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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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Mimosa pudica L.

Accepted
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
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Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
Mimosa pudica L.
🗒 Synonyms
synonymEburnax pudica (L.) Raf.
synonymMimosa hispidula Kunth
synonymMimosa pudica var. pudica
🗒 Common Names
Creoles and pidgins; French-based
  • Mamzelle Marie, Marie-honte, Herbe mamzelle, Honteuse femelle, Trompe la mort (Guyane)
  • Mari hont, Zèb Mamzèl (Antilles)
  • Radié lan mort, Sensitive (Guyane)
  • Sing-sing (Taki taki Guyane)
Créole Maurice
  • Sensitive
Créole Réunion
  • Sensitive
  • Trompe la mort
Créole Seychelles
  • Herbe sensible
  • Sensitive
  • Sansib
English
  • Touch me not, Sensitive plant, Common sensitive plant
French
  • Petite sensitive (Nouvelle-Calédonie)
Malagasy
  • Ramirena, Ramoregna, Rohitra, Roimena
Other
  • M'zungu n'tsuo, M'zungu (Shimaore, Mayotte)
  • Fatsiky ambili, Fodya rafino, Balabala maitso, Shibala maitso (Kibushi, Mayotte)
📚 Overview
Overview
Brief

Code

MIMPU

Growth form

Broadleaf

Biological cycle

perennial

Habitat

terrestrial
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ravi luckhun
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    Diagnostic Keys
    Description
    Global description
     
    Mimosa pudica is an herbaceous to woody plant, often spread on the ground but somtimes erect, reaching 20 to 50 cm, sometimes up to 1 m high. It is hairy to glabrous, with scattered curved spines. The leaves are compound, alternate, on the top of a petiole, 2 to 6 cm long. 2 to 4 principal leaflets arranged in finger like shape, which are in turn composed of 10 to 28 pairs of linear to oblong divisions of the leaflets. The leaves are sensitive and close when touched. The flowers are grouped in pink balls at the end of a stalk at the base of the leaves. The fruits are agglomerated flat pods, composed of 3 to 4 articles and with long stiff hairs on the edge.
     
    Cotyledons
     
    The cotyledons are shortly stalked, oval, glabrous, with the base and the top truncated.
     
    First leaves

    First  leaves are alternate, composed. The first leaf has only one pinnae with a petiole 6 to 14 mm long, glabrous and 3 pairs of divisions of the leaflets. The divisions of the leaflets are slightly asymmetric, 1 to 2.5 mm long with pubescent margin. The second leaf has 2 pinnae with 3 or 4 pairs of division of the leaflets.
     
    General habit
     
    Plant is diffuse, usually creeping but sometimes erect. It is a small sub-shrub with thorny branches, 20 to 50 cm in height, but can reach 1 m.
     
    Underground system
     
    The plant has a deep, robust taproot.
     
    Stem
     
    The stem is cylindrical, solid. It is rapidly lignified at the base, loosely hairy, and sparsely covered with weakly curved spines. It is well branched with branches reaching up to 2 m long, often vermillion or purple in colour.
     
    Leaf
     
    Leaves are alternate, composite, held by a petiole, 2 to 6 cm long, pubescent, smooth or with few thorns. At the base of the petiole are two linear stipules, 5 to 10 mm long, with parallel venation. The leaf lamina is composed of 1 to 2 pairs of almost digitate pinnae, 4 to 9 cm long and 0.8 to 1.5 cm wide. Each pinnae holds 10 to 28 pairs of elongate linear, sessile leaflets, 6 to 8 mm long and 1.5 to 2 mm wide. The leaflets are asymmetric at the base and apiculate at the top with a central mid-vein and secondary visible veins and with contour turning from green to purple. The margin is lined with applied stiff hairs. The upper side is glabrous while the underside bears fine hairs or some coarse hairs. The leaves are sensitive, leaflets and petioles are closed at night or when touched.
     
    Inflorescence
     
    Inflorescence consist of 1 to 4 axillary and terminal glomerules, 1 to 1.5 cm in diameter, held by a stem, 12 to 25 mm long. These inflorescences are pink to purple in colour, containing many flowers.
     
    Flower
     
    The flower consists of a tiny calyx and a campanulate corolla, 2 to 2.3 mm and with 4 stamens, 7 to 8 mm long, pink with white anthers. Between the stamens is a long white filiform style.
     
    Fruit
     
    The fruit is a sessile pod, linear-oblong, 1 to 1.5 cm long and 3 to 4 mm wide, flattened, margin with long, stiff hairs. The pods are agglomerated at the top of stalk. There are generally four articles per pod, containing a single seed. At maturity, the articles, indehiscent, separate.
     
    Seed
     
    Seed is oval to orbicular, 2.5 to 3 mm in diameter and 1 to 1.2 mm thick. Seed coat smooth and brown in colour.
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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Perenial
      Perenial

      French Guiana : Mimosa pudica is flowering and fruiting the year round because of continuous hot temperature and humidity.
      Mayotte: Mimosa pudica flowers and fruits all year round.
      New Caledonia: Seedling of Mimosa pudica is possible throughout the year when moisture conditions permits, but it usually occurs during the rainy season. The growth is rapid and the plant flowers in the year from the first heat of the dry season and the beginning of the warm period. A period of dormancy is often observed.

      Wiktrop
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        Reproduction
        Mimosa pudica is an annual or perennial plant. It reproduces by seeds. Fruits are disseminated by animals. The persistence and spread of sensitive plant is aided by its prolific seed set. There are approximately 110 seeds/g. It can complete its life cycle in 3 month. At nightfall the leaflets fold together and the rachises bend down. Flowers throughout the year. Seed germinate within 2 weeks (scarification helps); after 19 years storage, 2-percent still developed.
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          Morphology

          Growth form

          Prostrated
          Prostrated

          Leaf type

          Compound
          Compound

          Latex

          Without latex
          Without latex

          Root type

          Taproot
          Taproot

          Stipule type

          Lanceolate stipule
          Lanceolate stipule

          Pod type

          Compressed pod in section
          Compressed pod in section

          Lamina base

          asymmetric
          asymmetric

          Lamina apex

          apiculate
          apiculate
          obtuse
          obtuse

          Inflorescence type

          Pedonculate glomerule
          Pedonculate glomerule

          Stem pilosity

          Dense hairy
          Dense hairy

          Life form

          Broadleaf plant
          Broadleaf plant
          Look Alikes

           Look alikes

          Comparison between Mimosa
          M. pudica M. diplotricha
          Growth habit creeping lianescent
          Leaf-Number of pinnate 1 to 2 pairs 3 to 9 pairs
           

          Identification key for lianescent Fabaceae
          Bi-pinnate leaves 1 to 2 pairs of pinnate Mimosa pudica
          3 to 9 pairs of pinnate Mimosa diplotricha
          Pinnate leaves Clitoria heterophylla
          Trifoliate leaves Lateral leaflets are symmetrical Leaflets elliptic or oval Terminal leaflets similar to lateral leaflets Teramnus labialis
          Terminal leaftets larger than the lateral leaflets Venation very marked Cajanus scarabaeoides
          Venation slightly marked leaflets > 10 mm Desmodium adscendens
          leaflets < 10 mm Desmodium triflorum
          Extremity of leaflets wedged Leaflets narrowly lanceolate (2 cm) Macroptilium lathyroides
          Large acuminate leaflets Trigonal stem with scabrous angles Desmodium intortum
          Cylindrical stem Leaflets with a silver spot petiolule 2 mm Desmodium incanum
          petiolule 5 to 15 mm Desmodium uncinatum
          Leaflets uniform green foliole terminale large (7 cm) Centrosema pubescens
          Terminal leaflet very large (15 cm) Centrosema plumieri
          Lateral leaflets asymmetrical Terminal leaflet larger than its length Rhynchosia malacophylla
          Terminal leaflet as large as its lenght Large leaf (15 cm) Rhynchosia viscosa
          Small leaf (7 cm) Rhynchosia minima
          Terminal leaflets elongated Lateral leaflets with a single rounded lobe Macroptilium atropurpureum
          Lateral leaflet without lobe large stipules (6 mm) Lablab purpureus
          small stipules (2 mm) Mucuna pruriens

          Desmanthus virgatus and  Mimosa diplotricha can sometimes occurs in large mixed populations. These two species can then be confused for each other.
          Comparison ofMimosa diplotricha and Desmanthus virgatus
          Mimosa diplotricha Desmanthus virgatus
          Stem Prickles present Ptickles absent
          Petiole Prickles present Ptickles absent
          petiole No glands Leaf gland at the top of the petiole
          Leaflets (nb) 3 to 10 pairs 1 to 4 pairs
          Sensitive leaflets yes no
          Divisions of the leaflets (nb) 11 to 30 pairs 6 to 15 pairs
          inflorescence 1 to 3 dense, ovoid glomerules 1 spherical  tousled glomerule 
          Flower Pink white

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            Ecology

            Mimosa pudica is a weed of pastures, roadsides, coconut plantations, scrub; in fences on rivers banks; in sunny to rather heavily shaded, usually moist places; from 0-1000 m altitude.

            Comoros: Mimosa pudica is a species present on roadsides and fallow.
            French Guiana: Mimosa pudica is a characteristic species of degraded open habitats, particularly in grasslands, pastures, permanent vegetal cover in orchards, roadsides and agricultural plots. It is also very present along the dikes of the Mana rice polder.
            Madagascar: A common weed in marshland of low and medium altitude, mainly in the alluvial plains and temporarily flooded marshland.
            Mauritius: Species quite common on the island in vacant and uncultivated lots, the roadside. It is a weed of crops at all altitudes.
            Mayotte: Mimosa pudica is an exotic species widely naturalized and very common in secondarized areas of hygrophilic and mesophilic regions, along streams, ditches, roadsides, villages, crops and pastures.
            New Caledonia: This species introduced and sown as fodder in the late 19th century in New Caledonia by some breeders is now common mostly in damp places and appears along roadsides, in crops, gardens and in dry  forests .
            Reunion: M. pudica grows on very diverse types of soils. We find this plant in humid areas of Reunion Island: it grows mostly on the east side and the south of the island.
            Seychelles: M. pudica grows in all seasons and on all types of soil, mainly 0-300 m altitude.
            West Indies: Mimosa pudica is an indigenous species. It is a good indicator of overgrazed grasslands. It is undemanding on soil type but is more common and abundant in low to medium elevation wetlands. Competition from other herbaceous species has relatively little effect on its growth and development. It is common on roadsides, fallow land, wasteland and cultivated areas.

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              Miscellaneous Details
              Toxicity

              Mimosa pudica is suspected to be toxic as it contains an alkaloid, mimosin.
              Thomas Le Bourgeois
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                No Data
                📚 Habitat and Distribution
                Description

                Geographical distibution

                Madagascar
                Madagascar
                Reunion Island
                Reunion Island
                Comoros
                Comoros
                Mauritius
                Mauritius
                Seychelles
                Seychelles
                Origin

                Mimosa pudica is native to Tropical America.

                Worldwide distribution

                This species is now widely Pantropical.

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                  No Data
                  📚 Occurrence
                  No Data
                  📚 Demography and Conservation
                  Risk Statement

                  Global harmfulness

                  Mimosa pudica is a troublesome weed in the cultivation of corn, tea and rice. It can become a real plague in pastures. It is considered a weed of concern for crops in South America, the Caribbean, parts of East Africa, Mauritius, Asia and many islands in the South Pacific.
                   
                  Local harmfulness
                   
                  Benin: Rare and scarce.
                  Burkina Faso: Frequent and scarce.
                  Comoros: Species infrequent but can be locally abundant.
                  Ghana: Rare and scarce.
                  French Guiana: Mimosa pudica is a very common weed in pastures, it is an excellent species indicative of degradation of the pasture. Having a high capacity of colonization, it can very quickly develop at the expense of the fodder species. It is considered one of the most problematic weeds of pastures. It is favored by long periods of regrowth between two grazing periods. These long periods allow M. pudica to become lignified and makes subsequent consumption by livestock impossible and its trampling ineffective. It is also favored by large fluctuations in livestock load. Periods of heavy load tend to degrade the vegetation too much, favoring germination and weed development, while long low-load periods allow weeds such as M. pudica to become lignified, thus leaving them unpalatable. It is also a major species of fruit crops with permanent vehetation cover. It is present in nearly 50% of orchard fields, with an average abundance index greater than 3.
                  Madagascar: Mimosa pudica is a weed of relatively low frequency but can be locally abundant and makes (and expensive) soil preparation work (due to the weakness of the equipment of the farmers) and even access to fields culture after the rainy season, difficult.
                  Mauritius: occasional weed in crops with low harmfulness.
                  Mayotte: Mimosa pudica is a very common weed, present in 52% of cultivated plots, and abundant in fruit, forage and ylang-ylang plantations. It is particularly abundant in the north of the island.
                  Nigeria: Rare and scarce.
                  New Caledonia: At the moment it is mainly a weed of crops. It is often found in pastures where it is sometimes considered as extra forage. But it remains poorly productive, while taking the place of other forage species.
                  Uganda: Frequent and usually abundant.
                  Reunion: Present in 15% of the plots, it is quite common in the sugar cane fields, but it most often develops in the form of small scattered spots or along parcel, except in some humid  areas where it can have a dominant character. In this case, besides the danger it poses to the cane because of its competition, it hinders the maintenance work of plots by its very aggressive thorns.
                  Seychelles: common weed of cultivated areas, including tea plantations, market gardening, tubers and ornamental crops.
                  Senegal: Rare and scarce.
                  West Indies: Mimosa pudica is a very common weed species in crops. While it is not a major problem in sugarcane and banana crops, it tends to develop in dense patches in fruit and vegetable crops. It is particularly abundant and troublesome in overgrazed grasslands. It hinders the maintenance of plots and its management requires costly specific weeding when it is abundant.

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                    No Data
                    📚 Uses and Management
                    Uses
                    Medicinal : The infusion of the twigs with leaves of Mimosa pudica possess antipyretic, soothing and sudorifie properties. Apart from the toxicity from mimosine, the plant has shown a significant hypotensive, sedative and anti-inflammatory effect. An alcoholic extract has shown to lower blood glucose concentrations in diabetic rats. The alkaloidal extracts of the roots is antagonistic to both acetylcholine and histamine and histamine. A crude ethanol extract of the leaves was found to have insecticidal activity against one Trilobium species.

                    Agriculture: A crude leaf extract in ethanol shows insecticidal properties against a species of Trilobium.
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                      Management
                      Global control
                      Manual : Although the thorns od Mimosa pudica make hand pulling unpleasant, the plant can be controlled by hoeing. Cultivation can also held keep the plant under control.
                      Biological control: Individuals are often found attacked by a fungus Ramularia mimosae, which grows on the upper surface of the leaflets, forming irregular white spots.
                      Chemical : picloram (pasture), ametryne (cane), fenoprop, ioxynil plus 2,4 D.

                      Management recommandations for perennial broad-leaved weeds in rice fields: http://portal.wiktrop.org/document/show/26

                      Local control

                      French Guiana: In pasture, the best management of Mimosa pudica is to maintain a dense grassland (at least 20 cm high) with a strong livestock load over short periods and renewed very regularly. This practice allows during the load a high consumption of all the forage species and also the more or less desirable weeds, a strong trampling of the weeds, without exhausting the reserves of the meadow species. Thus the regeneration of grassland species during the period of livestock withdrawal can be very fast and very competitive compared to weeds like M. pudica. Cattle should be returned as soon as the meadow is reconstituted and as much as possible before M. pudica becomes woody and fructifies. In orchards with permanent vegetation, the rotary grinding should be intense (cutting a few cm above the ground) and renewed before the new period of seed production of M. pudica. To limit the development of this species, it is important to avoid overgrazing that triggers the germination of the seed bank and the explosionof weeds. Some forage species like Brachiaria decumbens or B. humidicola as well as Desmodium sp. and Arachis pintoi are able, if left to develop sufficiently, to limit the development of Mimosa pudica.
                      In spots of very high density, impeding the regeneration of graminaceous species, the use of a herbicide selective to  grasses (triclopyr-type woody broadleaf weeds) may be considered in young regrowth, some time after the rotary grinding. The treatment must be done in the form of a very fine spray (fog) to avoid closing the leaves.
                      Goats are the only farm animals to consume M. pudica, but their grazing requires good roasting fences.
                      Madagascar: The thorns make it very hard for manual control. Perennials, they require high doses of herbicide (glyphosate + 2,4-D). It is therefore very important to control as they occur on the edges to prevent their spread, especially as they are very harmful.
                      New Caledonia: Adequate management of pasture, avoiding overgrazing, should normally limit the extension of Mimosa pudica. It is important to keep livestock from infested plots in a quarantine plot to limit contamination of all plots of the rotation. Individuals can be removed manually. If too large stands are maintained, especially in wetlands, they should be controlled by means similar to Mimosa pigra. The combination of gyro-grinding with several applications of herbicide on the regrowths allows a good control. The active ingredients to be preferred are picloram + triclopyr.
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                        No Data
                        📚 Information Listing
                        References
                        1. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                        2. Henty E.E. and G.S. Pritchard. 1973. Weeds of New Guinea and their control. Botany bulletin No 7.
                        3. Soerjani M., Kostermans A. J. G. H., Tjitrosoepomo G. 1987. Weeds of rice in Indonesia. Balai Pustaka. Jakarta. Waterhouse, D.F. & K.R. Norris. 1987. Biological Control. Pacific Prospects. Inkata Press, Melbourne. 454 pp.
                        4. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                        5. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                        6. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                        7. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:313242-2
                        8. https://www.cabi.org/isc/datasheet/34202
                        1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                        1. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                        1. http://www.issg.org/database/species/ecology.asp?si=1002&fr=1&sts=sss&lang=FR
                        1. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad.http://idao.cirad.fr/applications
                        2. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                        3. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                        4. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                        5. https://www.feedipedia.org/node/631
                        1. Galinato M.I., Keith Moody, Colin M. Piggin. 1999. Upland Rice Weeds of Southeast Asia, IRRI.
                        2. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                        Information Listing > References
                        1. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                        2. Henty E.E. and G.S. Pritchard. 1973. Weeds of New Guinea and their control. Botany bulletin No 7.
                        3. Soerjani M., Kostermans A. J. G. H., Tjitrosoepomo G. 1987. Weeds of rice in Indonesia. Balai Pustaka. Jakarta. Waterhouse, D.F. & K.R. Norris. 1987. Biological Control. Pacific Prospects. Inkata Press, Melbourne. 454 pp.
                        4. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                        5. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                        6. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                        7. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:313242-2
                        8. https://www.cabi.org/isc/datasheet/34202
                        9. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                        10. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                        11. http://www.issg.org/database/species/ecology.asp?si=1002&fr=1&sts=sss&lang=FR
                        12. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad.http://idao.cirad.fr/applications
                        13. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                        14. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                        15. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                        16. https://www.feedipedia.org/node/631
                        17. Galinato M.I., Keith Moody, Colin M. Piggin. 1999. Upland Rice Weeds of Southeast Asia, IRRI.
                        18. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.

                        L'agroécologie pratique - Nos plantes hôtes

                        Cassandra Favale
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                        Thomas Le Bourgeois
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