contractis
were found long cells, trichomes, saddles, bilobates, and crescent plates. In the blade and
stem of S. cryptandrus were rondels, saddles, long cells, and trichomes and buliforms. In
the blade and stem of S. interruptus, long cells, trichomes, hair cells, saddles, bilobes, and
plates, among others, were found (McNamee 2013:98). Saddles, bilobates, and very few
rondels were in S. cryptandrus (Morris 2013:143-144). The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species
Cell
Type
Sporobolus Long
airoides
Long
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Phytolith
PI Figures
Sinuate epidermal with psilate texture, E1
Crenate epidermal, E6
Blocky epidermal, with lateral striations, E7
Elongate with pilate margins, L1
Elongate with entire margins, L2
Elongate with entire margins, granulate texture, L2a
Elongate with entire margines, psilate texture; needle-like, L2b
Elongate, aculeate margins, L5
Elongate, aculeate margins, granulate texture, L5c
Tracheid, V1
Chloridoid saddle, G2a
Gerneal ovoid rondel, G7a
General, polylobe/bilobe rondel, G7b
C
C
C
A
A
A
A
A
A
U
C
C
C
STIPA HYMENOIDES
Family: Poaceae, Graminae
Genus: Stipa
Species: hymenoides
Common Name: Indian ricegrass (Welsh et al. 2008:901).
Alternate Name: Oryzopsis hymenoides, Achnatherum hymenoides
Forest-Range Environmental Study Ecosystems:
FRES21 Ponderosa pine
FRES29 Sagebrush
202
4.4.E
4.5.O
4.6.B
4.10.D
4.10.K
4.10.O
4.10.Q
4.11.J
4.11.R
none
4.12.W
4.12.ff
4.12ii
FRES30 Desert shrub
FRES34 Chaparral-mountain shrub
FRES35 Pinyon-juniper
FRES38 Plains grasslands
FRES39 Prairie
FRES40 Desert grasslands
(Tirmenstein 1999a).
Production Time:
Flowers in spring with seeds ripening in the summer (Tirmenstein 1999a).
Frequent Associations:
Can hybridize with other needle grasses (Tirmenstein 1999a).
Archaeological Artifacts:
Stipa hymenoides seeds were found in the fill of Wolf Village (Dahle 2011).
Ethnographic Use:
Goshute, Western Shoshone, and Utah Southern Paiute ate the seeds (Fowler 1986:76;
Rainey and Adams 2004).
Collected:
Florets with seeds were gathered from Sego Lily Gardens, Sandy City. This sample was
not sonicated.
Microfossil Production:
According to Kerns (2001:286), Stipa species are in the Stipea tribe and produce Stipea
pyramids. They should also produce simple bilobates. Abundant phytoliths have been
found in Achnatherum hymenoides and A. nevadensis (Morris 2013:167). In the blade of
A. hymenoides, smooth and granulate long cells, trichomes, hair cells, wide saddles, short
shaft flat bilobes, short shaft with one flat and one round bilobes, stipa bilobes, stipa
crenulated lobates, and crescent, round, and square plates were found (McNamee
2013:99). The following table notes my findings, with PI referring to the production
index (see chapter 4).
Species
Cell
type
Stipa
Long
hymenoides Long
Long
Long
Long
Long
Long
Long
Long
Long
Long
Short
Short
Short
Short
Short
Short
Short
Short
Phytolith
PI Figure
Entire epidermal with psilate texture, E4
Blocky epidermal with lateral striations, E7
Elongate with pilate margins, L1
Elongate with entire margins, L2
Elongate with entire margins, granulate texture, L2a
Elongate with entire margines and psilate texture, needlelike L2b
Elongate, crenate margins, L4
Elongate, aculeate margins, L5
Elongate, aculeate margins, granulate texture, L5c
Elongate with sinuate margins, L6
Tracheid, V1
Round/oblong rondels, G1a
Trapeziform, G1e
Stipa bilobate, G3a
Lancelote style hair trichomes, G6a
Hair base trichome, G6b
Bulliform, G6c
Lancelote hair with granulate texture, unsegmented, H1d
Acicular hair, needle/rod like, H2d
U
U
C
C
C
U
4.5.G
4.6.A
4.10.E
4.10.L
4.10.P
4.10.R
C
C
C
C
U
A
A
A
C
C
C
C
C
4.11.D
4.11.K
4.11.S
4.11.T
none
4.12.D
4.12.U
4.12.X
4.12.dd
none
4.12.ee
none
none
203
TYPHA LATIFOLIA
Family: Typhaceae
Genus: Typha
Species: latifolia
Common Name: broad-leaved cattail (Welsh et al. 2008:943).
Forest-Range Environmental Study Ecosystems:
Occurs in disturbed moist or wet habitats, with cattail marshes being found around Utah
Lake and the Great Salt Lake, and in FRES41 Wet grasslands (Gucker 2008).
Production Time:
Flowers in the summer, with seed production following thereafter (Gucker 2008).
Frequent Associations:
Can hybridize with other cattail species (Gucker 2008).
Archaeological Artifacts:
Typha latifolia pollen has been found on groundstone from Wolf Village (Cummings
2011), Woodard Mound (Richens 1983:116), and Hinckley Mounds (Peterson 2016), and
in the fill of Smoking Pipe (Scott 1984).
Ethnographic Use:
The Western Shoshone and Utah Southern Paiute use the root, pollen, flowers, and stalks
(Fowler 1986:79). The Paiute would burn the cattail fluff to obtain seeds to eat. They
would encapsulate pollen in plant leaves and roast it, making the pollen hard and sweet
(Ebeling 1986:115). The Goshute ate the seeds, often roasted, after burning off the
bristles of spikes (Chamberlin 1964:341, 383). The Ute also ate the seeds and shoots
(Janetski 1991:38).
Collected:
Stalk, leaf, and a pollen head were collected from Red Butte Gardens. This sample was
sonicated, and samples required two digestions.
Microfossil Production:
No silica phytoliths were observed in this plant, but calcium oxalate crystals of raphide
bundles and randomly aligned raphides have been observed (Monje and Baran 2002). I
observed no silica forms, but I did observe CaOx crystals. The following table notes my
findings, with PI referring to the production index (see chapter 4).
Species
Plant Tissue
Typha latifolia Entire stalk
Pollen head
Pollen head
Pollen head
Pollen head
Pollen head
Pollen head
Phytolith
Single raphide, O1
Raphide bundle of same orientation, O1a
Styloid, single, O2
Styloid cluster, O2a
Crystal sand, O3
Prismatic, hexagon, single, O5b
VIGUIERA MULTIFLORA
Family: Asteraceae, Compositae
Genus: Viguiera
204
PI
NP
U
U
U
U
U
U
Figures
4.1.B
4.1.D
4.1.Q
4.1.W
4.2.G
none
Species: multiflora
Common Name: showy goldeneye (Welsh et al. 2008:278).
Alternate Name: Heliomeris multiflora
Forest-Range Environmental Study Ecosystems:
Often grows in “sagebrush, juniper, cottonwood, aspen, and spruce fir communities”
(Tilley 2012).
Production Time:
Golden-yellow flowers bloom in August (Anderson and Holmgren 1996).
Archaeological Artifacts:
Asteraceae pollen has been found in fill from Smoking Pipe (Scott 1984), in fill and on
groundstone from Hinckley Mounds (Peterson 2016), on groundstone from Wolf Village
(Cummings 2011), and on groundstone from Woodard Mound (Richens 1983:116).
Ethnographic Use:
The Goshute and Utah Southern Paiute ate the seeds (Chamberlin 1964:341, 371; Fowler
1986:71).
Collected:
Failed to record where flowers were collected. This sample was sonicated.
Microfossil Production:
Abundant phytoliths in the leaves, none present in the inflorescence or stem (McNamee
2013:33). In particular, polygonal epidermal, polygonal striated epidermal, blocky,
tracheid, single celled hairs with round bases, segmented hairs, and hair bases are found
(McNamee 2013:24-26). The following table notes my findings, with PI referring to the
production index (see chapter 4).
Species
Plant Tissue
Viguiera Inflorescence
multiflora Inflorescence
Inflorescence
Phytolith
Entire epidermal with psilate texture, E4
Lancelote hair with psilate texture, segmented, H1c
Tracheid, V1
PI
U
U
U
Figures
4.5.H
4.3.G
4.7.W
ZEA MAYS
Family: Poaceae, Graminae
Genus: Zea
Species: mays
Common Name: maize, corn (Welsh et al. 2008:906).
Forest-Range Environmental Study Ecosystems:
This is a cultivated species. Corn species tend to prefer full sun and well-drained soils.
Production Time:
A good time to plant corn is when soil temperatures are above 60oF. Corn matures
between 60 to 90 days from planting.
Archaeological Artifacts:
Kernels were found in the fills of Smoking Pipe (Billat 1985:91; Forsyth 1984:17),
Hinckley Mounds (Peterson 2016), Wolf Village (Dahle 2011), Woodard Mound
(Richens 1983:111), Kay’s Cabin (Puseman and Cummings 2001), and American Fork
Cave (Hansen 1941:11). Cobs were found in West Canyon (Wheeler 1968:66), Smoking
Pipe, and Wolf Village. Pollen was found on groundstone from Wolf Village (Cummings
2011) and Woodard Mound, and in the fill of Smoking Pipe (Scott 1984).
205
Ethnographic Use:
The Goshute, Southern Paiute, and Plateau Shoshoneans consumed this plant (Lowie
1924:200; Rainey and Adams 2004). The Ute and Shoshone planted, harvested, and
stored this plant. They roasted the kernels in husks, boiled on the cob, ground then into a
flour. A meal was made from green maize or from flour, which was also used to make
breads, mush, and dumplings (Stewart 1924:255-256).
Collected:
No samples were collected for this study given the prolific publication and analysis that
currently exists on Zea mays.
Microfossil Production:
Phytoliths are produced in the cupules and glumes, in the cobs, but not in the kernels.
There are four kinds: wavy-top rondel, ruffle-top rondel, half-decorated rondel, and
irregular with short protrusions (IRP). The three rondel types are indicative of all Zea
species (Pearsall et al. 2003:619, 621), although the half-decorated form is only observed
in half of Zea species.
The wavy-top has a flat base that is longer than the body is tall. It is circular to oval in
shape, and the top is a single complete wave equal to or less than the base. The edges are
not ruffled, and one or both sides are concave. There are no horns or spikes, but it is
almost keel-like. No bilobates, saddles, or rectangles have been observed as wavy-top
maize types (Pearsall et al. 2003:613).
The ruffle-top has a flat, oval to circular shaped base that is longer than the body is tall.
The edges at the top are ruffled (cf undulating), and these ruffles are not sharp or angular.
The top of this rondel is flat and larger than the base. When viewed from above, the
ruffles can be seen expanding out and over the edges (Pearsall et al. 2003:613).
The half-decorated form has a circular to oval to square shaped base, and the upper part
(cf top) is round to square (cf puffed out). The top is decorated with no more than four
projections that are neither horns nor spikes. These decorated rondels in the cobs are
similar to short cells with aculeate processes. (Pearsall et al. 2003:613).
IRPs (irregular with short protrusions) can be observed as wide rectangles with long
parallel sides. They can be two-dimensional or three-dimensional in shape with undulate
but no crenate sides. The projections can be regularly or irregularly patterned, and sparse
or plentiful in number. The projections are often round with a bead-like head. They are
not aculeate in texture. Other IRP forms include non-rectangle shape, robust globular
body shape, gracile spherical body, tubular body, and oblong half decorated body
(Pearsall et al. 2003:616-618). IRPs are only indicative of a few species (Pearsall et al.
2003:619) and the IRP forms usually all occur together (Pearsall et al. 2003:620)
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Data Analysis Sheet
Appendix E
Site: _________________________ Artifact #: ______________________
Analyst:______________________ Slide #:_________________
Date: _______________
Page ___ of ___
L. Elongates
G. Grass Short Cell Forms
G1. Pooideae types
G1a. round/oblong:
G1b. Square/rectangular:
G1c. Keeled:
G1d. Pyramidal:
G1d1. aculeated:
G1e. trapeziform, sinuate:
G1f. Reniform:
Subtotal:
G2. Chloridoid type
G2a. Saddle:
Subtotal:
G3. Stipa types
G3a. Bilobate:
G3b. Buliform:
Subtotal:
G4. Panicoideae types
G4a. Cross:
G4b. Bilobate:
Subtotal:
G5. Zea mays types
G5a. Wavy top:
G5b. Ruffle top:
G5c. IRP, rectangular:
Subtotal:
G6. Trichomes
G6a. lancelote style hairs:
G6b. hair bases:
G6c. bulliform cf:
Subtotal:
G7. Rondel, general
G7a. circular/ovoid:
G7b. polylobes/bilobes:
Subtotal:
Other forms observed:
L1. Elongate with pilate margins. Grass-type.:
L1a. Achillea type:
L1b. pilate to clavate margins. Grass type.
L2. Elongate with entire margins. Grass-type.:
L2a. granulate texture. Grass-type.
L2b. psilate texture, needle like. Grass
type.
Subtotal:
L3. Elongate, dendritic margins. Grass-type
L4. Elongate, crenate margins. Grass-type
L5. Elongate, aculeate margins. Grass-type
L5a. curled. Grass-type
L5b. non-grass type.
L5c. granulate texture. Grass-type.
Subtotal (grass):
L6. Elongate, sinuate margins. Grass-type
L7. Elongates, echinate margins. Grass-type
L7a. granulate texture. Grass-type
Subtotal:
Other forms observed:
Total (grass):
Total (non-grass):
E. Epidermal, Articulate
E1. Sinuate epidermal, psilate texture
E1a. striate texture
E1b. heavy or light striations
E2. Polygonal epidermal, psilate texture
E2a. granulate texture
E3. Ligulate epidermal, psilate texture
E3a. striate texture
E3b. ligulate to collumnate
E4. Entire epidermal, psilate texture.
E4a. striate texture
E5. Favose epidermal
E6. Crenate epidermal
E7. Blocky epidermal, lateral striations.
Grass-type
E8. Amoeboed epidermal, Prunus type.
Other forms observed:
Total (grass):
Total (non-grass):
Total:
P. Papillae
P1. Papillae with ligulate margins:
P1a. tuberculated:
P1b. pitted edges. Grass-type.
Other forms observed:
Total (grass):
Total (non-grass):
207
Site: _________________________ Artifact #: ______________________
Analyst:______________________ Slide #:_________________
H. Hairs
S. Spheroid, Polygonal
H1. Lancelote hair
H1a.striate texture, unsegmen.
H1b. psilate texture, unsegmen.
H1c. psilate texture, segmented
H1d.granulate texture, unseg.
H2. Acicular hair
H2a. striate texture, unsegmen.
H2b. psilate texture, unseg.
H2c. ovoid base with tuberculate
processes. Elymus type
H2d. needle like.
Other forms observed:
Total (grass):
Date: _______________
Page ___ of ___
S1. Irregular sub-spheroid forms
S1a. ruminate texture
S1b. ruminate to facetate
S1c. granulate texture
S2. Blocky, psilate texture
S2a. psilate to granulate
S2b. facetate texture
S3. Spheroids
S3a. granulate texture
S3b. ruminate texture
S4. “crescents”, or half nuclei
S5. Ellipsoids with tuberculate processes.
Elymus type.
Other forms observed:
Total (non-grass):
C. Sclerids
Total:
C1. Astrosclerid
Other forms observed:
Total:
V. Vascular Tissue
V1. Tracheids
V2. Vascular tissue, unknown
V3. Stomata
V4. Trichome
V4a. Umbrella peltate
V4b. Trichome base (cf non-grass)
Other forms observed:
O. Calcium Oxalates
O1. Raphide, single: needle shaped w/ one end
pointed
O1a. bundle, same orientation
O1b. bundle, differ. Orientation
O1c. small raphide-types connected
together and of differ orientations
O1d. “dunmbbell” bundle
O2. Styloid, single
O2a. cluster
O3. Crystal sand
O4. Druse: several facets radiating from a central
core
O4a. druse-like
O5. Prismatic, rectangular, single
O5a. rectangular cluster
O5b. hexagon
O5c. hexagon cluster
O6. Rhombohedrals, Prunus-type.
Other forms observed:
Total:
Y. Parenchyma
Y1. Parenchyma
Total:
Other forms observed:
Total:__________
Total:
208
Appendix F
Phytolith and starch granule analysis of groundstone from the Wolf Village site (42UT273),
Utah Valley, Utah
by
Chad L. Yost
Ph.D. Candidate
Department of Geosciences
University of Arizona
Paleoscapes Archaeobotanical Services Team (PAST), LLC
Technical report 16022 prepared for
Madison Pearce
Graduate Studies
Department of Anthropology
Brigham Young University
April 2017
209
INTRODUCTION
Six groundstone artifacts from the Wolf Village site (42UT273) were analyzed for phytolith and starch
microremains to determine plant resources processed using these tools. Site 42UT273 is a Fremont
culture village site situated on a series of ridges north of the mouth of Goshen Canyon on the west side
of Currant Creek. The site was primarily occupied in the AD 1000s; however, some structures may have
been occupied as early as the AD 600s. Previous investigations have identified the use of both wild
gathered and cultivated plant resources (Dahle 2011; Puseman and Yost 2017).
MICROFOSSIL REVIEW
Although phytolith and starch extraction protocols are optimized for the recovery of starch and biogenic
silica from plant cells, other useful microremains and particles are often present in the extracts.
Phytoliths, starch granules and some of these other microremains recovered from the groundstone
samples are reviewed here.
Phytoliths
Phytoliths are biogenic silica (SiO2-nH2O) in-fillings and casts of plant cells (Blinnikov 2013). They are
produced by many plants, including grasses, sedges, and many economically important plants (Piperno
2006). Phytoliths range in size between 5 and 200 μm, and a single grass plant can produce 105 to 106
phytoliths (Yost and Blinnikov 2011). When plant matter decays, phytoliths are released and
incorporated into soils and sediments. One gram of dry soil typically yields 104 to 106 grass phytoliths,
and one cm3 of lake sediment typically yields 104 to 105 grass phytoliths (Yost, et al. 2013). Because of
their decay-in-place taphonomy, phytoliths tend to represent a very localized vegetation record,
whereas pollen records are influenced by both local and regional vegetation. However, depending on
the geomorphology of the study area, a proportion of the phytolith record may have been deposited
some distance from its initial place of formation due to wind or water transport. Phytoliths also differ
from pollen in that they can be produced in many different plant tissues such as roots, stems, leaves,
and seeds.
Starch granules
Starch is a plant energy storage substance composed of crystalline and non-crystalline regions made up
of amylose and amylopectin. Some of this starch forms globular, spherical, or polyhedral bodies referred
to as either grains or granules. Starch granules are found in many plant parts, but are often
concentrated in seeds, fruits, roots, and tubers. Starch granules range in size between 1 and 100
microns, and can persist in soil, artifact surfaces, cooking residue and dental calculus for tens of
thousands of years. A single plant species often produces a variety of starch granule shapes, sizes and
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forms. Some of these morphotypes overlap with those produced by other plants; however, there are
often morphotypes that are diagnostic of specific plants at various taxonomic levels. When viewed using
polarized light microscopy, starch granules appear bright white against a black background, a
phenomenon called birefringence. Chemical extraction, cooking, drying and environmental degradation
can result in reduced or even the complete loss of birefringence (Gott, et al. 2006).
Chrysophytes
Chrysophyte stomatocysts are biogenic silica structures produced by chrysophycean algae (classes
Chrysophyceae and Synurophyceae) during the resting stage of their life cycle (Wilkinson, et al. 2001).
Like diatoms, these organisms are often preserved in soils and sediments and can be used to reconstruct
past environmental conditions. Chrysophytes are primarily unicellular or colonial organisms that are
abundant in freshwater habitats throughout the world. Chrysophytes are related to diatoms, but are
distinct organisms. Chrysophyte stomatocysts are most common in fluctuating freshwater habitats of
low to moderate pH and that experience some winter freezing. Many stomatocyst types are found in
specific habitats, such as montane lakes, wet meadows, ephemeral ponds, perched bogs, and the moist
surfaces of rock and plant substrates. Chrysophyte stomatocysts can also be found in sites that are only
wet during certain seasons, such as snowmelt ponds and low swales (Adam and Mahood 1981). In coolcold temperate lakes, chrysophytes are most common in the spring, when acidic snowmelt dominates
the water chemistry. Chrysophytes are intolerant of eutrophic lake conditions (Cohen 2003).
Diatoms
Diatoms are single-celled algae with a biogenic silica cell wall. They grow in a wide range of habitats,
including the surfaces of wet plants and rocks, damp soils, marshes, wetlands, mudflats, and all types of
standing and flowing aquatic habitats (Spaulding, et al. 2010). Their silica cells, or frustules, often are
preserved in sedimentary deposits. Because individual taxa have specific requirements and preferences
with respect to water chemistry, hydrologic conditions, and substrate characteristics, the presence of
diatoms in soils and sediments can provide information about the nature of the local environment or
the sources of water used for cooking and food processing.
Sponge spicules
Freshwater sponges (Porifera: Spongillidae) are primitive members of the animal kingdom. They use
biogenic silica to form skeletons comprised of spicules and other structures such as spherasters for
support and for reproduction. Freshwater sponges inhabit a wide variety of wet habitats that include
ponds, lakes, streams, and rivers; however, they need a hard stratum for growth like submerged logs,
aquatic plant stems and rocks. They typically thrive in water that is slightly alkaline (above pH 7), and
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their abundance is negatively correlated with increasing turbidity, sediment load and salinity (Cohen
2003; Harrison 1988).
Fungal spores
The Ascomycota are the largest division of the Fungi kingdom and contain organisms such as mold,
smut, rust, yeast and mushrooms. These organisms produce microscopic sexual structures called
ascospores (fungal spores), which can often be used to identify the fungus from which they arose.
Fungal spores can persist for long periods of time in the environment and can be recovered using many
pollen, starch and phytolith extraction methods. In paleoecological and archaeological investigations,
fungal spores can be used to identify the presence of crop damaging molds like ergot (Claviceps sp.),
wheat smut (Ustilago sp.), and corn smut (Ustilago maydis), or the presence of dung fungus like
Sporormiella, which further indicates the presence of browsing and grazing animals (Davis and Shafer
2006; van Geel, et al. 2003).
METHODS
Starch granules and phytoliths were extracted from the use surfaces of the groundstone tools in
tandem, then separated and concentrated based on their unique physical properties.
Starch granule extraction
A pre-treatment wash of the groundstone tools using a wet brush, reverse osmosis deionized water
(RODI), and 10% hydrochloric acid (HCl) was first conducted to remove post-use soil and sediment. The
HCl solution was slowly dripped onto the tool surface to remove any carbonate rind that may have
developed post-use, and water was used to flush silts, clays, and carbonates liberated from the tool. This
aggressive pre-treatment step, which is based on the assumption that microfossils directly related to
tool use will be bonded and preserved within microscopic crevasses, pits and pores on the groundstone
use surfaces, is intended to remove any environmental signal that could confuse, dilute, or otherwise
obscure the food processing record.
After pre-treatment, each tool was placed within a plastic container with enough RODI water to
make contact with its use surface, and then floated within an ultrasonic water bath. Each use surface
was exposed to 5 minutes of sonication, which often required repositioning of the tool. The washes
from each artifact were collected in 50 mL centrifuge tubes and subjected to short-duration spins of 30
seconds at 3000 rpm to remove clay-sized particles that remained in suspension. This step was repeated
until the supernatant was clear. Finally, the samples were dried at room temperature under vacuum.
Lithium metatungstate (LMT) heavy liquid was set to a density of 1.8 g/ml and added to each of
the dried samples, which were then mixed using a vortex mixer and centrifuged at 1500 rpm for 10
minutes. Starch granules and microcharcoal particles, which were suspended at the surface of the heavy
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liquid, were decanted into new centrifuge tubes, rinsed with RODI, and transferred to storage vials with
99% ethyl alcohol.
Phytolith extraction
The heavy fraction from the heavy liquid step above containing most of the phytolith fraction and other
minerals was rinsed with RODI to remove the LMT and then mixed with 70% nitric acid and heated at
80° C for four hours to remove any remaining organic matter. The samples were rinsed to neutral pH
using RODI and then dried at room temperature under vacuum. The dried samples were mixed with LMT
set to a density of 2.3 g/ml and centrifuged at 1500 rpm for 10 minutes. Phytoliths, which were
suspended at the surface of the heavy liquid, were decanted into new centrifuge tubes and rinsed with
RODI. The phytolith extracts were transferred to 1.5 ml storage vials using 99% ethyl alcohol and dried
under vacuum for weight calculations (Table 1).
Microscopy and microfossil counts
For the starch samples, all extracted residue was mounted onto microscope slides using Permount in
approximately 1 mg aliquots, resulting in 1 to 4 starch slides per sample (Table 1). The slides were
scanned for starch at 200X using normal and cross-polarized light (Table 2). Microscopic charcoal
particles (microcharcoal) were present in extremely high concentrations and were not counted. Any
fungal spores observed were counted. A starch granule percent relative abundance diagram based on
the total starch count for each sample was produced (Figure 1).
For the phytolith samples, all extracted residue was mounted onto microscope slides using
Permount in approximately 1 mg aliquots, resulting in 1 to 8 phytolith slides per sample (Table 1).
Phytolith counts, and counts for other microremains, were completed for one entire microscope slide
per sample, and additional slides, if applicable, were scanned for rare and economically significant
phytoliths (Table 3). A phytolith percent relative abundance diagram based on the total phytolith count
for each sample was produced (Figure 2).
RESULTS AND DISCUSSION
Microfossil recovery
Phytoliths
Phytoliths were well preserved and counts of just over 300 per sample were easily achieved. Phytoliths
derived from cool climate C3 grasses dominated the assemblages. Phytoliths derived from warm season
C4 grasses were present but at low levels of relative abundance. Most of these C3 and C4 grass phytoliths
are likely derived from the environments that these tools were used in and ultimately abandoned.
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Although aggressive pretreatment during the extraction step is intended to remove as much of the
environmental signal as possible, not all of these phytoliths can be removed without loosing a portion of
the tool use signal.
Wavy-top rondel phytoliths unequivocally diagnostic of maize (Zea mays) glumes were
recovered from all of the tools, indicating that maize kernels were ground (Figure 3a,b). To be
considered a maize (Zea mays) wavy-top rondel, all of the requirements as outlined by Pearsall et al.
(2003) must be met. The main characteristics are that maize wavy-top rondels have a circular to oval
base in outline (top view) that is flat, not concave in side view; the base must be longer than the body is
high or tall; the top (the side opposite the rondel base) is a single, complete wave that is equal to or less
than the length of the rondel base; and the peak or sides of this wave are not horns or spikes. Wavy-top
rondels with the “cf” prefix in Table 1, Figure 2 and Figure 3d,e typically lack a strong expression of one
of these characteristics, yet they are likely to be derived from maize. However, modern reference
material from western wheatgrass has yielded wavy top rondels that look very similar to Zea mays, so
phytolith workers in the Great Basin need to be extra carful when identifying maize using wavy-top
rondels.
Other diagnostic maize phytoliths observed include IRP-types (rectangular phytoliths with
irregular short protrusions; Figure 3i) and ruffle-top rondels (Figure 3g). IRP phytoliths are the product of
epidermal silicification in the fruitcase, cupule, glume, and other inflorescence tissues of maize, teosinte,
and some non-Zea (wild) grasses found in the tropics and subtropics (Piperno and Pearsall 1993). Ruffletop rondels have been observed from both North American (Bozarth 1993) and South American
(Pearsall, et al. 2003) varieties of maize. The main characteristics are that maize ruffle-top rondels have
a circular to oval base in outline (top view); the width of the body must be longer than the body is high
or tall; the edges of the top (the side opposite the rondel base) are ruffled or undulating, and the top
does not have any acute or sharply angled edges. Ruffle-top rondels can be produced in large numbers
in the glume material for some varieties of maize, and can be completely absent in other varieties.
Lastly, pyramidal rondels with a double-wall in top view (Figure 3f,h; Figure 4a) that are not diagnostic of
maize but that are highly likely to be derived from maize were recovered from all of the groundstone
tools.
Dendritic phytoliths (dendriforms) diagnostic of grass (Poaceae) inflorescence material were
recovered from all of the tools, indicating that seeds from local grasses were also ground with these
tools. Dendriforms originate in the bract material (lemmas, paleas and glumes) that surrounds the seed
(caryopsis) of many wild and domesticated grasses. Dendriforms can sometimes be found on the
surfaces of grinding tools because the dendriform-bearing plant material that encapsulates the grass
seed is never entirely removed from all of the grains during parching, winnowing, and other pre-grinding
steps. Dendriforms were either recovered as disarticulated dendritic morphotypes (Figure 4d) or
articulated in sheets of silicified epidermis (Figure 4b,c).
Stipa-type bilobates diagnostic of tribe Stipeae grasses were recovered from 5 of the 6
groundstone tools. Indian rice grass (Achnatherum hymenoides [syn. Oryzopsis hymenoides]) is a Stipeae
214
grass that produces this type of phytolith in its leaf and inflorescence structures, and most likely the
grass represented by this morphotype.
Rondel phytoliths derived from common reed (Phragmites australis) were recovered from most,
and reed canarygrass (Phalaris sp.) phytoliths were recovered from all groundstone tools. However, it is
unclear if these phytoliths are derived from the environment or are part of the subsistence signal, as
they are produced in leaves and sheaths, and not unique to inflorescence structures.
Phytoliths diagnostic of various sedge (Cyperaceae) plant parts were observed on all of the
tools, sometimes at high levels of relative abundance. This is interesting since lightly silicified sedge
phytoliths are typically underrepresented in paleoenvironmental phytolith assemblages due to poor
preservation. Although these phytoliths could be derived from the surrounding environment, the
recovery of sedge phytoliths diagnostic of seeds (Figure 5g) and starchy rhizomes (Figure 5h,i) suggests
intentional use by the site’s occupants. The most common sedge phytolith type recovered was a
morphotype termed “thin with ridges”, which is derived from sheath and leaf epidermis (Figure 5j,k).
The lowermost portion of sedge sheath structures attach very close to the starchy root crown and would
have easily been ground together with the rhizomes, which have a very fibrous texture.
Starch granules and fugal spores
Starch granule recovery was good to excellent, and ranged from a low of 8 to a high of 135 granules.
Polyhedral granules with a centric hilum were the dominant starch morphotype recovered (Figure 6d).
The roots of some sedges (Cyperaceae) and many C4 grasses produce a variety of polyhedral types, in
particular some members of the subfamily Chloridoideae and many members of the subfamily
Panicoideae including Setaria, Andropogon, Panicum, and maize (Zea mays) produce these polyhedrals
(Messner 2011). Polyhedral types are not as common in cool season C3 grasses, and when they do occur,
as with Phalaris, they tend to be very small. However, Indian rice grass (Achnatherum hymenoides)
produces polyhedral types that are typically larger than those produced by most Cyperaceae, C3 and C4
grasses, but smaller than those produced by some varieties of maize. Based on the phytolith record,
maize, Indian rice grass, and sedges may all be the source of these starches. Polyhedral faceted starches
diagnostic of maize (Zea mays) were also very numerous and recovered from all of the tools (Figure 6ac).
Starch granules derived from cool season C3 Triticeae grasses were recovered from all of the
groundstone tools (Figure 6f-h). The Triticeae is a tribe of grasses within the Pooideae subfamily that
includes many wild and domesticated cereal grains such as wheat, barley and rye. Wild Triticeae seeds
commonly used in North America for subsistence include barley (Hordeum sp.) and rye (Elymus,
Leymus). Because many starch grain morphologies overlap between Triticeae taxa, secure identifications
can be difficult to obtain. This problem is exacerbated when grinding and cooking damage is present. It
does not appear that Hordeum is represented in any of the Triticeae starches recovered, as Elymus and
Leymus, and possibly Pascopyrum appear to be the best matches (Perry and Quigg 2011).
215
Starches diagnostic of roots were recovered from 5 of the 6 tools analyzed, and indicate the use
of Apiaceae (celery) family (Figure 6j), Typha (cattail), Liliaceae (lily) family roots and tubers (Figure 6i,k).
It is noteworthy that elongated Liliaceae starches derived from either Calochortus or Fritillaria were
recovered from 5 of the 6 tools. Although Calochortus is the most likely source for the elongated root
starch (Herzog 2014; Scholze 2011), Fritillaria is also a possibility, as it shares a similar starch
morphology (Shujun, et al. 2007). There is a need for a comprehensive comparative study of Calochortus
and Fritillaria root starch for western US taxa. Also, eccentric shaped Triteleia-type starches were
recovered from one groundstone tool. Triteleia is closely related to Brodiaea and Dichelostemma, other
Liliaceae taxa that produce edible tubers (corms). In addition to Triteleia, Dichelostemma also occurs in
Utah, so additional comparative work is needed to securely identify these Triteleia-type root starches.
Interestingly, Ustilago maydis (corn smut) fungal spores were recovered from two of the tools
(Figure 6o). Ustilago spp. typically infect the inflorescences of many wild and domesticated grasses
(Fischer 1937), including maize, and their spores can be found on processing tools and incorporated into
prepared and stored foods.
Diatoms, sponges, and Chrysophytes
The silica remains of various aquatic organisms were also recovered from the groundstone surfaces.
Diatoms (siliceous algae) and sponge spicules (freshwater sponges) are indicative of wet environmental
conditions and the presence of ponds, lakes and streams, but also can be derived from water used for
processing foods prior to milling. Chrysophytes are another type of siliceous algae, and their presence
indicates oligotrophic water conditions or water sourced from spring snowmelt.
Groundstone tools
FS 219
Groundstone FS 219 is characterized by the lowest starch granule recovery and the highest relative
abundance of sedge phytoliths observed from all of the tools. Sedge phytoliths recovered include root
and achene (seed) types, indicating that sedges were processed with this tool. Grass inflorescence
dendriforms were also numerous, indicating grass seeds were processed. Many phytoliths either
diagnostic of maize or highly indicative of maize were observed.
One phytolith diagnostic of Commelina cf. angustifolia was observed (Figure 5e). Commelina is a
weedy plant of disturbed areas and agricultural fields. It is possible that the presence of Commelina in
archaeological samples may be an indication of agricultural activities. Eichhorn (2010), in their study of
Commelina plants and phytoliths in West Africa, have linked the occurrence of difference species of
Commelina with specific types of anthropogenic disturbance, such as cultivated fields, heavily fertilized
fields, periodically inundated fields, village vegetation, ruderal plant communities, and fallow fields.
216
This groundstone had the lowest total starch granule recovery (8), and a low concentration (0.08
per cm ) but yielded a diverse assemblage of granules derived from maize kernels, various grass seeds,
and starchy roots and tubers from Typha (cattail), and a member of the lily family
(Calochortus/Fritillaria)(Figure 6k). Two large lenticular starch grains with visible lamellae were
recovered. These characteristics are consistent are consistent with western wheatgrass (Pascopyrum
smithii); however, it is possible that these grains are Elymus types that were enlarged and lamellae
made visible due to grinding, as this type of grinding damage has been observed (Crowther 2012; Henry,
et al. 2009).
2
FS 2357
Groundstone FS 2357 is characterized by a relatively high recovery of grass dendriforms (Figure 4d) and
dendritic epidermis sheet elements, indicating that grass seeds were processed with this tool. Some of
the dendritic sheet elements exhibited curvilinear (Figure 4b) or straight line breaks (Figure 4c) across
the short axis of the long cells that are not typical of natural breakage of the silica sheet, but rather
suggest breakage from cutting or rolling pressure from the grinding process. Further, many of these
sheet elements are slightly darkened from exposure to fire, suggesting parching of the seeds before
grinding. A relatively high occurrence of rondel phytoliths derived from canarygrass (Phalaris sp.)(Figure
5c) and common reed (Phragmites australis)(Figure 5d) is noteworthy, as is the presence of a sedge root
phytolith.
This groundstone tool also yielded a variety of maize-type phytoliths that included ruffle-top and
wavy-top rondels, and an IRP-type. Also, two epidermis sheet elements with maize-type pyramidal
rondels were recovered (Figure 4a). Thus, maize kernels were also ground with this tool.
Perhaps the most interesting phytolith find was the recovery of a hemispherical scalloped
phytolith (Figure 5l) produced in the exocarp of squash (Cucurbita sp.), suggesting that dried squash may
have been processed with this tool.
A total of 93 starch granules were recovered from this groundstone, yielding the second highest
concentration at 0.53 per cm2 of use area washed. The starch assemblage was dominated by polyhedral
morphotypes (Figure 6d) that could be derived from a variety of plants, including maize, Indian rice grass
and other grasses, and sedge roots. Polyhedral starches diagnostic of maize were also numerous (Figure
6a,b), including two compound clusters of maize starch (Figure 6c). Other starch granules recovered
included those from Elymus (Figure 6h) and Leymus (Figure 6f,g) grass seeds, Typha roots, Liliaceae roots
(Calochortus/Fritillaria), and root starches derived from members of the Apiaceae family (Figure 6j).
FS 11975
Groundstone FS 11975 is characterized by the lowest number of maize-type phytoliths recovered from
the entire set of groundstone analyzed. Of these nine maize types, none were the polyhedral type
217
unequivocally diagnostic of maize, but they are likely to be derived from maize. Two sedge achene
(seed) phytoliths and a sedge root phytolith were recovered, indicating that sedges were processed with
this tool. One Phalaris and one Phragmites phytolith were also recovered, but it is unclear if these are
part of the background environmental record or part of the tool use record, as they are produced in
leaves and sheaths, and not unique to inflorescence structures.
Starch granule recovery was relatively low and consisted of the typical mix of polyhedral
granules that could be derived from a variety of plants and the polyhedral types that are diagnostic of
maize. A few starches diagnostic of Elymus were observed, as were spherical granules typical of grass
seeds in general. One Liliaceae (Calochortus/Fritillaria) root starch was observed. Perhaps most
interesting was the recovery of two starch grains exhibiting cooking damage, likely from parching prior
to grinding. One of these damaged grains can be seen in Figure 6n.
FS 15814
Groundstone FS 15814 yielded numerous phytoliths diagnostic of maize, as well as the polyhedral and
pyramidal morphotypes most likely derived from maize. The most interesting aspect of the phytolith
record was the recovery of 4 clusters of pyramidal rondels that appear to be fused or altered in a way
that is similar to damage from exposure to high heat (See Figure 3h for one example). However, it’s
possible that exposure to alkaline conditions such as those achieved during nixtamalization may also
cause this type of damage. Also of note was the recovery of a maize wavy-top rondel that was darkened
from exposure to fire (Figure 3c), and a maize ruffle-top rondel.
Starch granule recovery was relatively high at 38, yielding a starch concentration of 0.34
granules per cm2 of use area washed. Numerous starches diagnostic of maize kernels and other grass
seeds were observed. A celery family (Apiaceae) root starch and a Liliaceae (Calochortus/Fritillaria) root
starch were recovered. Also of note, a starch granule with damage consistent with cooking was
observed (Figure 6l,m), suggesting some type of processing such as parching prior to grinding.
FS 16494
Groundstone FS 16494 yielded the highest number of maize-type phytoliths of the six groundstone tools
analyzed, including an IRP-type and a ruffle-top rondel. Of the 11 wavy-top maize rondels and the 10 cf.
wavy-top maize rondels recovered, four of each were darkened from exposure to fire, suggesting
processing prior to grinding. Numerous dendritic phytoliths from grass inflorescence structures were
also observed, indicating that grass seeds were ground with this tool.
Despite having a washed use surface approximately 4 times the size of all the other tools, only
12 starch granules were recovered. No root starches were observed, but starches diagnostic of maize
kernels and grass seeds were observed. Interestingly, two corn smut (Ustilago maydis) spores were
recovered (Figure 6). Though highly speculative, the high number of burned maize phytoliths and
218
presence of corn smut suggests that infected kernels may have been parched prior to grinding to
remove fungal toxins (Guzmán-de-Peña 2010).
The most interesting aspect of microfossil record from this tool was the presence of small plant
fibers (Figure 6p) that dominated the starch extraction. The fibers were so numerous they were not
counted. Also, chrysophyte cysts, golden algae found in oligotrophic waters, were present at a very high
level of relative abundance. It’s possible that fibrous edible roots from an aquatic plant that does not
produce phytoliths or starch granules, was most recently processed with this tool.
FS 16642
The phytolith record from this groundstone yielded evidence for maize and grass seed processing, as
well as strong evidence for sedge achene and root processing. One corn smut (Ustilago maydis) spore
was recovered. The most striking aspect of this tool was the very high number of starch granules
recovered, which yielded the highest starch concentration of all the groundstone at 1.06 granules per
cm2 of washed use surface. Polyhedral starches diagnostic of maize dominated the starch record.
Starches diagnostic of Elymus were also very numerous. Root starches were well represented by those
diagnostic of Apiaceae and Liliaceae (Calochortus/Fritillaria) tubers. In addition, two starch granules
possibly derived from Triteleia were recovered (Figure 6i). Triteleia is closely related to Brodiaea and
Dichelostemma, other Liliaceae taxa that produce edible tubers (corms). In addition to Triteleia,
Dichelostemma also occurs in Utah, so additional comparative work is needed to securely identify these
Triteleia-type root starches.
CONCLUSIONS
The combined use of phytolith and starch analysis to identify some of the plants that were processed
with these groundstone tools was successful in providing complimentary and confirming evidence for a
multitude of plant and specific plant-part processing. Evidence for maize (Zea mays) and grass seed
processing (Elymus, Leymus, and others) was ubiquitous. Fungal spores from corn smut (Ustilago
maydis) were recovered from two of the tools. Phytoliths from Phalaris, Phragmites and a tribe Stipeae
grass, most likely Indian ricegrass (Achnatherum hymenoides) were also ubiquitous, but present at low
levels of relative abundance. A squash (Cucurbita sp.) phytolith was recovered from FS 2357, suggesting
that dried squash was ground. Phytoliths from sedge (Cyperaceae) achenes were recovered from some
of the tools.
Because they are highly perishable, edible roots, tubers, corms and rhizomes are rarely
recovered during macrofloral investigations and are likely underrepresented in the archaeobotanical
record. However, this study demonstrated that phytolith and starch analysis can be used to detect the
219
presence of these perishable tissues. Phytoliths indicate that at least 5 of the 6 groundstone tools were
used to process sedge roots, possibly from a species of Scirpus or Schoenoplectus. Root starches
observed included cattail (Typha sp.), celery family (Apiaceae), lily family (Calochortus and/or Fritillaria),
and a Brodiaea type starch, possibly derived from a species of Triteleia. When both the starch and
phytolith results are combined, diagnostic evidence for starchy root use was observed in all 6
groundstone samples analyzed.
And lastly, the presence of burned maize and grass phytoliths, and starch damaged from wet
and/or dry heat suggest that some foods were processed before they were ground. Nixtamalization and
parching of seeds to remove chaff and fungal toxins are possible pre-grinding processes observed here.
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2011 Starch Remains and Stone Boiling in the Texas Panhandle Part II: Identifying
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222
Table 1. Ground stone washes, extract weights, and microscope slide prep
Starch Extraction (1.8 g/ml heavy liquid flotation)
Sample (FS)
Approx. area
Extract
No.
washed (cm2)
weight (mg)
219
96
0.3
2357
160
4.0
11975
102
0.7
15814
112
4.1
16494
400
2.2
16642
126
2.0
Microscope
slides made
2
4
1
2
3
2
Phytolith Extraction (Nitric acid and 2.3 g/ml heavy liquid flotation)
Sample (FS)
Approx. area
Extract
Microscope
No.
washed (cm2)
Weight (mg)
slides made
219
96
0.8
2
2357 (L)*
160
4.3
8
Slides Analyzed
All slides scanned
Slides Analyzed
#1 counted and scanned
#1 counted and scanned,
#2 and #3 scanned
2357 (H)*
10.2
0
11975
102
0.7
2
#1 counted and scanned
15814 (L)*
112
0.3
1
#1 counted and scanned
15814 (H)*
4.6
0
16494
400
1.7
2
#1 counted and scanned
16642
126
0.5
2
#1 counted and scanned
*Groundstone matrix released platy particles that did not sink using LMT, so a second flotation
using Potassium/Cadmium Iodide at 2.3 g/ml density was conducted. The heavy fraction (H) from
this separation was saved separately from the light (L) fraction.
223
Table 2. Starch granule counts
Starch granule types
219
Polyhedral: Achnatherum, maize, other grasses, sedge roots
1
Compound Polyhedral: Grasses, maize, sedge roots
Polyhedral: faceted, acute angles: Maize (Zea mays)
1
Lenticular: Elymus type
1
Lenticular: Leymus type
Lenticular: Pascopyrum smithii type
2
Spherical w/centric hilum: Poaceae
1
Bell shape w/ecc. hilum: Apiaceae
Small spherical/polyhedral: Typha
1
Eccentric - Calochortus/Fritillaria type
1
Eccentric-Triteleia type
Gelatinized (swollen) starch
Ustilago sp. fungal spore
Total starch granules counted
8
Wash area starch concentration (# per cm2) 0.08
224
2357
42
2
29
9
6
2
1
2
93
0.58
Groundstone sample
11975 15814 16494
6
13
7
16642
42
2
2
13
3
2
2
58
18
3
3
6
1
1
7
1
1
1
2
1
16
0.16
38
0.34
2
12
0.03
2
2
1
1
134
1.06
Table 3. Phytolith, diatom, sponge spicule and chrysophyte counts
Phytolith type
Trapeziform sinuate: Pooideae
Rondel-keeled: Pooideae
Rondel-angular keel: Phalaris
Rondel-plateau saddle: Phragmites
Bilobate-Stipa type: Tribe Stipeae
TOTAL C3 SHORT CELLS
Cross: Panicoideae
Bilobate: Panicoideae
TOTAL Panicoideae
Saddle: Chloridoideae
TOTAL C4 SHORT CELLS
Rondel-Pyramidal: C3 & C4 Grasses
Grass epidermis
Bulliform
Trichome
Elongate-psilate
Elongate-echinate
Trichome base
TOTAL GRASS/SEDGE TYPES
Dendriform
Dendriform [SCAN]
Dendritic epidermis frag.[SCAN]
TOTAL GRASS INFLOR. TYPES OBSERVED
Thin w/ridges: Sedge stem
Irreg. w/proj: Sedge root
Irreg. w/proj: Sedge root [SCAN]
Sedge cone cell: Cyperus-type
TOTAL SEDGE TYPES OBSERVED
Domed cylinder: Commelina sp.
Pyrimidal-cf. Tradescantia sp.
Hemispherical scalloped: Cucurbita sp.
Globular-psilate
Globular granulate: trees/shrubs
Rondel-Pyramidal: cf. Maize
Rondel-Pyramidal: cf. Maize [SCAN]
Rondel-Wavy top: cf. Maize
Rondel-Wavy-top: cf. Maize [SCAN]
Epidermis w/rondels: cf. Maize [SCAN]
Rondel-Wavy top: Maize
219
91
8
1
4
8
112
9
17
26
6
32
25
1
4
13
9
13
1
41
0
5
3
8
88
1
0
1
90
1
0
0
0
0
3
7
1
8
0
1
2357
140
16
3
4
4
167
2
2
4
2
6
39
4
3
24
41
20
0
92
1
3
5
9
7
1
0
0
8
0
0
1
1
0
2
12
1
3
2
1
Table 3. (Continued)
225
Groundstone sample
11975
15814
16494
92
145
84
40
23
39
1
1
1
1
0
0
5
14
8
139
183
132
1
2
10
11
2
14
12
4
24
17
3
4
29
7
28
37
47
42
0
0
0
4
0
3
32
8
27
38
29
43
25
11
14
2
1
1
101
49
88
0
0
1
1
1
2
0
1
1
1
2
4
30
4
12
1
0
1
0
0
0
2
0
0
33
4
13
0
0
0
0
0
1
0
0
0
0
0
0
6
6
5
5
1
5
0
11
4
4
1
5
0
1
5
0
0
1
0
0
3
16642
100
33
1
1
0
135
9
17
26
9
35
41
0
0
19
32
24
0
75
0
1
1
2
42
0
2
1
44
0
0
0
0
2
1
6
1
3
0
3
Phytolith type
Rondel-Wavy top: Maize [SCAN]
Rondel cluster: Maize
Rondel: Ruffle-top [SCAN]
IRP-type: Maize glume [SCAN]
TOTAL MAIZE TYPES OBSERVED
PHYTOLITHS COUNTED (for percentages)
219
4
0
0
0
24
306
2357
0
0
1
1
23
318
Other microremains observed
Sponge spicule
Sponge spheraster
Sponge gemmosclere
Diatom-Aulacoseira sp.
Diatom-pennate
Diatom-centric
Chrysophyte cysts
3
1
2
5
2
0
1
3
0
1
1
17
1
2
226
Groundstone sample
11975
15814
16494
0
3
8
0
4
0
0
1
1
0
0
1
9
22
33
354
302
323
6
14
0
6
1
2
2
0
1
0
2
20
0
13
3
5
0
0
14
1
137
16642
5
0
0
1
20
335
2
6
0
0
9
2
13
Figure 1. Starch granule relative abundnace based on total starch count
227
228
Figure 2. Phytolith and aquatic organism relative abundance based on the total phytolith counts. Taxa counts observed during microscope slide
scans are listed as numbers in the
Figure 3. Selected maize (Zea mays) and maize-type phytoliths. The white 10-μm scale bar in image I
applies to all images. A-B) Wavy-top rondel phytoliths diagnostic of maize glumes. C) Burned wavy-top
maize rondel in side (left) and top (right) views; note double walls in top view. D-E) Wavy-top rondels
likely derived from maize (cf. maize). F) Pyramidal, double walled rondel phytolith typical of maize but
not unequivocally diagnostic of maize. G) Ruffle-top rondel diagnostic of maize in top (left) and oblique
(right) views. H) Sequence of maize-type pyramidal rondels. I) Sheet of IRP-type phytoliths diagnostic of
maize.
229
Figure 4. Selected epidermis sheet elements and dendritic long cell phytoliths recovered from
groundstone FS 2357. The white scale bar equals 10-μm. A) Sheet element with wavy-margin long cells
and double-walled rondels typical of maize (Zea mays). B) Dendritic epidermis sheet element with a
curvilinear break across the short axis of the long cells, which are slightly darkened from exposure to
fire. C) Dendritic epidermis sheet element with a linear break across the short axis of the long cells,
which are darkened from exposure to fire. D) Disarticulated and broken dendritic long cell.
230
Figure 5. Selected phytolith and diatom micrographs. The white 10-μm scale bar in image O applies to all
images. A) Trapeziform sinuate derived from grass subfamily Pooideae leaves. B) Keeled rondel derived
from grass subfamily Pooideae leaves and inflorescence structures. C) Plateau saddle in top (left) and
side (right) views, derived from Phragmites australis leaves. D) Angular keeled rondel derived from
Phalaris sp. leaves. E) Domed cylinder (bottom view) derived from Commelina cf. angustifolia seed coats
F) Pyramidal phytolith in top (upper) and side (bottom) views, derived from Tradescantia sp. seed coats.
G) Achene (Seed) cone cell derived from a sedge (Cyperaceae) species. H-I) Asteriform phytoliths
derived from sedge roots/rhizomes (cf. Scirpus, Schoenoplectus, or Cyperus). J-K) Thin with ridges
phytoliths derived from sedge (Cyperaceae) leaf and sheath epidermis. L) Hemispherical scalloped
phytolith in top (left) and side (right) views, derived from squash (Cucurbita sp.) exocarp. M) Aulacoseira
sp. diatom. N) Pennate diatom fragment (lower left) and freshwater sponge gemmosclere roulette
(upper right). O) Epithemia sp. diatom.
231
Figure 6. Selected starch granules, fungal spores, and plant fiber micrographs. The white 10-μm scale
bar in image P applies to all images. Darkfield images are viewed using cross-polarized light. A-B)
Polyhedral faceted and angular starch granules diagnostic of maize (Zea mays). C) Cluster of maize
232
polyhedral starch granules. D) Polyhedral starch morphotype that can be derived from a wide variety of
plants including grass seeds (Poaceae), maize (Zea mays) kernels, and sedge (Cyperaceae) roots. E)
Spherical starch granule diagnostic of grass seed (Poaceae). F-G) Leymus-type lenticular grass seed
starch; these granules could be damaged Elymus-type starches. H) Lenticular starch diagnostic of Elymus
sp. seed. I) Elongated polyhedral with eccentric hilum root starch most likely derived from the Liliaceae
genera Triteleia or Brodiaea. J) Bell-shaped starch granule typical of Apiaceae (Celery) family root starch,
and possibly derived from biscuitroot (Lomatium sp.). K) Large, elongated Liliaceae root starch with
eccentric hilum most likely derived from Calochortus or Fritillaria. L) Damaged starch granule from
cooking (wet/dry heat) prior to grinding. M) Same starch granule as in L but viewed from the side. N)
Damaged (gelatinized starch granule). O) Corn smut (Ustilago maydis) fungal spore. P) Unknown plant
fibers from FS 16494.
233